d ' '-' r> ^' '• /^. a: /f . -. NEW YORK ACADEMY OF SCIENCES SCIENTIFIC SURVEY OF Porto Rico and the Virgin Islands \ VOLUME III— Part 1 Tertiary Mollusca from Porto Rico — C. J. Maury a n^ =r n ! ^! ii NEW YORK: Published by the Academy 1920 NEW YORK ACADEMY OF SCIENCES SCIENTIFIC SURVEY OF Porto Rico and the Virgin Islands VOLUME III Part 1 Tertiary Mollusca from Porto Rico — C. J. Maury NEW YORK: Published by thf: Academy 1920 I, \ TERTIARY MOLLUSCA FROM PORTO RICO AND THEIR ZONAL RELATIONS By Carlotta Joaquin a Maury INTKODUCTION The collection of Porto Rican fossil mollusca upon which the results embodied in this paper are based, was made by Dr. Chester A. Reeds of The American Museum of Natural History, during the summer of 1915, under the auspices of the New York Academy of Sciences and the Porto Rican Government, The American Museum cooperating. Unlike the beautifully preserved fossils in the northern part of the neighboring island of Santo Domingo, where the shells in the , Miocene blue clays of the Yaqui Valley are as perfect as those on the recent beaches, the fossils from Porto Rico are nearly all in the form of external or internal molds. The majority have been preserved in a creamy or light bufl:' limestone from which all the substance of the shells has been removed by leaching, leaving merely either the imprint of the exterior impressed upon the stone, or else the hardened mud which filled the internal spaces of the shell. Add to this the fragmentary state of the material, and the difficulty of correct determination can be imagined. By making artificial casts of the external molds and comparing them very care- fully with actual shells from , Santo Domingo, Jamaica and else- where, I could confront myself with undoubtedly true identities. The internal molds were very unsatisfactory and the majority were simply left untouched. But in a few cases where these molds showed the exact form and peculiarities of muscle scars, or colu- . mella plications in reverse, or other internal features, one could feel reasonably sure they were the internal casts of certain shells. Sometimes the situation was saved by finding among a great many internal molds a single imprint of the exterior which clinched the identification. I beg to offer my sincerest thanks to Dr. W. H. Dall, Honorary Curator of the Department of Molluscs, United States National Museum, and to Professor G. D. Harris, Department of Paleontol- ogy, Cornell University, for their valuable suggestions, as to the stratigraphic age of the faunas from the Collazo shales and the 2 SCIENTIFIC SURVEY OF PORTO RICO Campanile Zone. I am also indebted to i)r. Dall for his help oji certain very puzzling identifications of some imfigured species h.v comparisons with types at the National Museum. CHARACTERISTICS OF THE FAUXAS Disappearance of Genera^ and Pacific Affinities As in other Antillean Tertiar)^ collections one notes in this from Porto Rico two striking peculiarities; first, the presence of certain genera which have now entirely vanished from the whole Caribbean region, and second, the singular fact that in some cases the recent species most like the fossil form is not now living in the surround- ing seas but is in the distant waters of the Pacific. This is due tc miration through the Isthmus: the establishment of West Coast and Pacific colonies; and the extinction of the ancestral Antillean stock. The cause of this extinction is a complete mystery, for the larger portion of analogous recent species are Atlantic and West Indian, and are evidently descendants of Oligocene and ]\[iocene forms which have continued on unchanged in the Antillean seas. This fact prevents our calling upon any violent or profound changes to explain the annihilation of the genera and species concerned, and the question remains open. Range of Species In brief, the vertical range of muUuscan species through rlie generalized Porto Rican section is short and limited. Only three percent of those I studied are common to the green shales and the limestone. Of those the omnipresent form is Teredo incrassala (Gabb) which ranges throughout the section and is extremely abun- dant. From the distribution of this species in the type region, Santo Domingo, I had thought of it as a ^NLiocene form only; but Dr. T. W. Vaughau now lists it from the Upper Oligocene of the Emperador limestone, in the Panama Canal Zone. So it is not surprising to find these Teredo tubes also in the Porto Rican Oligo- cene horizons. ' Area umhonata Lamarck occurs in the Collazo green shales and in the Quebradillas and Ponce limestones. This is in harmony with the wide range of this species from the Upper Oligocene of the Tampa formation to the recent fauna, althougli I have carried it down lower by placing the shales tentatively in the Middle MAURY, TERTIARY MOLLUSC A FROM PORTO RICO 3 Oligocene. Ostrea collazica, new species, is the third case common to the green shales and the limestone. This finely tinted oyster is highly characteristic of and abundant in the green shales, but at one locality Dr. Reeds found it extending up into the limestone. Doubtless more complete collections would give a few other species in common between the shales and limestones, but in any case I am sure they will be found to be exceptional. SIGNIFICANCE OF THE FAUNAE CHANGE The striking change in the molluscan life at the close of the depo- sition of the Collazo shales clearly marks an important interval of time and change. If the Collazo shales are correctly referred to the Middle Oligocene, this interval would mark the transition from Middle to Upper Oligocene time. CORRELATION OF THE FAUNAE ZONES As a brief review of the conclusions of recent authors on Porto Rican paleontology and stratigraphy has already been given in an earlier paper^ the matter need now only be summarized by saying that the Collazo green shales were generally referred to the Eocene and the Arecibo limestone to the Oligocene period. In 1917 I suggested, reasoning merely from analogies between Berkey's description of the Porto Rican beds- and my Santo Domingan sec- tions'^ that the Collazo (Lares) shales might go with the Orthaulax zone, or with the probably older Monte Cristi range limestone, and be included in the Oligocene, w^hile the limestones about the shales might correspond to my Dominican Aphera and Sconsia formations and be of Miocene age. The present detailed study corroborates this hypothesis, and indi- cates furthermore the presence of a number of distinct faunal zones to he interpolated between the Collazo shales and the highest beds , of the Arecibo limestone. In determining these zones a hundred and one species and varieties have been worked out, of which fifty- two are known and forty-nine are new. These are discussed in the systematic part of this paper. 'C J. Maury — On the Correlation of Porto Rican Tertiary Formations with Other Antillean and Mainland Horizons. Amer. Jour. Sci., XLVIII, pp. 209-21.';, September, 1919. 2C. P. Berkey— Ann. New York Acad. Sci., XXVI, pp. 1-70, Pis. I-IU, 1915. ^C. J. Maury — Bull. Amer. Paleontoloc;y, No. 30, pp. 1-43, Pis. I-III, and Correlatiotl,- Table, 1917. f 4 SCIENTIFIC SURVEY OF PORTO RICO A description of the faunal zones was also published in my earlier paper. On the north side of Porto Rico they consist of the following groups placed in descending geological order: (1) Que- bradillas limestone with many Bowden and Dominican Miocene forms; (2) Aguadilla limestone with Orthaulax aguadillensis; (3) Lares limestone with Campanile (Portoricia) laricum ; (4) Collazo shales with dementia rabelli. On the south side are (a) the Ponce limestone with Ostrea cahohasensis Pilsbry and Brown; (b) Guanica limestone with Ostrea antigiiensis and (c) Juana Diaz shales. The true time relation between the beds on the north and south sides of the island is very difficult to determine and is only tentatively expressed. The correlation also of the Collazo shales with the Lower Culebra formation is suggested but awaits further knowledge for proof. In my report, On the Correlation of Porto Rican Tertiary For- mations with Other Antillean and Mainland Horizons (]\Iaury, 1919), I followed Dr. Dall in correlating the Flint River beds near Bainbridge with the Tampa formation of Florida. The moUuscan evidence would seem to justify this relationship of these horizons especially since they both contain the striking forms, Orthaulax in- ornatus and 0. pugna-x. Vaughan, however, believes that the Flint River beds are older than the Tampa formation and that they are stratigraphically equiv- alent to the Antigua formation. His conclusion is based on tield sections and on evidence derived from the corals and foraminifera. In the letter dated October 7, 1919, Dr. Vaughan writes: ''My stratigraphic placing of the Flint River coraliferous chert is based on several different kinds of evidence. One is the comparison of numbers of sections made along Flint River from Bainbridge down stream to below River Junction, Florida, and stratigraphic studies across Florida eastward from Apalachicola River. With regard to the coral fauna in the Flint River chert, there is very little in com- mon with the coral fauna of Tampa — Orthaulax inornatus and Orthaulax pugiiax are both species of considerable stratigraphic range and neither one of them can be used as a basis for close geologic correlation." Now, if we lower the Flint River beds to the Antiguan, should we also drop the Lares limestone with Campanile laricum down to the same level? This would in turn suggest dropping the Collazo shales into Lower instead of Middle Oligocene. Beds of Lower TENTATIVE CORRELATION OP THE PORTO RICAN TERTIARY FORMATIONS Porto Rico Santo Domingo Haiti Jamaica Angullla Antigua Panama Florida Georgia Cuba O o 5 o s I 44 O ■<-( CO North Side Quebradlllas limestone with bowden Fauna and Metis trin- Itaria South Side Upper or Sconaia formation Gatua beds vflth Sconsia La Cruz marl Lower or Aphera rorraa- tion Mais- sade beds .-^ Bowden marl ? Lower Gatun horizon Oak Grove sands Cbipola marls Marl at Baracoa 8 o ■ C3 ■ Aguadllla limestone with Orthaulax aKuadlllen - sis Ponce chalky beds with Ostrea Orthaulax inomatuB cahoba- sensls Lares lime- stone with Campanile T Portorlcl ^ larlcum " Las Cahobes beds and Thomonds beds with O strea cahoba' Angullla foimation Emp era- dor lime- stone Upper part of Culebra formation Tampa fonnatlcn with Orthau- lax In- ornaTuB and O. pugnax Various localities of Cuba sensls f-5 o a s ^ a o A c o •>^ CQ GO « Oi- 14 c MMU Rio Collazo Shales with Clementla rabelli Guanica shaly lime- stone with Ostrea antl- ?Monte Crlsti range lime- stone Antigua fomatlon with Orthaulax guensls pugnax and Ostrea antleuen- Lower part of Culebra formation Flint River chert below Balnbrldge with Orthaulax Inomatus and 0. pugnax Coral reef at Guanta- namo ( Vaughan ) sis MAURY, TERTIARY M0LLU8CA FROM PORTO RICO 5 Oligocene age are represented by Hill's Moiitpelier white limestone and synchronous formations which are thought to be present in Cuba and Haiti. There is a structural argument for postulating a Lower Oligocene age for the Collazo shales in that they show evidence of disturbance which has greatly deformed the fossils they contain. This might well have taken place during the period of Antillean diastrophism between the deposition of the Upper Eocene and the Middle Oliogo- cene. On the other hand, the paleontological evidence suggests a younger age since the most abundant fossil is a dementia like C. dariena from Gatun. According to the field observations, however, of Berkey and of Reeds, the shales must be much older than the Gatun Forma- tion, which is about equivalent to the Quebradillas limestone, for they underlie the limestone series. The genus dementia is gen- erally thought to have originated in the Oligocene, but Professor Harris^, has latelv referred a St. Maurice Eocene shell, Dosinia mercenaroidea Aldrich, to dementia. This, however, was a primi- tive form just differentiating from its relatives, while the shell from the Collazo shales is a typical, well defined dementia, cer- tainly of the same group as dementia dariena and possibly identical, though more likely ancestral. Two of the Areas in the shales are forms which might run back even into Early Oligocene time; but one species is a later type resembling a Culebra shell. In view of these faunal relations, I do not think that the Collazo shales will prove older than Middle Oligocene. The purely paleontological results given herein must be weighed by other investigators and harmonized with the actual succession showed by the field sections. It will very likely then be found that certain of the faunal zones in my correlation table should be some- what raised or lowered. Vaughan's recent geological survey of the island of Santo Domingo will doubtless reveal the presence there of new faunal horizons some of which may bring out more clearly the interrelation of the Porto Rican horizons. Meantime the following correlation table is sug- gested as an approximation towards the true sequences and relation- ships of the Tertiary molluscan zones of Porto Rico. *Bull Amer. Paleontology, No. 31, p. 151. 1919. 6 SCIENTIFIC SURVEY OF PORTO RICO SYSTEMATIC DESCRIPTION OF SPECIES CLASS PELECYPODA Ordee Prionodesmacea Genus Area Linngeiis We are much indebted to Dr. Pearl Sheldon of Cornell University for critically examining and sending iis valuable notes upon the Areas. In a letter, dated June 27th, 1919, Dr. Sheldon writes: There are at least fourteen different species, many of them new, in the Porto Rican collection, but only a few of them can be described (owing to their very imperfect state), and only one certainly identified as an old species, A. umbonata. The single valve of Area cf. cumanensis is the best preserved specimen in the collection, but one of the hardest to identify, because the two valves would be discrepant, and it might be A. cumaiien:is which is unfigured and only briefly described. The collection seems to be very different from the Santo Domingan Areas There are no species in common that can be identified except A. umbonata, and Barbatia reticiclata. . . . The comparisons are chiefly from Panama or other distant localities. Concerning this striking diii'erence between the Porto Rican and Dominican Areas, as observed by Dr. Sheldon, it is important to note that the specimens were sent to her with station numbers only, and that very few happened to be from the horizon I had identified, unknown to her, as equivalent to the Dominican and Bowden Mio- cene. Of these latter she refers one to A. umbonata and the other to a new species nearest a Dominican shell that I described as A. cibaoica. Dr. Sheldon's results from the Areas thus strengthen the evidence I have gathered from the other molluscan genera, namely that in the Porto Rican collection we have to do with several distinct faunas, only one of which is like that of Bowden. Area umbonata Lamarck Area umbonata Lamarck, 1819, Anim. s. Vert.. VI, p. 37. Area umbonata Philippi, 1847. Abbild. u. Beschr., Ill, p. 13, PI. XVII. h. figs. 3 a-c. Area imbricata (Bruguiere) Gabb (in part) 1873, Trans. Amer. Phil. Soc. XV, p. 254. Area umbonata Arango, 1879, Fauna Malacologica Cubana, p. 261. Area umbonata Ball, 1898, Trans. Wagner Inst. Sci., Ill, pt. 4, pp. 620, 659; 1900, Idem pt. 5, PI. XXXVIII, figs. 4, 4 a. Area umbonata Sheldon, 1916, Paleontographica Americana I, p. 8, PI. I, figs. 12-17. Area umbonata Maury, 1917, Bull. Amer. Paleont., No. 29, p. 164, PI. XXX, fig. 11. Area umbonata Vaughan, 1919. Bull. 103 U. S. Nat. Mus., p. 564. MAURY, TERTIARY MOLLUSCA FROM PORTO RICO 7 Lamarck described the type of this species from a recent shell from Jamaica. Yet, curiously, it has not turned u}) in the Miocene (Bowden) beds of that island, although 1 collected it from syn- chronous deposits in Santo Domingo on the Rio ('ana, and now it appears in Reeds's collection from Porto Rico. Area umbonata has a wide geological and geographical range. It is found in the upper Oliogocene of the Tampa formation; the lower Miocene of the Chipola marls. Pleistocene of the Florida Keys and the Antilles, and is living from Cape Hatteras to Santa Caterina, Brazil, and also in the Antilles. The Porto Rican molds of this ark-shell are readily distinguished from the other Areas by the sharp umbonal ridge characteristic of this species. Because of its nestling habit the form of the shell varies considerably. This species is one of the best represented Areas in Porto Rico, and the localities show its presence in both the green shales and the limestones, and on the northern and southern sides of the Island. Apparently the shell was comparatively common as contrasted with its presence in the Dominican blue clays in which we found but a single valve, although other Areas were wonderfully abundant and varied. Localities*. — Rio Collazo, near San Sebastian, Stations 35, GO; near Quebradillas, Station 134; near Ponce, Station 292. Also a mold from Station 111 of which Dr. Sheldon notes ''I am sure this is the young of A. umbonata but cannot entirely prove it." The others were all identified as unquestionably this species. Of the specimen from Station 35 Dr. Sheldon writes ''Besides the general form this specimen has the coarse ribs on the posterior slope and the conspicuously finer, cancellated ribs anterior to the ridge which characterize A. umbonata and its Pacific allies." Subgenus Barbatia Gray Barbatia (Acar) reticulata Gmeliii Area reticulata Gmelin, 1792, Syst. Nat. VI, p. 3311. Area reticulata Chemnitz. Conch. Cab., VII, p. 193, PI. LIV, fig. 540. Barbatia (Acar) reticulata Dall, 1898, Trans. Wagner Inst. Sci., Ill, pt. 4, p. 629. Area (Barbatia) reticulata Dall and Simpson, Bull. U. S. Fish Comni. XX, pt. 1, p. 460. *The locality numbers mentioned in this paper are those assigned in the field by Dr. Re-ds. Their exact position will be recorded by him in a forthcominj report. The American Museum accession number, 440, is understood to precede all the Station numbers. 8 SCIENTIFIC SURVEY OF PORTO RICO Area (Barbatia) reticulata Sheldon, 1916, Paleontographica Americana, I, p. 20, PI. IV, figs. 8-12. Barbatia (Acar) reticulata Maury, 1917, Bull. Amer. Paleont., No. 29, p. 1G6, PI. XXX, fig. 16. There is a fragmentary imprint in a limestone block apparently made by a valve of Barbatia reticulata^ since a cast from it agrees well with shells of this species. This richly and deeply sculptured ark-shell was first developed in the Jacksonian Eocene of Mississippi, It continued on in the Oligocene of Tampa, Miocene of Bowden, of Matura, Trinidad, and of the Chipola River, Florida; Pliocene of Florida; Pleistocene of the Antilles; Recent from Cape Hatteras to the Gulf of Cam- peche. It is one of the very few fossil species still living on the Porto Rican shores and is reported by Dall and Simpson from the reefs at Ponce, from Mayaguez, Guanica, San Juan Harbor and Hucares. In 1917 I found a single valve in the Miocene bluffs of Cercado de Mao, Santo Domingo. The single external mold in the present series shows that the species was also present in Porto Rico, but seems then to have been a rare shell. Locality. — Near Ponce, Station 288. Area (Scapharca) gnajatica Sheldon and Maurj^, new species Plate I, Figure 3 Shell very small, oblong, much inflated, marked by a shallow sulcus ending at about the middle of the ventral margin. The sculpture consists of twenty-seven or twenty-eight narrow, rounded ribs, widely spaced over the anterior and posterior parts of the valve, but finer and closer in the sulcus. Cardinal area partly concealed by the limestone matrix, but probably narrow. Margin of valve crenulate. Length of shell 7 mm., height 4 mm., semi-diameter 3 mm. Five examples of this little form show that it is not the young of any other Ark in the collection, and it appears to be a new species. Its most conspicuous feature is the convexity in the umbonal region. This species recalls Scapharca cibaoica Maury from the Yaqui Val- ley, Santo Domingo, but it has a broadly inflated umbo instead of the small pointed beaks of that shell; the beaks are less anterior (at about two-fifths the length of the shell), the direction of the sinus is less oblique, so too is the general aspect of the shell, and finally the number of ribs is greater in the Porto Rican species. From Bowden also a small species, though larger than either of our Domin- MAURY, TERTIARY MOLLUSCA FROM PORTO RICO 9 ican or Porto Rican little Areas, has been described by Dall as A. howdeniana, but its general appearance is not like this Porto Ricac shell. Localities. — Four specimens from near Quebradillas, Station 130. and a mold, probably this species, from Station 128, same region. Area (Scapharca) a^adica Sheldon and Maury, new species Plate I, Figure 2 Shell small, flattened, umbo not much inflated; beaks at the anterior third; cardinal area long and narrow; ribs about twenty-seven or twenty- eight, those on the posterior third appearing as low, narrow, rounded ridges on the mold, anterior ribs probably more closely spaced, ornamentation un- known. The outline of the shell is oblong, the base being nearly parallel to the hinge which is almost as long as the shell. The posterior margin of the best preserved specimen is straight, nearly at right angles to the hinge and basal margin, and curved only at the corners. More worn specimens have an elliptical posterior margin, yet the outline remains almost bilaterally symmetrical, instead of the usual produced and pointed A7'ca form. Lrength 17 mm., height 9.5 mm., diameter 7 mm. This species may be distinguished from the young of tlie group to which Area secticostata Reeve and A. henekeni Maury belong, and from the Area transversa group, by its smaller and less inflated umbo, and especially by its flat, high, oblong and symmetrical pos- terior end. The collection affords five internal molds, showing both valves in place. Although the exterior has been dissolved away, the form of the mold is so characteristic as to be recognizable, because the species does not closely resemble any described from the Antillean region. Locality. — Near Aguada in the northwestern part of the island, Station 124. Area (Seapharea) eollaziea Sheldon and Maury, new species Plate I, Figure 4 Shell oblong, closely-ribbed, the ribs numbering thirty-two to thirty-six, simple, as wide as, or wider than the interspaces, and of almost the same size and spacing over the entire shell; anterior ribs beaded; cardinal area long and narrow, beaks at the anterior two-sevenths. Length of shell 28 mm., height 15 mm., semi-diameter 4 mm. The green shales of Rio Collazo yielded several valves of a small Scapharca of common form, but with distinctive ribbing, unlike 10 SCIENTIFIC SURVEY OF PORTO RICO that of related species. Like other specimens from these shales they have sulfered distortion from pressure. Because of the distortion of the valves it is uncertain whether the posterior end is more like that of Area oronlensis Gabb (Journ, Acad, Nat. Sci. Phila., (2), VIII, p. 346, PI. XLIV, fig. 21) from the Miocene of Panama and Costa Rica, and like Area dariensis Brown and Pilsbry (Proc. Acad. Kat. Sci. Phila., LXIII, p. 3G2, PI. XXII, fig. 10, 1911) from the Miocene of Panama, or like that of .4. gatunensis Toula ( Jahrb. der k. k. (ieol. Reichsanstalt, Wien, LXI, p. 493, PL XXX, fig. 4, 1911), also from the Gatun Miocene. Of Antillean species it re- sembles Area inc&quilateralis Guppy (Quart. Jour. Geol. Soc. Lon- don, XXII, p. 293, PL XVIII, figs. 2a, 2b, 1866) from the Miocene of Bowden, which more nearly agrees in size, and Area loscjiieniadiea Maury (BulL Amer. Paleont., 'No. 29, p. 172, PL XXX, fig. 1, April, 1917) from the Dominican Miocene. All these Isthmian and Antillean species, however, except Area oronlensis, have ribs which are narrower on the central part of the shell and wider and divided by a groove on the anterior and posterior areas of the valve, while in the Porto Rican species the ribs are entire and nearly the same size over the entire shell. From Area oronlensis the fossil differs in its smaller size, more numerous, and probably less beaded ribs. Localities. — Rio Collazo near San Sebastian, Stations, 53, 54, 60. Area (Scapharca) sansebastiana Sheldon and Maury, new species Plate I, Figure 5 Shell small, oblong, rather inflated, with the umbonal ridge ve-y sharply defined; beak full, incurved; hinge-line rather short, its characters concealed by the matrix. The entire surface of the valve is sculptured with about twenty-five, apparently uniform ribs. No beading is shown on the ribs, but this may be due to the imperfect preservation of the shell. Length of right valve 18 mm., height 12 mm., semi-diameter 5 mm. This species is most nearly akin to Area balboai Sheldon (Palaeon- tographica Americana I, p. 69, 1 9 17=A rca dalli Brown and Pilsbry, Proc. Acad. Nat. Sci. Phila., LXIV, p. 510, PL XXIV, fig. 4, January, 1913, name preoccupied), from the Isthmus of Panama. But the Porto Rican shell is smaller, longer in proportion to its height, much less inflated, and with the umbonal ridge much more sharply angled. Tlio ribs also are fewer, numbering twenty against MAURY, TERTIARY MOLLUSCA FROM PORTO RICO 11 twenty-eight. Clearly, however, the affinities of this i*orto Rican species are with the Isthmian shell, which was obtained from a lignitic clay at the bottom of the Culebra Cut, Las Cascades. The new species is founded upon a single specimen of entirely different aspect from Scapharca coUazica, with which it was asso- ciated. Locality. — Rio Collazo near San Sebastian, Station ')'■). Area (Scapharca) cf. actinophora Dall Ai'ca (Scapharca) actinophora Dall, Trans. Wagner Inst. Sci., Ill, pt. 4. p. G47, PI. XXXIII, fig. 2G, 1898. There are in the collection two specimens, probably belonging to the same species, one being the internal mold of both valves with broken posterior margin, the other the worn exterior of the umbonal rea'icn. The latter recalls Area auriculafa Lamarck, but the ribs are wider and the form of the mold is different. The ribs are wider than the interspaces, varying but little over the shell and entire as far as the fragment indicates. This species is most like Scapharca actinophora Dall from Panama, but is smaller and is less attenuated posteriorly. It bears some re- semblance also to A. gatunensis Toula, but is larger, the beaks higher, and more anterior, and the ribbing seemingly was different. This is probably a new species, but too poorly preserved to warrant specific description. Localities. — ISTear Aguada, in the northwest part of the island, Station 117 (internal mold) ; near Ponce, in the south central part of the island. Station 293 (exterior, probably the same). Area (Cunearca) cf. cumanensis Dall ' Area (Cunearca) cumanensis Dall, Trans. Wagner Inst. Sci., Ill, pt. 4, pp. C33, 634, 1898. There is a single right valve of a small Cunearca intermediate between the recent A. incongrua Say and A. chemnitzi Philippi, but nearer the former species. This Porto Rican shell answers the description of Area cumanensis from the Miocene of Cumana, Venezuela, excepting that the posterior ribs near the hinge are nodulose. It is allied also to Area chemnitzoides Maury from Trinidad (Jour. Acad. Nat. Sci. Phila., (2), XV, p. 44, PL VjI, figs. 13, 14, 15, PL VIII, fig. 1, 1912), which is nearer A. chem- 12 SCIENTIFIC SURVEY OF PORTO RICO nitzi. Probably this species is identical with A. cumaneTisis which has never been figured. Locality. — Near Aguada, Station 123. Genus Glycymeris Da Costa Glycymeris acuticostata Sowerby Pectunculus acuticostatus Sowerby, 1850; Quart. Jour. Geol. Soc. London, VI, p. 53, PI. X, fig. 13. Pectunculus acuticostatus Guppy, 18G6, Quart. Jour. Geol. Soc, London, XXII, p. 293; 1867, Proc. Sci. Assoc. Trinidad, p. 164. Axinea acuticostata Gabb, 1873, Trans. Amer. Phil. Soc, XV, p. 255. Pectunculus acuticostatus Guppy, 1874, Geol. Mag. London, p. 443; 1876, Quart. Jour. Geol. Soc. London, XXXII, p. 532. Glycymeris acuticostata Dall, 1898, Trans. Wagner Inst. Sci., Ill, pt. 4, p. 608. Glycymeris acuticostata Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila., p. 364. Glycymeris acuticostata Maury, 1917, Bull. Amer. Paleont., No. 29, p. 180. PI. XXVI, fig. 12; Idem, No. 30, pp. 10, 11, 20, 23. Glycymeris acuticostata Vaughan, 1919, Bull. 103 U. S. Nat. Mus., p. 561. In the collection is a single internal mold probably of Glycymeris acuticostata, because artificial casts made from the interior of a valve of similar size of this species from Santo Domingo show the same angle of slope of the hinge teeth and bear the same general aspect. This species was originally described from the Yaqui Valley, Santo Domingo, where Colonel Heneken first collected it. The Maury expedition, 1916, found it quite common in the Sconsia Icevigata formation of the Mao and Gurabo rivers, Santo Domingo, but did not obtain any specimen from the Aphera formation. The species is also known from Cumana, Bowden and Gatun, but is now for the first time found in Porto Rico. Locality. — Near Quebradillas, Station 130. Glycymeris cf. jamaicensis Dall Pectunculus pennaceus Guppy, 1866, Quart. Jour. Geol. Soc. London, XXII, p. 293. Not of Lamarck. Axinea pennacea Gabb, 1873, Trans. Amer. Phil. Soc, XV, p. 255. Pectunculus decussatus Guppy, 1876, Quart. Jour. Geol. Soc. London, XXXII, p. 532. Not of Hanley. Glycymeris jamaicensis Dall, 1898, Trans. Wagner Inst. Sci., Ill, pt. 4, p. 608; 1903, Idem, III, pt. 6, p. 1586. Glycymeris jamaicensis Maury, 1917, Bull. Amer. Paleont., No. 29, p. 181, PI. XXVI, fig. 13; Idem. No. 30, pp. 24, 27. MAURY, TERTIARY MOLLUSCA FROM PORTO RICO 13 Partly imbedded in the limestone matrix is a Glycymeris which is sculptured with fine, impressed, radiating lines, giving under the lens the effect of obsolete ribs. This sculpture is characteristic of Glycymeris jamaicensis Dall from Bowden, which we also found in the Dominican Miocene, Aphera formation. It is quite probable that the Porto Rican shell is a young specimen of G. jamaicensis but it is too imperfect to identify decisively. The recent analogue of this species is Glycymeris pennacea Lamarck, while the corre- sponding Gatun species is G. carhasina Brown and Pilsbry. Locality. — Near Aguada, Station 117. Genus Pteria Scopoli Pteria inomata (Gabb) Avicula inomata Gabb, 1873, Trans. Amer. Phil. Soc, XV, p. 253. Pteria inomata Dall, 1898, Trans. Wagner Inst. Sci., Ill, pt. 4, p. 669. Pteiia inomata Maury, 1917, Bull. Amer. Paleont., No. 29, p. 181, PI. XXVI, fig. 14; Idem, No. 30, pp. 11, 13. Two internal molds of a Pteria match very well with shells of P. inomata Gabb obtained by the writer on Santo Domingo. This attractive little shell is somewhat variable in form, just as these two molds are not quite alike, but Dominican specimens can be chosen that conform very exactly to each. Well preserved shells are marked with concentric wavy lines of chestnut brown, otherwise the surface is quite smooth. Usually one finds them without the outer layer and then thej justify Gabb's name inomata. The sub- stance of the shell has been entirely dissolved away from the Porto Rican specimens. This is the first time that this species has been reported from outside of the Dominican Republic. The recent Porto Rican shell, Pteria radiata Leach, is larger and is ornamented with radiating rows of scales; but d'Orbigny's Pteria candeana is of much the same general aspect as the fossil. Locality. — Seilor Rabell's Ranch on Rio Guajataca, Station 110. Genus Ostrea (Linnaeus) Lamarck Ostrea coUazica, new species Plate I, Figure 7; Plate II, Figure 1 Shell irregularly oval, valves very dissimilar in form and sculpture. The lower valve is very convex and irregularly ridged and deformed because of the large area of attachment usually in front of and below the beak. Some specimens show internal marginal crenulations anterior and posterior to the 14 SCIENTIFIC 8URTEY OF PORTO RICO ligameniai area. The beak is recurved The convex valve is sculptured with close, somewhat regular, wrinkled flutings, in younger shells about five to a space of ten millimeters, in old valves about three to the same distance. On young valves this radial sculpture is very noticeable, in old valves it becomes more or less obscured but traces can be seen on some areas. Some convex valves are greatly thickened, one measuring 05 mm. along the ligament area. The close, radial, wrinkled flutings, convex, irregu- larly ridged form, and large, deforming attachment area at once distinguish these valves from those of any other species in Porto Rico. The flat valve is marked only with exceedingly close, fine, wrinkled, concentric growth lamella?. Beak very curving, often with a strong spiral twist as in Exogyra. Marginal crenulations are usually in front of and behind the beak. A medium-sized shell with both valves in place measures 70 mm. in height, 55 mm. in width and 40 mm. in thickness. Estimated length of an old valve ICO mm., width 105 mm. Although extremely characteristic of and common in tlie green Collazo shales, this species was found also at one locality (Station 56), in the Arecibo limestone. Localities. — E.io Collazo near San Sebastian, Stations 22, 24, 25, 26, 27, 28, 36, 38, 45, 46, 47, 48, 49, 50, 51, 52, 53, 55, 62, 63, 95, 107; near Lares, Station 56. ISTear Arecibo, Station 452, Dr. Reeds collected an enormously heavy Ostrea, possibly this species. Ostrea berkeyi, new species i'late 1, Figure 6 Shell radially sculptured with smooth, veil definid divaricating, radi:il ribs, divided into two unequal series by a main prin^ary lib. The shell (lifters from Ostrea coUazica in beins; smaller and in jiaving no wrinkled and irregular llutings. In its size and type of sculpture this Ostrea recalls the Claibornian Eocene species, Ostrea divaricaia Lea. Height of sliell 40 mm., greatest width 25 mm., greatest thickness of convex valve l.') nnn. This species is named in li(/n(>r of Professor ('. P. Herkey of Columbia University, in recognition of his valuable contributions to the geology of Porto Kico. Locality. — Rio Cnlla/o near San Sebastian, Station 107. Ostrea sansebastiana, new species Plate I, Figure 1 Shell nearly circular in outline, concave. Beak incurved. Exterior marked by fine, irregularly wrinkled lamellae not unlike those of a flat valve of Ostrea coUczica. Practically enlire internal margin consjiicuously crenu- late. Height of valve (J5 mm., breadth C5 mm., semi-diameter 15 mm. MAURY, TERTIARY MOLLUSCA FROM PORTO RICO 15 I shoidd consider this shell to be a specimen of Ostrca collazica that had through some accident of growth assumed a rounded and concave form', were it not for the strong marginal crenulations which lead me to differentiate it under a special name. Locality. — Rio Collazo near San Sebastian, Station 23. Ostrea cahobasensis Pilsbry and Brown Ostrea cahobasensis Pilsbry and Brown, 1917, Proc. Acad. Nat. Sci. Phila., LXIX, p. 40. PI. VI, fig. 8. Many specimens of a very heavy, thick oyster, of crude aspect, and in nearly every case badly broken, were collected on the south side of the island. The shells were much like the description and figure of Ostrea cahobasensis Pilsbry and Brown, the type of which is from Haiti, in the mountains north of Lake Assuei, on the trail to Las Cahobas, and the species is also found south of Thomonde. In both these Haitian localities the valves of this Ostrea form extensive beds. Because of its very evident importance in correlation. Dr. H. A. Pilsbry by request compared one of the Porto Rican shells with the Haitian type, which is at the Philadelphia Academy, and stated that the Porto Rican shell, "shows no characters differentiating it from Ostrea cahobasensis Pilsbry and Brown. I am disposed to identify it provisionally with that Haitian species ; yet the condition of your specimen does not admit of positive identification." There are, however, among the number some valves precisely like the figure of the type. Dr. Dall also remarked of our shell: "We have fragments of this from the mountains between Haiti and San Domingo." In Porto Rico this oyster is very abundant in and characteristic of the Ponce chalky beds. The affinities of Ostrea cahobasensis seem to be with 0. vaughani Dall, from the Tampa silex beds or the limestone immediately above them. Localities. — N'ear Ponce, Stations 292, 293, 295. Ostrea antignensis Brown Ostrea antignensis Brown, 1913. Proc. Acad. Nat. Sci. Phila., LXV, pp. G03, C14, 615, PI. XIX, fig. 7, PI. XX, figs. 1, 5, G. On the island of Antigua Dr. A. P. Brown found in the hard limestone along the sea shore at Hodge's Bay, High Point and 16 SCIENTIFIC SURVEY OF PORTO RICO Wetherill's Bav, manv isolated ovster shells which he first referred to Ostrea gatunensis Brown and Pilsbrj, but on further examination decided were new and named Ostrea, antigiiensis. On the south side of Porto Rico, east shore of Guanica Harbor, Dr. Reeds collected a quantity of beautifully preserved oyster shells from a shaly for- aminiferal limestone. I was at first disposed to refer these speci- mens provisionally to Ostrea gatunensis, as a few valves bear con- siderable resemblance to the figure of the Gatun oyster, but the majority are broader and of a different aspect, so that I have concluded that the two species are distinct. Dr. Brown's figures of the type of Ostrea antiguensis are of specimens so worn as to be hardly recognizable. Hence a specimen of the Guanica shell was sent to Dr. Pilsbry, who compared it with the type which is at the Philadelphia Academy. In a letter dated July 7th, 1919, Dr. Pilsbry writes: Your oyster is in my opinion, Ostrea antiguensis A. P. Brown. The original specimens are poorly preserved, being in hard rock. In the type there are indications of some ribs additional to the 7 mentioned by Brown. The wide depression, mentioned as characteristic of antiguensis (convex valve), is well marked in your shell, which is far better preserved than the type. I have no doubt of the identity. The relationships of this oyster are with Ostrea haitensis Sowerby from the Miocene of Santo Domingo, 0. gatunensis Brown and Pilsbry from Panama and 0. veatchi Gabb from California. In addition to the strong, radiating, radial ribs of 0. haitensis and 0. veatchi there are upon the ribs of the Guanica oyster spinose flutings, formed by the raised concentric lamella\ The sculpture of the flat valve is similar to that of the convex but is less bold. Localities. — Guanica Harbor, Stations, 368, 369, 370, 371, 373, 374, 375, 376, 381. Genus Pecten Miillor Pecten thetidis Sowerby Pecten irieiiuis sowerby, 1S50, Quart. Jour. Geol. Soc. London, VI, p. 52. Pecten thetidis Gabb, 1873, Trans. Amer. Phil. Soc, XV, p. 2.^6. Pecten thetidis Guppy, 1876, Quart. Jour. Geol. Soc. London. XXXII. p. 532. Pecten (Aequipecten) thetidis Dall, 1898, Trans. Wagner Inst. Sci., Ill, pt. 4, p. 714; 1903, Idem, pt. 6, p. 158G. Pecten (Aequipecten) thetidis Maury. 1917, Bull. Amer. Paleont., No. 29, p. 185, PI. XXXIV, fig. G; Idem, No. 30, p. 27. MAURY, TERTIARY MOLLUSCA FROM PORTO RICO 17 lu the Porto Rican collection are several Pectens which corre- spond to specimens of Fecten thetidis Sowerby that my party col- lected from the type region in the Yaqui Valley, Santo Domingo. Dr. Dall compared our Dominican shells with his Bowden shells, which in turn had been compared with Sowerby's type in the British Museum. This species bears very small spiny scales, visible only under a lens. These decorate the ribs and the radiating threads on the ears. The interspaces also are squamose in older shells. Pecten thetidis has been recorded from Curagao and Jamaica as well as Santo Domingo. Its range is now extended into northern Porto Rico. Locality. — Xear Aguada, Station 122. Pecten inaequalis Sowerby Pecten incequalis Sowerby, 1850, Quart. Jour. Geol. Soc. London, VI, p. 52. Pecten incequalis Guppy, 1866, Quart. Jour. Geol. Soc. London, XXII, p. 294, PI. XVIII, fig. 6; 1867, Proc. Sci. Assoc. Trinidad, p. 164. Pecten incequalis Gabb, 1873, Trans. Amer. Phil. Soc, XV, p. 256. Pecten incequalis Guppy, 1874, Geol. Mag. London, p. 443; 1876, Quart. Jour. Geol. Soc. London, XXXII, p. 532. Pecten inwqualis Dall, 1898, Trans. Wagner Inst. Sci., Ill, pt. 4, p. 714; 1903, Idem, pt. 6, p. 1586. Dr. Dall has identified one of the specimens in this collection as probably Pecten incequalis Sowerby. The shell is incomplete but agrees well with Sowerby's description of the type, which was col- lected by Heneken in the Yaqui Valley, seventy years ago. Guppy later identified the species in Jamaica and figured a shell from that island. Gabb remarked that Pecten incequalis was common in the Cibao (south of the Yaqui River in the northern part of Santo Domingo) and had been found by Bonaczy, one of his assistants, at Loma Cristina on the south side of the island, near San Cristobal. Dall notes that Pecten incequalis is the most common and widely distributed of the Antillean Miocene Pectens. It is recorded, in addition to the Dominican type locality, from Jamaica by Guppy and Bland, from Curagao by the United States Fish Commission, and from the Isthmus of Panama by R. T. Hill. The mollusca associated with this specimen in Porto Rico so strengthen Dall's probable identification as to leave very little doubt that we can now extend the range eastward to Porto Rico. Locality. — ISTear Aguada, Station 124. 18 SCIEXTIFIC SURVEY OF PORTO RICO Pecten guanicus, new species Plate IV, Figure 4 Shell rather small, obliquely fan-shaped; right valve with about nineteen ribs. On the upper part of the valve towards the beaks, the ribs are cari- nated by a single radial thread and the narrow interspaces are smooth. Somewhat further down the single radial thread on each rib becomes accompanied by a second, this by a third and a fourth. About the same time that the tertiary radials appear on the ribs one or two radials develop on the interspaces. Hence on the ventral margin of the shell each rib is ornamented with four radial threads and each interspace with two. An- teriorly the ribs tend to become flatter than on the posterior part of the valve. Approximate altitude 24 mm., breadth 2G mm. The single specimen is partly concealed by the matrix but the characteristics of sculpture are well preseived. In general form and in the sudden development of more ornate sculpture this species resembles Pecten hatoviejonis Maury from the trail between Hato Viejo and Potrero, Santo Domingo, but that shell is decidedly squamose whereas this is entirely without scales. Brown's^ figure 6 of Pecten anguiUensis Guppy bears a slight resemblance to the Porto Rican shell, but his figures 4, 7, 8 of P. anguiUensis are quite unlike our specimen. The Antigua shell like ours has secondary ribbing but the ribs are far broader and much less numerous. The Porto Rican shell is decidedly more like the Dominican species. Dr. Dall has examined this Pecten and he notes that the National Museum has it from Santo Domingo but not named. Locality. — East shore Guanica Harbor, Station 369. Pecten meseticus, now species Plate III, Figure 2 Operculate valve perfectly flat and nearly circular in outline. The ribs number about twenty-five and the perfect ear shows six sharply defined radiating threads. In the umbonal region and upper part of the shell, the ribs are almost linear and very sharply carinated by a radial thread. Less than half way to the ventral margin two more radial threads develop, one on either side of the carinating thread, so that the rib becomes ornamented on top with the three radials. The interspaces are deep set with perpendicu- lar margins and have no radial threads. They are, however, ornamented with sharp, very delicate raised growth lines hardly visible without a lens and as fine as spider webs. These also pass over the ribs but are most apparent in the interspaces. Altitude of shell 35 mm., breadth the same. "Proc. Acad. Nat. Sci. Phil., 1913, PI. XVIII. MAURY, TERTIARY MOLLUSC A FROM PORTO RICO 19 We are much indebted to Dr. Dull for compariii!^' this shell with Pecten soror Gabb. The hitter species has never been figured, Mjut Dall places this Porto Rican shell close to F. soror as identified by Guppy. According to Gabb's description the left valve of his type was concave, but our operculate valve is perfectly flat. The ribs of P. soror are fewer, nor does Gabb mention any radiating threads upon them. Doubtless, however, though not identical, the affinities of this Porto Rican Pecten are with the Dominican P. soror. This is the most exquisite of the Porto Rican Pectens. LocaVity.- — -East shore of Guanica Harbor, Station 375. Pecten camuycencis, new species Plate IV, Figure 5 Shell rather small, subcircular with nearly equal ears. The ribs, omitting the very small anterior and posterior ones, number twenty. Towards the umbonal region the ribs are entire and rather rounded, then about half way towards the ventral margin these on the central part of the valve become medially grooved while those near the anterior and posterior margins remain entire. An exceedingly delicate radial thread develops in the interspaces. It is visible only with a lens and then chiefly because it is accentuated by fine, sharp, V-shaped growth lines, the apices pointing forwards. These growth lines also swing over the ribs and form in the medial grooves U-shaped arches directed slightly backwards. The ears are marked by radial threads, roughened by the fine growth lines which cross over them. Altitude of shell 25 mm., width 2G mm. Dr. Dall has compared this shell with the Pectens in the National Museum and says that it is not represented in the collection there. In its sculpture Pecten camuyceiisis is of the same general type as Pecten gabhi Dall (= paranensis Gabb, 1881, not of d'Orbigiiy, 1849) from Santo Domingo and Antigua, but that shell is larger , and of a much less symmetrical and elegant form than the Porto Rican shell. Locality.- — Xear Arecibo, Station 511. Pecten sansebastianus, new species Plate III, Figure 1 Shell of medium size, oblique, ovate, somewhat inflated. The sculpture consists of about twenty-four ribs which do not extend quite to the anterior and posterior margins. Over the anterior region of the valve each rib bears on either side of its kfel a radial thread, but posteriorly the ribs may 20 SCIENTIFIC SURVEY OF PORTO RICO have four of these radii. Interspaces each with one radial thread. Ears of about the same size. The anterior rather coarsely radially striated, the posterior more finely so. Greatest height of shell, measured obliquely, 35 mm., width 32 mm. Dr. Dall on comparing this Pecten with the collection in the National Musenm says it is not represented there. In sculpture it resembles our Porto Rican species F. guanicus but otherwise is quite different. The type showed no scales or nodules upon the ribs but other specimens show microscopic spines, and it may be that these have been abraded from the type. The species is com- mon and variable. Localities. — Rio Collazo, near San Sebastian, Stations 49, 63. Specimens, probably this species, were also found at nearby Sta- tions 27, 38, 47, 107 and a doubtful shell from 105. Pecten sansebastianus laresianus, new variety Plate IV, Figure 3 Shell with sculpture of the same general plan as that of Pecten sanse- bastianus, but the central keel of the ribs bears minute tear-like nodules. On either side is a radial thread and one also occupies every interspace. The ear is handsomely ornamented with well-defined nodose radii. Altitude 31 mm. This appears to be a variety of Pecteii sansebastianus, but more material may prove it distinct. Locality. — Near Lares, Station 59. Pecten gnajatacus, new species Plate IV, Figures 1, 2 Shell rather large, suborbicular, very inaequivalve; ribs thirteen to sixteen, tending to become obsolete near the anterior and posterior margins of the shell. The convex valve is moderately inflated but not extremely convex. The opposite valve is operculate in form. Altitude of largest speci- men 55 mm., width approximately 53 mm. Dr. Dall after examination notes that these shells are identical with specimens named Pecten heermanni Conrad in the National Museum collection. He did not mention where the latter specimens had been collected, but I presume from the Antillean region. If 80, the label would seem to have been a provisional one, for a com- parison of our specimens with Conrad's description of the type MAURY, TERTIARY MOLLUSC A FROM PORTO RICO 21 indicates that the Antillean and Califoruian fossils are of the same P. eboreus group, but specifically distinct. The true, western P. heermanni has twenty-one square ribs, and, judging from the de- scription, is more compressed than the Porto Rican shell. For these reasons I would suggest that the latter should be differentiated by the name P. guajatacus. Locality. — Senor Rabell's Ranch on Rio Guajataca, Station 113. Genus Amusium Bolten Amusium papyraceum (Gabb) Pleuronectia papyracea Gabb, 1873, Trans. Amer. Phil. Soc, XV, p. 257; 1881, Jour. Acad. Nat. Sci. Phil., (2) VIII, p. 347. Amusium papyraceum Dall, 1898, Trans. Wagner Inst. Sci., Ill, pt. 4, p. 718; 1903, Idem, pt. G, p. 1586. Amusium papyraceum Maury, 1917, Bull. Amer. Paleont, No. 29, p. 190, PI. XXVI, fig. 22; Idem. No. 30, pp. 23, 28. A single fragment of an Ainusium evidently when complete about the size of A. papyraceum is in the Porto Rican collection. The substance of the shell is preserved and the internal surface, showing the characteristically paired ribs of Amusium papyraceum, lies up- ward in the matrix so that the ribs are clearly seen. They are spaced so as to allow two pairs to ten millimeters. Each pair occu- pies about four and each interspace about three millimeters. Gabb's type of this species measured about two inches. The probabilities are very strongly in favor of the fragment being identical with Gabb's shell, since the other Miocene form, Amusium lyoni Gabb has the internal ribs solitary, not paired. Amusium papyraceum was. first described by Gabb from Santo Domingo. It also occurs at Bowden. The shell is very closely related to, and perhaps identical with, Amusium mortoni Ravenel, living in the Antillean and Gulf waters. Locality. — Xear Arecibo, Station 475. Section Propeamusium De Gregorio Propeamusium hollicki, new species Plate III, Figures 3, 4 Shell rather small, suborbicular, thin and semi-transparent, valves dis- similar, the lower gently convex, the upper flat and operculate, ears nearly equal. Outer surface of both valves smooth, but prettily diversified by the 22 SCIENTIFIC SURVEY OF PORTO RICO shining through of the delicate, internal, radiating riblets. Thfse riblets occur in pairs which near the basal margin of the valve measure about one millimeter across, and are separated by interspaces about tv.o and a half millimeters wide. Towards the beak the riblets converge. Each valve has usually eight pairs of riblets which are distributed over the central part of the shell, the anterior and posterior areas being smooth. The most perfect flat valve measures 16 mm. in altitude and 17 mm. in width. A small convex valve is 12 x 12.5 mm., and the largest 20 x 21 mm. This seems to be the maximum size attained, as we have a number of specimens from various localities. No species resembling this has ever been recorded from the Autillean Miocene. It has somewhat the general aspect of Propea- musium pourtalasianum tnarmoratum DalP, a living form but differs among other respects in the lower valve being smooth, not concen- trically sculptured, and the riblets being paired, not club-shaped. The Porto Rican shell is referred tentatively to the section Propea- musium. It appears most closely allied to this group, yet all of the characters are not revealed. According to Dall, the living members of this section are in deep waters in temperate and tropic seas, from 805 to 13 fathoms and at bottom temperatures of 39"' F. to 82° F. which is a very unusual range. This, the most fragile and exquisite member of the genus Amusium yet discovered in the Antillean Mio- cene, is named in honor of Dr. Arthur Holliek. Localities. — Rio Collazo near San Sebastian, Stations, 27, 36, 53, 54, 60, 63, 107. Genus Spondylus Linnaeus Spondylus bostrychites Giippy Spondylus bifrons Sowerby, 1850, Quart. Jour. Geol. Soc. London, VI, p. 53. Not of Goldfuss, 1835 Spondylus bostrychites Guppy, 18C7, Proc. Sci. Assoc. Trinidad, p. 17G. Spondylus bostrychites Gabb, 1873, Trans. Amer. Phil. Soc, XV, p. 257. Spondylus bostrychites Dall, 1898, Trans. Wagner Inst. Sci., III. pt. 4, p. 758; 1903, Idem, pt. 6, p. 1586; 1916, Proc. U. S. Nat. Mus., LI, 1917, p. 493. Spondylus bostrychites Maury, 1917, Bull. Amer. Paleont., No. 29. p. 190, PI. XXXII, fig. 4; Idem. No. 30, pp. 10, 20, 23. Professor Gabb remarked of this shell: "A beautiful convex, almost equivalve Pecteniform species with remarkably narrow areas and a thin shell." This is a welcome addition to Sowcrby's laconic, Latin, three-line description of the type, Spondi/lus ]){fronH, re- «Bull. i7, U. S. Nat. Mus., PI. IV, Fig. i. MAURY, TERTIARY MOLLUSCA FROM PORTO RICO 23 named, because preoccupied, by Guppy. Dall also adds a short description of this Autillean Miocene species in describing his vari- ety chipolanus from Florida. I.ater he referred this to the typical species. In 187-i, Guppy listed Spondylus bosirychites from Jamaica, xlnguilla and Haiti. By Haiti he really meant the Yaqui Valley, Santo Domingo, where Heneken obtained the type. The Porto Rican specimens are nearly all poorly preserved molds but several seem undoubtedly identical with shells my party collected in Santo Domingo and figured as this species. Localities. — Seiior Rabell's Ranch, Station 113; near Aguada, Station 117; near Arecibo, Station 499; near Ciales, Station 500. Spondylus gumanomocon Brown and Pilsbry Spondylus americanus Gabb, 1873, Trans. Amer. Phil. Soc, XV, p. 257. Not (S. americanus Lamarck. Spondylus gnmanomocon Biown and Pilsbry, 1912, Pro?. Acad. Nat. Sci. Phila., p. 514 (Footnote). Spondylus gumanomocon Maury, 1917, Bull. An^.er. Paleont., No. 20, p 191; Id€7n. No. 30, p. 22. This Dominican Miocene species has never been figured, but when adult is easily known by its very large and heavy valves. A speci- men collected by Gabb weighed over thirty-two ounces. Unfortu- nately, one does not always find these ponderous examples and one is perplexed by variations/ of form and growth stages. We have from Porto Rico some internal molds showing that the cavities of the two valves were nearly equal, while others show the typically flatter and the more convex valve, and finally, some of the specimens have the convex valve somewhat twisted. Similar variations of form appear in a series of recent Spondylus americanus in -the American Museum and suggest that the same took place in the Mio- cene member of this group, Spo7idylus gumanomocon. , ,; Localities. — Near Arecibo, Stations 463, 471; near Ciales, Sta- tions 492, 493, 495, 496, 501, 502, 505. Spondylus lucasi, new species Plate V, Figure 1 Shell oval, oblique, small for the genus. The cardinal area is defective and broken, but shows traces of the isodont hinge of Spondylus. The sculpture consists of stronger, low, primary, radiating, rounded threads 24 SCIENTIFIC SURVEY OF PORTO RICO bearing spines at intervals, and between every two of these primaries are eight to ten or twelve much more delicate radiating lines. The central one of these is slightly stronger than the rest. It does not stand out sharply but the eye can discern that it is a shade thicker. The fine radii between the primaries nearly always alternate in strength, the finest lines of all being visible only with a lens. All of the fine radii are ornamented with minute scales, giving them a beaded appearance, especially on the anterior part of the valve. Approximate altitude of shell 33 mm., estimated breadth 28 mm. Among the specimens of Spo)idylus were some fragmentary ex- ternal molds remarkable for the extreme fineness and delicacy of their sculpture which was imprinted in reverse upon the limestone. A single shell shows the same delicate sculpture in the positive; the actual substance of the valves being retained. The exquisite sculpture distinguishes this species from the speci- mens that my party collected in Santo Domingo. Our young shells of Spoiidylus bostrychites have very much coarser and less numer- ous radial ribs between the spinose primaries. The large shell, Spondylus gumobnomocon has the primaries but little differentiated and frequently the spines are no longer than the coarse, imbricating scales which cover all the ribs alike. Dall has lately described Spoiidylus filiaris from the Flint River, Georgia, Oligocene. His figure of the young shell (Proc. U. S. Nat. Mus. LI, PL LXXXIII, fig. 5) bears at first glance quite a resemblance to the Porto Rican fossil, but the Georgian species has no spinose processes, the radii are quite smooth, and the sculpture not so delicate. There is, however, a very beautiful Spondylus in tlio Calcaire grossier of Grignon, France. It is Lamarck's 81. radula. In form it is quite unlike Spondylus lucasi, but its sculpture is so strikingly like as to be almost exactly parallel. The European Eocene shell is more sharply spinose and more prickly. This shell is named in honor of Dr. Frederic A. Lucas. Localities. — Senor Rabell's Ranch, Rio Guajataca, Station 110; East Shore of Guanica Harbor, Station 370; near Arecibo, Station 509. Genus Lithophaga Bolten Lithophaga nigra (d'Orbigiiy) Lithodomus niger d'Orbigny, 1845, In De la Sagra's Hist. Pol. y. Nat. Isla de Cuba, V. pt. 2, p. 351; 1855, Idem, Atlas, VIII, PI. XXVIII, figs. 10. 11. Modiola cariba-a Philippi, 1847, Abbild. u. Beschr., Ill, p. 20, PI. II. fig. ."S. MAURY, TERTIARY MOLLUSCA FROM PORTO RICO 25 Lithophaga nigra Ball, 1898, Trans. Wagner Inst. Sci., Ill, pt. 4, p. 799; 1915. Bull. 90 U. S. Nat. Mus., p. 129. Lithophaga nigra Maury, 1917, Bull. Amer. Paleont., No. 29, p. 194; Idem, No. 30, p. 26. A single specimen of this shell is present in this series. It is fragmentary but unquestionably this species, since it shows tlie entire anterior part of the two valves, which have remained in place, and the characteristic cross striations ending abruptly along an oblique line. In size it corresponds almost perfectly to d'Orbigny's figure of the type which was a recent specimen. The earliest recorded appearance of this species is in the Upper Oligocene, Dall having identified it in the Tampa formation. My party found it in the Lower Miocene of the Yaqui Valley, Santo Domingo and it is now living from Florida to St. Thomas, West Indies. The fossil shells in all respects except color are like the recent specimens. Locality. — Near Lares, Station 56. Order Ais^omalodesmacea Genus Thracia Blainville Subgenus Cyathodonta Conrad Cyathodonta reedsi, new species Plate V, Figure 2 Shell rather elongate, beaks subcentral, right valve gently convex, with its anterior end somewhat produced and rounded, posterior end broad, abruptly truncated; posterior dorsal area flattened, nearly smooth and marked off by a rounded carina passing from the beak to the base of the truncation; dorsal line in front of the beak very sloping, but behind nearly straight; beak low, acute, curving over the hinge and evidently in life very close to, or touching, the opposite beak, its apex appears perforated but this may be due to an accident and not to contact with the opposite beak. Surface of valve anterior to the carina ornamented with close-set, nearly uniform, slightly wavy, concentric ripples, about four to every five milli- meters These ripples near the center of the disc are often broken off and then continued as though unskillfully spliced, and sometimes they anastomose with adjacent ripples. Nevertheless, their appearance is rather regular and elegant. In front of the carina is a shallow broad, inconspicuous sulca- tion. Hinge entirely concealed by the limestone matrix. Length of shell 32 mm., altitude 20 mm., thickness of one valve 7.5 mm. The genus Thracia has never before been recorded from the Mio- cene of the Greater Antilles. Heneken, Guppy, Gabb and the writer 26 SCIENTIFIC SURVEY OF PORTO RICO all failed to find a Thracia in Santo Domingo or Jamaica, nor does Dall list this genus in the Henderson-Simpson collection from the Bowden beds of Jamaica. Dr. J. W. Spencer, however, collected two species from the Lesser Antilles, both on the Island of Guade- loupe. These were described by Dall as Cyathodonta guadalupensis and C. spenceri, and were referred to an Oligocene horizon. Sinffularlv enoueh, the Porto Rican Thracia is not like either of the Guadeloupe species, but resembles far more the older shell, Cyathodonta vickshurgiana Dall, from the Oligocene of Vicksburg, Mississippi. In general aspect it corresponds strikingly with C. vickshurgiana, but is proportionally higher, more convex, with beaks subcentral, instead of at the posterior third, and the ripples are less crowded, there being four instead of five to every five millimeters. To a less extent our fossil recalls Toula's Isthmian Cyathodonta gatunensis, which among other respects differs in its broader an- terior end, less elongate form, and its crude aspect. This rare and more delicate shell is dedicated to Dr. Chester A. Reeds, by whom it was discovered. Localities. — Near Quebradillas, Stations 126, 129, 131; near Ponce, Station 293. Genus Cuspidaria Nardo Cuspidaria islahispaniolse Mamy Plate V, Figure 3 HJea-ra aUernata Gabb, 1873. Trans. Amer. Phil. Soc. XV.. p. 248. Not Sphena alternata d'Orbigny. 1845. Newra ornatissima Guppy. 1876. (in part), Quart. Jour. Geol. Soc. London, XXXII, p. 530. Not Sphena ornatissima d'Orbigny, 1845. Cuspidaria islahispaniohr Maury. April 29, 1917. Bull. Amer. Paleont., No. 29, p. 196, PI. XXVI. fig. 20; Idem. No. 30. p. 13. Cuspidaria gabbi .Johnson and Pilsbry. May 5, 1917, Proc. Acad. Nat. Sci. Phil , LXIX, p. 195. A specimen in the collection matches so perfectly with shells of my Dominican Cuspidaria islahispaniota from Bluff 3, Cereado de Mao, that there seems to be no doubt of their complete identity. Gabb and Guppy both referred the Miocene Cuspidarias from Santo Domingo to the recent species C. alternata and C. ornatissima of d'Orbigny. These they considered identical with each other and with the fossils. MAURY, TERTIARY M0LLU8CA FROM PORTO RICO 27 Johnson and Pilsbry's revision of the Gabb collection in the Phila- delphia Academy differentiated two species of Cuspidaria as C. ornatior (like ornatissima) and C. gahbi (like alternata). This harmonizes with Gabb's remark in 187H that he had specimens grading into the two species. Johnson and Pilsbry nnfortunately gave no figures, but judging from their description, C. gahbi is synonymous with C. islahispaniohv. We have no representatives of C. ornatior. Locality. — Near Juana Diaz, Station 178. Cuspidaria juanadiaza, new species Plate V, Figure 4 Shell small, plump, with full beaks and a long rostrum. Surface marked by rather coarse concentric growth lines as in some small Ledas. Exfoliated over central part of valve and under layers cracked and veined, but along the base and at the anterior and posterior ends the outer layer is preserved and shows no radial ribbing whatever. There is only a weak radial thread defining the rostrum. Hinge characters concealed. Length of shell 10 mm., height 6 mm. This species is about the same size as Cuspidaria islahispaniolce, but is at once differentiated by the absence of ali radial ribbing. This feature also separates it from Dall's two Bowden species C. craspedonia and C. distira and also from ('. costellata Deshayes. It is evidently a shell of the same type as the recent Cuspidaria laniel- losa Sars, which lives in fifty to five hundred and fifty-two fathoms off the Norwegian and Khode Island coasts. This is the first Cus- pidarian of this group ever found in the Antillean Miocene. Locality. — Near Jauna Diaz, Station 176. Okder Teleodesmacea Genus CrassateUites Kriiger CrassateUites juanadiazus, new species Plate IV, Fig-ures 6, 7 Shell elongated, anterior end rounded, posterior end produced and bluntly pointed. The substance of the shell was thin and this has been increased by exfoliation. The original form has been altered by pressure which apparently has greatly increased the inequality of the valves giving tne shell a CorbwJa-like form, with one valve very gibbous and incurving 28 SCIENTIFIC SURVEY OF PORTO RICO at its base, and the other valve flattened and seeming to fit into the larger. This appears to be due to distortion. On the convex valve a pronounced carina passes from the beak to the posterior angle; beak anterior to the center; nepionic shell, as far as can be seen, with very close, sharp, con- centric ridges. With age these become progressively rounder and somewhat more distant and ornament the entire shell. Hinge concealed by the matrix. Length of shell 28 mm., height 16 mm., thickness of convex valve 7 mm. Gabb's Crassatellites reevei from the Miocene of Santo Domingo is a very much larger shell, fifty-six millimeters in length, and more coarsely sculptured, I do not think this Porto Rican shell could be the young of that species as it has a very much more delicate and different aspect and appears adult. The Bowden species, Cras- satellites jamcuicensis Dall, is wholly unlike this fossil in both form and sculpture. The convex valve of the Porto Rican shell is, how- ever, strikingly like Crassatellites flexurus (Conrad) from the Jack- sonian Eocene of Jackson, Mississippi, but the latter shows no noticeable inequality of its valves, while the Porto Rican shell even before distortion would seem to have had discrepant valves. A single specimen only was found. Locality. — i^ear Juana Diaz, Station 182. Genus Venericardia Lamarck Venericardia collazica, new species Plate VI, Figure 6 Shell elongate, very inequilateral, much inflated in the umbonal region, beaks very near the anterior end, very high, acute, curving over the hinge line. Ribs thirteen or fourteen showing traces on the mold of crenulations. The ribs are simple without secondary threads nor are there Interstitial riblets in the interspaces. The interspaces are about as wide as or a little narrower than the ribs which are broadest and more distant on the central portion of the valves and become suddenly finer and closer on the posterior dorsal areas. Length of shell approximately 34 mm., height 20 mm., di- ameter 21 mm. In general form this species resembles Veyiericardia nc^su^a 'Dall from the Eocene of Alabama (Trans. AVagner Inst. Sci., Ill, pt. 6, p. 1425, PL IJTI, fig. 9, 1903), but that shell has lateral threads accompanying the anterior and medial ribs, and the ribs number twenty-five against fourteen in the Porto Rican shell. The species is founded on a single internal mold from the CoUazo shales, still retaining in places a coating of the shell substance, which shows MAURY, TERTIARY M0LLU8CA FROM PORTO RICO 29 both valves together of a shell resembling Carditamera in its elon- gated form, but with the prominent, prosogyrate beaks characteristic of V enericardia. Locality. — Rio Collazo near San Sebastian, Station 53. Venericardia rabelli, new species Plate VI, Figure 7 Shell rather small, sub-circular with low, acute beaks, strongly prosogy- rate. Surface ornamented with many close-set ribs, numbering about twenty- one, and separated by narrower interspaces. The ribs of the anterior and central portion of the valve develop on either side of their central keels, fine threadlike secondary riblets which are just visible without a lens. On the posterior part of the valve these secondary threads become weaker and finally disappear. Traces yet remain of nodules on the anterior ribs. The char- acters of the hinge are concealed by the matrix. Length of shell 20 mm., height 17 mm., semi-diameter 10.5 mm. This pretty little Venericardia recalls in some respects Y. cer- rogordensis Maury, from Santo Domingo. That shell is of similar form and size, and its anterior and central ribs also have two ac- companying lateral, threadlike riblets, but there are only seventeen ribs in the Dominican species and they are far more distantly spaced. This Venericardia is dedicated to Seiior Narcisso Rabell of San Sebastian. Locality. — Weathered green shale, which has become reddish, in a roadside cut, Rio Collazo near San Sebastian, Station 62. Also a fragment, probably this same species from the nearby Station 54. Venericardia juncalensis, new species Plate VI, Figure 5 Shell when complete ovate-cordate in form moderately convex, com- pressed by pressure to which the Collazo shales were subjected. Substance rather thin; beaks low, acute; ribs fourteen when perfect strongly nodular. All the ribs except those of the posterior area are accompanied on either side by a radiating thread; at the base of the shell a medial rib with its two radiating threads measures about three millimeters across. Length of shell approximately 30 mm., height 29 mm., diameter of crushed shell 11 mm. Although this species has the same general type of sculpture as the preceding shell and as V. cerrogordensis Maury, yet its aspect is unlike those two species. Locality. — Rio Collazo near San Sebastian, region of Juncal, Station 94. 30 SCIENTIFIC SURVEY OF PORTO RICO Genus Echinochama Fischer Echinochama antiquata Dall Chama arcinella Moore, 1853, Quart. Jour. Geol. Soc. London, IX, p. I'M). Not C. arcinella Linnaeus, 1767. Chama arcinella Guppy, 18G6, Quart. Jour. Geol. Soc. London, XXH, p. 294: 1867, Proc. Sci. Assoc. Trinidad, p. 163. Chama arcinella Gabb, 1873, Trans. Amer. Phil. Soc. XV, p. 251. Chama arcinella Guppy, 1874, Geol. Mag. London, p. 442; 1870, Quart. Jour. Geol. Soc. London, XXXII, p. 531. Echinochama antiquata Dall, 1903, Trans. Wagner Inst. Sci., III. pt. 6, pp. 1404, 1586, PI. LIV, fig. 9. Echinochama antiquata Maury, 1917, Bull. Amer. Paleont., No. 29, p. 201, PI. XXXIII, fig. 10; Idem,, No. 30, p. 10. Echinochama antiquata Jones, 1918, Jour. Geol., XXVI, p. 738. This Miocene species was recorded from the Yaqiii ^ alley, Sauto Domingo and the Bowden beds of Jamaica by Guppy and Gabb as the recent shell Chama arcinella which has much more prominent and spiny ribs. Dall first discriminated the fossil and referred it to Fischer's genus Echinochama. The closest recent ally of E china- chama antiquata is a Pacific shell and not Echinochama arcinella which Dall and Simpson record from Mayaguez Harbor, Porto Rico and is abundant elsewhere in the Antilles. The type of Echinochama antiquata is a Bowden shell. My party collected fine specimens from Cercado de Mao, Santo Domingo, and Jones has now found it in the Las Cahobas beds, south of Thomonde, Haiti. From Porto Rico Reeds collected a single internal iiuild which fits very perfectly into a fine shell of Echinochama antiquata in this Museum collected by F. C. Nicholas from Bowden. The only other Echinochama which it might be is E. yaquensis Maury, but that tends to be smaller. It is almost certainly the internal mold of Echinochama antiquata. Locality. — Near Arecibo, Station 445. Goniis Lucina Binguioro Lucina chrysostoma (Mcusc'lioii) iMiili})])! Tellina crysustoma Meuschen, 1787, Mus. Gevers., p. 482, (Typographical error). Lucina chrysostoma Philippi, 1847, Abbh. und Beschr. Conchy.. II, p. 206, PI. I, fig. 3. MAURY, TERTIARY MOLLUSCA FROM PORTO RICO 31 Venus edentula Chemnitz, 1784, Conch. Cab., VII, PL XL, figs. 427-429. Not of Linnaeus, 1758. Lucina edentula Reeve, 1850, Conch. Icon., Lucina, PI. II, fig. 9. Lucina edentula Heilprin, 18SG, Trans. Wagner Inst. Sci., I, pp. 102-103. Lucina chrysostoma Dall, 1903, Trans. Wagner Inst. Sci., Ill, pt. 6, p. 1354. Lucina chrysostoma Maury, 1917, Bull. Amer. Paleont., No. 29, p. 202, PI. XXXV, fig. 2; Idem. No. 30, pp. 13, 26. Lucina chrysostoma Jones, 1918, Jour. Geol. XXVI, pp. 740, 742. This is one of the most abundant species in the Porto Rican collection. Among a great number of internal molds which show the plump form, elongated adductor scar and variable radial lines characteristic of the interior of shells of Lucina chrysostoma, are several very perfect impressions of the exterior of the shell, showing also the edentulous hinge. Artificial casts of these when compared with recent and Pleistocene specimens of Lucina chrysostoma showed no points of difference. This is the apricot shell of col; lectors, — so named because the interior of living shells has a fine apricot blush. Dall does not record this species from beds earlier than Pliocene; but it seems that it extended into the Miocene since both these Porto Rican specimens and mine from the Yaqui Valley, Santo Domingo, appear in all respects like the recent. Jones also reports it from the Maissade beds of Haiti. Localities. — Xear Juana Diaz, Station 174; near Ponce, Stations 285, 287, 288, 289, 290, 291, 292, 293, 297, 298, 299, 301; near Guavanilla, Station 320. '■') Genus Phacoides Blainville There are several varied forms of this genus in the collection, but nearly all are internal molds, upon which no specific determinations can be based. Subgenus Here Gabb Phacoides (Here) quebradillicus, new species Plate \a. Figure 8 Shell moderate in size, convex, with the posterior and anterior dorsal areas very sharply marked off by narrow, deep sulci. The beak is full, acute and strongly turned forwards. The entire surface of the valve is sculptured with very close-set, nearly uniform and equidistant concentric ridges, so fine as to be hardly visible without a lens. Characters of the hinge unknown. Length of shell 24 mm., approximate altitude 21 mm., semi-diameter 7 mm. 32 SCIEXTIFIC SURVEY OF PORTO RICO This species is much less convex than the subglobular Bowdcn shell, Phacoides podagrinus Dall, which is also a member of the Pliocene to recent, P. peniisylv aniens group. Phacoides domin- gensis Dall is flatter with less fine and regular, concentric sculpture, and is of the Phacoides pectinatus stock. The Porto Rican shell, which is represented bv a single, nearly perfect external mold of the left valve, is so finely sculptured that the cast of its imprint appears almost smooth to the eye. The delicate, nearly regular, concentric ridges stand out only with a lens. In its general aspect this species resembles Phacoides (Here) wacissanus Dall, from the Tampa silex beds more than any with which I am acquainted. Locality. — Xear Quebradillas, Station 126. Subgenus Lucinisca Dall Lucinisca hoveyi, new species Plate VII, Figure 2 Shell small, rather plump, suborbicular, with an acute, strongly prosogy- rate beak. Valve very beautifully latticed by the intersection of fine, sharp, close-set concentric ridges, with the somewhat stronger radial ribs. Ribs thread-like, nearly equidistant and about uniform in strength with no secondary riblets intervening. Anterior dorsal area marked by a change in sculpture as the ribs are obsolete and only the concentric ridges are present. Posterior dorsal area defined by a slight umbonal ridge beyond which the area is a little compressed. Hinge characters unknown, but the shell is referred tentatively to the subgenus Lucinisca because of its form and sculpture. Length of most perfect external mold 11 mm., height 10.5 mm., semi-diameter 4.5 mm. This species is at once differentiated from the Santo Domingan Miocene shells, Lucinisca liispaniolamv Maury and L. cercadica Maury, by its lack of secondary riblets. It is closer to Dall's Floridian Lucinisca calhounensis from the Tampa and Chipola horizons, for that has all the ribs subequal ; but the Porto Rican shell is less compressed and differs in form, being plumper and more cordate. The external molds retain every detail of sculpture in reverse in the limestone rock, and show that this shell was one of the most exquisite on the Miocene beaches of Porto Rico. It is dedicated to Dr. E. O. Hovey. LocaZi^tes.— Near. Quebradillas, Stations 126, 129, 131, 133, 136; near Ponce, Station 292. MAURY. TERTIARY MOLLUSCA FROM PORTO RICO 33 Subgenus Miltha H. and A. Adams Miltha cf. smithwoodwardi Maury Phacoides (Miltha) smithwoodwardi Maury, April, 1917, Bull. Amer. Paleont., No. 29, p. 204, PI. XXXV, fig. G. As in the Dominican Miocene waters, so in the Porto Rican, two species of Miltha were represented. Unfortunately, the collection affords only internal molds of the Porto Rican shells. One of these molds has the very striking subquadrilateral and flattened form of Miltha smithwoodwardi Maury, from Bluff 2, Cercado de Mao, Santo Domingo. The mold is somewhat larger, measuring 53 mm. in length, 60 mm. in height and 18 mm. in di- ameter, while the corresponding measurements of the type were 45 X 50 X 12 mm. Nevertheless it was clearly either identical with or very closely akin to Miltha smithwoodwardi, for the striking posterior truncation, and the angles of the ventral margin are ex- actly parallel to the outlines of the Dominican shell. The beak also was markedly prosogyrate. The main difference in form is that the anterior dorsal margin of the mold is sloping and not angu- lated as in the Dominican shell. This species is represented in the living fauna by Miltha childreni Gray, in the Gulf of California. The only other living Miltha is found off Mozambique. The sub- genus was at its maximum during the Eocene and has since de- clined. Dall calls attention to the interesting fact that in most horizons down to the Pliocene, species representing three groups Qccur — namely the group of M. hillsboroensis Heilprin, that of M. childreni Gray, and that of M. pandatus Conrad. The first two groups are represented in the Porto Rican Miocene. Localities. — I^ear Quebradillas, Station 137. A smaller, broken internal mold, probably the same species, from Senor Rabell's Ranch, Station 112. Miltha cf. hillsboroensis (Heilprin) Lucina hillsloroensis Heilprin, 1887, Trans. Wagner Inst. Sci., I, pp. 117, 120, PI. XVI, fig. 62. The other internal mold of Miltha under consideration has the general form of the Floridian species M. hillsboroensis Heilprin from the Oligocene and Miocene, and of the related Pliocene shell, M. disciformis Heilprin. It also resembles the Santo Domingan mem- 34 SCIENTIFIC SURVEY OF PORTO RICO ber of this group, Miltha riocane7isis Maury. Of all these, the mold corresponds in size best with M. disciformis, but that is a much later species. As the surface characters of the Porto Rican shell are quite unknown no specific identification is possible. We know, however, that a member of the M. hillsboroensis group was living in the Miocene of Porto Rico. Localities. — Near Ponce, Station 289. Also a larger, fragmen- tary mold 60 mm. in width, probably the same species, from Sta- tion 292. Genus Divaricella von Martens Divaricella prevaricata Guppy Divaricella prevaricata Guppy. 1896, Pioc. U. S. Nat. Mus., XIX (1897), No. 1110, p. 327, PI. XXX. fig. 6. Divaricella prevaricata Dall. 1903. Trans. Wagner Inst. Sci., Ill, pt. G, pp. 1389, 1587. Divaricella prevaricata Maury, 1917, Bull. Amer. Paleont., No. 29, p. 20 1. PI. XXXV, fig. 10; Idem, No. 30, pp. 13, 24, 28. The type of this Miocene species was described by Guppy from Bowden, Jamaica. My party collected shells corresponding to Guppy's description and fig-ure in the Miocene of Santo Domingo, in the Apfiera islacolonis formation in the bluffs of the Rios Mao, Gurabo and Cana. In the present collection there is a single, but very perfect, imprint of this shell in the limestone so that the species has now been found on three of the Greater Antilles. Divari- cella prevaricata is smaller and more delicately sculptured than the recent representative, Divaricella quadrisulcata d'Orbigny, recorded by Dall and Simpson from Hucares, Porto Rico. Locality. — Near Quebradillas, Station 125. Genus Cardium (Linnaeus) Brugiiiere This genus is represented in the collection by a considerable number of specimens, but nearly all in the form of poorly preserved molds. Probably several correspond to Jamaican and Santo Do- mingan Miocene species. Section Trigoniocardia Dall Cardium (Trig^oniocardia) sancti-sebastiani, new species Plate VI, Figure 4 Shell sub-trigonal, e.xceedingly oblique, anterior end very short, posterior end produced and roundly pointed; beak very anteriorly placed, high, acute. MAURY, TERTIARY MOLLUSC A FROM PORTO RICO 35 curving forwards; posterior slope from the iimbonal region to the posterior basal margin not carinated, but gently and evenly rounded, so that the posterior area is not v, ell defined. On the body are ten radial ribs which show traces of having been crenulated, on the posterior area are five ribs closer, and finer than those on the body of the shell. Length 12 mm., altitude from beak to anterior basal angle 6 mm., semi-diameter 5 mm. The Antillean Tertiary has yielded the following Cardiiims of the section Trigoniocardia: C. castum Guppy (Quart. Jour. Geol. Soc. Lond, XXII, pi. XXVI, tig. 4, 1866) from the older Tertiary of Manzanilla, Trinidad, a small subovate shell with twenty-two ribs and beak nearly central ; C. haitense Sowerby, from the Miocene of Santo Domingo and Bowden and its more oblique, more strongly carinated variety, C. cercadica Maury, both have fourteen ribs -on the body and ten on the truncation ; C. aminense Dall from Rio Amina, Santo Domingo, is a strongly carinate, very convex shell, with ten ribs on the body and ten also on the truncation, and C. maturense Dall from the later Tertiary of Matura, Trinidad, is not carinate, but is only slightly oblique and has twelve body and eight posterior ribs. The Porto Rican shell is clearly specifically distinct from any of these forms. Locality. — Rio CoUazo near San Sebastian, Station 53. Cardium (Trigoniocardia) haitense Sowerby Cardium haitense Sowerby, 1850, Quart. Jour. Geol. Soc. London, VI, p. 52, PI. 10, fig. 11. Cardium haitense Guppy, 186G. Quart. Jour. Geol. Soc. London, XXII, p. 293; 1867, Proc. Sci. Assoc. Trinidad, p. 163. Cardium (Fragum) haitense Gabb, 1873, Trans. Amer. Phil. Soc, XV, p. 251. Cardium haitense Guppy, 1874, Geol. Mag. London, p. 442; 187G, Quart. Jour. Geol. Soc. London, XXXII, p. 531. Cardium (Trigoniocardia) haitense Dall, 1900, Trans. Wagner Inst., Sci., Ill, pt. 5, pp. 1103, 1105. Cardium {Trigoniocardia) haitense Maury, 1917, Bull. Amer. Paleont., No. 29, p. 211, PI. XXXVI, figs. 5, 5a. In 1917 I noted that Sowerby's figures of the type of Cardium haitense included two forms; the upper representing a subquadrate shell with roundly angulated umbonal slope, the lower a trigonal shell, posteriorly produced with a sharp umbonal ridge. As the description fitted the first figure I suggested, restricting the name haiteiise to that form and diiferentiating the second by the varietal name cercadicum. An external mold in this collection shows the same form and number of ribs as American Museum specimens of 36 SCIENTIFIC SURVEY OF PORTO RICO C. haitense from Bowden, Jamaica, and mine from the Yaqiii Valley, Santo Domingo. As long ago as 1867, Guppy recorded this shell from Jamaica and Cumana, Venezuela, aa well as from the type locality which was the Yaqui Valley. To this distribution we may now add Porto Rico. Localities. — ISTear Quebradillas, Station 133 ; near Arecibo, Sta- tion 476. Specimen from Station 476 poor and identification un- certain. Cardium (Trigoniocardia) haitense cercadicum Mauiy Cardium (Trigoniocardia) haitense variety cercadicum Maury, 1917, Bull. Amer. Paleont., No. 29, p. 212, PI. XXXVI, fig. 6; Idem, No. 30, p. 11. Several internal molds of what appear to be this variety are in the Porto Rican collection. Traces of about fourteen ribs on the body and of about ten on the truncation still remain. The umbonal ridge is sharply angulated and the outline of the shell is trigonal. Localities. — Near Quebradillas, Station 131 ; near Ponce, Sta- tions 288, 289, 291. Subgenus Laevicardium Swainson Cardium (Laevicardium) serratum Linnaeus Cardium serratum Linnaeus, 175S, Systema Naturae, ed. X, p. 680; 1767, ed. XII, p. 1123. Cardium Icvvigatum Lamarck, 1819, Anim. sans Vert., VI. pt. 1, p. 11. Not of Born, 1780, nor of Linnagus 1758. Cardium { La-vicar dium) serratum Dall, 1900, Trans. Wagner Inst. Sci., Ill, pt. 5, p. 1110; 1903, Idem, pt. G, p. 1587. Cardium (Lwvicardium) serratum Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila., p 3C7. Cardium (Lwvicardium) serratum Maury, 1917, Bull. Amer. Paleont., No. 29, p. 212, PI. XXXVI, fig. 8; Idern. No. 30, pp. 10, 22, 23, 24. Cardium {La;vicardium) serratum Vaughan, 1919, Bull. U. S. Nat. Mus. No. 103, p. 5C2. My party found in the Miocene Bluifs of the Gurabo and Mao Rivers, Santo Domingo, fossil shells of Lcevicardium serratum pre- cisely like recent specimens on the beach at Monte Cristi. Dall records this species from Bowden, J amaica, and Alum Blulf , Florida ; the Pliocene of Florida and South Carolina ; Pleistocene of Florida and the Antilles, and living from Cape Hatteras to Bahia, Brazil. MAURY, TERTIARY MOLLUSCA FROM PORTO RICO 37 Among the molds of Cardium from Porto Rico collected by Reeds are some which seem undoubtedly to represent this species. We mav thus extend its range eastward to Porto Rico. Localities.— ^ ear Quebradillas, Stations 126, 131; near Ponce, Station 291. Genus dementia Gray This tropical genus is now reduced to some half dozen species living chiefly on the Philippine, Korean, and Australian coasts, the genotype being Clementia yayyracea Gray. A single authentic re- cent species C. solida Dall, is on the West coast of Mexico. It is the only living member of the genus in American waters. During the Tertiary, however, Cleme^itia lived on the East coast of America. Professor G. D. Harris has lately referred Dosinia mercenaroidea Aldrich, from the St. Maurice Eocene beds at the base of the Clai- borne bluff, to Clementia. During the Miocene Clementia inoceri- formis inhabited the Maryland shores and probably ranged north to Martha's Vineyard, Mass., since a mold like it has been found in the Gay Head Miocene clays. Clementia grayi Dall lived during the deposition of the Oak Grove sands, Florida, and C. tceniosa Guppy has been preserved in the Savaneta beds of Trinidad Island. The isthmian species was C. dariena of which we have now found an ancestral form in the Middle Oligocene of Porto Rico. This genus has never before been recorded from the Greater Antilles. The Trinidad species, C. twniosa is subtri angular with the beaks nearlv central. Clementia rabelli, new species Plate VI, Figures 2, 3 Shell closely resembling Clementia dariena Conrad, but far less ventricose and smaller than this later, Isthmian species. Original substances of shell very delicate and papery, handsomely sculptured with wave-like, concentric, rounded ridges, resembling the undulating sculpture of the modern genus Raiita and its Eocene precursor, Pteropsis. Older shells of the Clementia lose this sculpture and near their ventral margins show only close, con centric growth lines just as in aged specimens of C. dariena. Dr. Reeds collected many molds of this common and very char- acteristic fossil of the Collazo shales. The molds are greatly dc^ formed bv disturbances of the strata in which they were deposited. 38 SCIENTIFIC SURVEY OF PORTO RICO Many are flattened, while in others the pressure has acted on the anterior-posterior axis, fracturing the shell medially and compress- ing it into an oblong triangular form with a false, superficial aspect of Inoceramus. Measurements of various shells are as follows: Oval specimens — altitude 40 mm., length 45 mm., diameter 15 mm. ; 33 X 38 X 12 ; 34 x 45 mm. Triangular compressed shell — 38 x 24 X 20 mm. This Clementia is an ancestral Oligocene precursor of the Miocene Gatun C. dariena. The affinity is so close that, had there been a less distant stratigraphic relationship, I would have called the Porto Ricau shell a variety of the Gatun. But the Porto Rican Clementia, according to Dr. Reeds, underlies the Orthaulax zone. Apropos of this Vaughan's recent listing of an unidentified Clementia in the Culebra formation at Panama is interesting. This shell would be far more nearly synchronous with the Porto Rican Clementia. Of recent shells, Clementia rabelli is most like specimens in the American Museum of C. papyracea from ISTew Holland. This specie? is dedicated to Seiior Xarcisso Rabell Cabrero, the scientist and natu- ralist of San Sebastian, w^ho most generously showed his collections to Dr. Reeds and pointed out the rich fossiliferous localities along Rio Collazo. The Clemeniia and its associated fauna were obtained from the green shales outcropping on the Collazo in the vicinity of the beautiful waterfalls on Sefior Rabell's property. Localities. — Rio Collazo near San Sebastian, Stations 23, 53, 54, 60, 61, 62. Genus Antigona Sclminacher Antigona tarquinia Dall Venus magnifica Heilprin, 188G. Trans. Wagner Inst. Sci.. I, p. IIG. Not V. magnifica Sowerby, 1S53. Venus tarquinia Dall. 1900, Trans. Wagner Inst. Sci., Ill, pt. 5, p. 1104. PL XXXVIII, figs. 2, 2a. Figure only. Cytherea tarquinia Dall, 1903, Trans. Wagner Inst. Sci., Ill, pt. 6, p. 1274. Antigona tarquinia Dall, 1915, Bull. 90, U. S. Nat, Mus., p. 147, PI. XXVI, figs. 1, 2. From the Collazo shales come two imperfect specimens which have the striking and very characteristic sculpture of Aniicjona tarquinia, described by Dall from the Tampa formation, and listed by him also from Santo Domingo. The species is remarkable for its very handsome sculpture which is Corhis-Wk^, and consists of MAURY, TERTIARY MOLLUSCA FROM PORTO RICO 39 imiform, close-set, radial threads interrupted by regularly-spaced, concentric ridges. One of the shells has been crushed into an eliptical form by pressure acting on the dorsal region. This enhances its false re- semblance to a Corhis. The other is not distorted but is broken at the ends. Yet the shell shows a decided likeness in form, sculpture, and size to the Floridian type and the posterior dorsal margin also is nearly straight. The altitude is -iO mm. and diameter 25 mm., the corresponding measurements of the type were 41 x 26 mm. Localities. — Rio Collazo near San Sebastian, Station 46. A crushed shell, probably this species from Station 63, same locality. From the limestone near Aguada, Station 117, is an internal mold, retaining traces of sculpture, which may be this species. Antigona tarquinia antillica, new variety Antigona tarquinia Maury, 1917, Bull. Amer. Paleont., No. 29, p. 217, PI. XXXVII, fig. 4; Idern, No. 30, p. 27. Not A. tarquinia Ball, 1900. Shell of medium size, roundly trigonal, with the posterior doisal margin sloping rapidly; beaks low; surface sculptured with narrow, concentric ridges between which are conspicuous radial threads. Length of shell 3G mm., altitude 33 mm., semi-diameter 10 mm. Collected by my party at Zone H, Rio Cana, Santo Domingo, in 191G. In describing this Dominican shell I noted that more material might prove it w^orthy of varietal rank. The Porto Rican collection made by Dr. Reeds includes a single external mold in the limestone which strikingly resembles the Do- minican valve. The umbo is broken away, but the mold agrees in its sloping posterior margin and in size and general aspect with the valve from the neighboring island. This form appears to constitute a variety. Locality. — ^ear Quebradillas, Station 130. Antigona, species indet. A very imperfectly preserved shell shows that a large Antigona was present resembling somewhat in form and sculpture A. willcoxi Dall from the Florida Pliocene, but smaller. In size the shell re- sembled the Miocene A. caesarina Dall from the Chipola marls, but in outline it is nearer A. willcoxi. Unfortunately it is too poorly preserved to describe. .-•'~'' Locality. — Xear Ciales, Station 504. ■<- 40 SCIENTIFIC SURVEY OF PORTO RICO Genus Callocardia A. Adams Callocardia riocoUazica, new species Plate VI, Figure 9 Shell rather small, cordate, moderately convex, beaks rather low and curving forward over the small lunule. Hinge characters concealed by the matrix, hence the reference of the shell to this genus is tentative. Surface sculpture of very fine, even ridges barely seen without a lens. Length of shell 28 mm., altitude 20 mm., semi-diameter 8 mm. Callocardia gatunensis Dall from the Miocene of the Isthmus and of Santo Domingo, is larger and the curve of the anterior basal margin is quite unlike that of the Porto Rican shell. Locality. — Rio Collazo near San Sebastian, Station 53. Genus Chione Mergele von Miihlfeldt Chione woodward! (Guppy) Venus ivoodwardi Guppy, 1866, Quart. Jour. Geol. Soc. London, XXII, p. 292, PI. XVIII, fig. 1; 1867, Proc. Sci. Assoc. Trinidad, p. 162. Chione u-oodwardi Gabb, 1873, Trans. Amer. Phil. Soc, XV, p. 249. Venus ivoodwardi Guppy, 1874, Geol. Mag. London, p. 442; 1876, Quart. Jour. Geol. Soc. London, XXXII, p. 536. Chione ivoodwardi Dall, 1903, Trans. Wagner Inst. Sci., Ill, pt. 6, pp. 1291, 1587. Chione woodivardi Maury, April, 1917, Bull. Amer. Paleont., No. 29, p. 218, PI. XXXVII, fig. 6. Chione woodicardi Pilsbry and Johnson, May, 1917, Proc. Acad. Nat. Sci. Phila., p. 199. Guppy first described this species from a Jamaican shell, later he identified it at Cumana, Venezuela, and in 1876 found it among the left over specimens of the Heneken Collection from the Yaqui Valley, Santo Domingo. Dall also cites it from Bowden, Jamaica and Santo Domingo. In 1916, my party collected Chione wood- ivardi in abundance in Santo Domingo. The specimens correspond in all respects to casts made from many external molds in the Porto Rican limestone. These molds show the characteristic single radial riblets of C. ivoodwardi which are never paired, nor split into smaller ribs, as in related species. Curiously, Gabb failed to find C. ivood- wardi in Santo Domingo, and that is doubtless why Pilsbry and Johnson remark that it does not appear to occur there, — as it is not in the Gabb collection at Philadelphia. But Guppy, Dall, and the MAURY, TERriARY MOLLUUCA FROM PORTO RICO 41 writer have all noted its presence in Santo Domingo. This pretty shell was evidently quite widespread in Porto Rico. Localities.— 'Ne&T Quebradillas, Stations 125, 129, 136; near Ponce, Station 299; near Ensenada, Stations 392, 393, 394; near Manati, not in situ, from boulders, Station 474. Chione quebradillensis, new species Plate VII, Figure 4 Shell subtrigonal, anterior end rounded, posterior end somewhat pointed. The surface is handsomely sculptured by about fifteen concentric, crenulated ridges which are cut by the deeply impressed lines defining the lunule, and then continue on over the lunule itself. Over the anterior and posterior areas of the valve the radial sculpture consists of very fine and close-set riblets more or less uniform, only the posterior are somewhat finer and closer. Upon the central portion of the valve the radial ribs are less crowded, although still close, and usually show a definite alternation of a compara- tively broad, unpaired, rib with a very fine, linear rib. Length of shell 21 mm., height 16 mm., semi-diameter 9 mm. This species is related to Chione walli Guppy, from the Manzanilla beds of Trinidad and also recorded from Bowden, Jamaica ; but that shell has a great many more concentric ridges, and the central rib- lets alternating with the small intermediate ribs are paired. Lately Pilsbry and Johnson have briefly described, but not figured, a Dominican shell, Chione saniodomingensis, which is apparently of the same group. The Floridian analogue is C. chipolana Dall. The Porto Rican collection afforded a single but very perfect external mold of this shell imprinted in the limestone. Locality. — Near Quebradillas on Rio Guajataca, Station 134. Chione quebradiUensis guajatica, new variety Plate VII, Figure 5 A fragmentary external mold was collected by Reeds which re- sembles the species described as Chione quebradillensis, but the ribs over the central area of the valve are paired and alternate with the fine, intermediate rib. This character is as in Guppy's Chio7ie walli, but in the Porto Rican shell the concentric crenulated ridges are distant, while in C. ivalli they are very close, almost crowded. Locality. — Near Quebradillas, Station 133. 42 SCIENTIFIC SURVEY OF PORTO RICO Genus tellina (Linnaeus) Lamarck This genus is well represented in the collection, but nearly all the specimens are internal molds, too imperfect for specific identifica- tion. From the Quebradillas limestone comes a fragmentary imprint of a Sciasula near S. cercadica Maury but probably not that species. Subgenus Angulus (Mergele) Dall Tellina (Ang-ulus) disparoides, new species Plate VI, Figure 1 Internal mold with short elliptical form; comparatively straight dorsal margin on either side of the low, a?ute beaks. Hinge characters and sur- face markings unknown. On comparing this mold with recent species in the American Miiseiun it was found to be strikingly like specimens labelled Tellina {Tellinula) dispar Conrad from California and from the Hawaiian Islands. When a shell of similar size of Tellina dispar was placed upon the mold it coincided at nearly every point. Failing any evidence regarding the surface characters of the Porto Rican fossil, we can only say that judging from the form alone its relationship is seemingly with this recent Pacific species. Length of fossil 25 mm., altitude 18 mm., semi-diameter 6 mm. Localities. — Rio Collazo near San Sebastian, Station 60. Im- perfect mold, probably the same species, from Station 57. A smaller mold, perhaps identical, from Station 62. Genus Metis H. and A. Adams Metis trinitaria Dall Tellina Uplicata Guppy. ISGG, Quart. Jour. Geol. Soc. London. XXII. p. 588; 18G7, Proc. Sci. Assoc. Trinidad, p. 161: 1874. Geol. Mag. London, p. 441; 1876, Quart. Jour. Geol. Soc. London. XXXII, p. 530. Not Tellina bipHcata Conrad, 1834. Jour. Acad. Nat. Sci. Phila , VII, p. 152. Tellina sagrw Guppy and Ball, 1896, Proc. V. S. Nat. Mus., XIX, p. 329. Not Tellina sagrcr d'Orbigny. Metis trinitaria Dall, 1900, Trans. Wagner Inst. Sci., 111. pt. 5, p. 1041, PL XLVI, fig. 24. Metis trinitaria Maury, 1917, Bull. Amer. Paleont., No. 29, p. 226. Internal molds of Metis are very abundant. Three external nuilds occur, casts of which correspond with DalFs figure of Metis trini- taria, except they are less truncate posteriorly. Still they vary in this respect, some being less produced than others. In one case MAURY, TERTIARY MOLLUSCA FROM PORTO RICO 43 the internal mold still rested within the external, thus removing all possible doubt as to whether the internal molds were of the same species. This shell is easily known by its striking form, the right valve being medially constricted as though pressed in by the thumb, while the left valve is convex. This gives a peculiar aspect, different from any of the other Porto Rican fossils. The internal molds show an immense, rounded pallial sinus, which is slightly larger and higher in the left than in the right valve. The posterior end of both valves is sulcated, that of the right being noticably bisulcate. Surface smooth except for concentric lines of growth. Dr. Guppy referred this Antillean shell to Tellina biplicata Conrad from the Miocene of Maryland, and in 1874 listed it from the Miocene of Barbuda, and Santo Domingo, from the Caroni Series of Trinidad, and the Tertiary of Cuba. In 1900, Dall differentiated the Antillean fossil by the name M. trinitaria and recorded it from near Santiago de Cuba, at about two hundred and fifty feet above sea level along the ore railway. A specimen was seen by Guppy in the Heneken collection from the Yaqui Valley, Santo Domingo, but both Gabb and the writer failed to collect it there. It has not been reported from Jamaica, but it ranged from Trinidad north to Barbuda and thence west to Porto Eico, Santo Domingo and southeastern Cuba. The number of speci- mens indicate very favorable conditions in Porto Rico. On the one hand Metis trinitaria resembles the Chesapeake Mio- cene M. biplicata and on the other the Pleistocene and recent M. intastriata Say. The Tellina sagrce of Orbig-ny's unpublished Pale- ontologica Cubana pi. IV, figs. 8, 9, was an internal cast, according to Dall probably of M. intastriata. Localities. — Rabell's Ranch, on Guajataca River, Station 110; ' near Quebradillas, Stations 125, 127, 128, 129, 130, 131, 133, 135, 136, 137, 138, 144. Also a very poor internal mold possibly thisi species from near Guayanilla, Station 320. Genus Semele Schumacher Semele, species indet. A number of distorted specimens of a Semele somewhat resembling Semele chipolana Dall come from the Collazo shales, but the shells 44 SCIENTIFIC SURVEY OF PORTO RICO are too deformed and imperfect for specific description. Attention is simply called to the presence of the genns in this formation. Localities. — Rio Collazo near San Sebastian, Stations 36, 53, 54. Genus Tagelus Gray Section Mesopleura Conrad Mesopleura hubbardi, new species Plate VII, Figure 3 Shell transversely oblong-elliptical, compressed; anterior and posterior ends evenly rounded; basal margin gently arched; beaks very low, almost exactly central; muscle scar prominent; the most conspicuous character is the very strong, nearly straight, internal rib which is directed downward from the beak a varying distance towards the ventral margin. In the internal molds this rib appears in reverse as a strongly-marked groove. Length of shell 35 mm., altitude 14 mm. The thickness varies with the pressure to which the specimens have been subjected after deposition in the shales. A number of specimens of a Tagelus somewhat resembling Tage- his {Mesopleura) divisus Spengler, and probably an ancestral form of that Pliocene to recent species were found in the Collazo shales. This species is named in honor of Mr. Bela Hubbard, as a token of appreciation of his very accurate and valuable paleontological work in northwestern Porto Rico and on Mona Island. Localities. — Rio Collazo near San Sebastian, Stations 53, 62. Genus Psammosolen Risso Psammosolen cf. sancti-dominici Maury Psammosolen sancti-dominici Maury, 1917, Bull. Amer. Paleont., No. 29, p. 228, PI. XXXVII, fig. 13; Idem., No. 30, p. 14. A very imperfect mold shows the general characters of Psam- mosolen, and retains very clearly the characteristic, oblique surface markings of this genus. The shell may have been identical with the form we collected at Bluff ,3, Cercado de Mao, Santo Domingo, and named Psammosolen sancti-dominici, as it is about the size, but it is too poorly preserved for specific identification. Locality. — Near Quebradillas, Station 131. Genus Corbula (Bruguiere) Lamarck Corbula collazica, new species Plate VI, Figures 10, 11 Shell ovate-triangular, of medium size for the genus, anterior end rounded, posterior end produced, right valve somewhat larger than the MAURY. TERTIARY MOLLUSC A FROM PORTO RICO 43 left, its posterior end obliquely truncate, its surface sculptured with about twenty-five to thirty fine, close-set, even, subequal concentric ridges, alter- nating with linear interspaces; left valve somewhat smaller and pointed posteriorly to fit within the truncated end of the right valve. Concentric sculpture of both valves similar. Length of shell 13 mm., height 8 mm.. semi-diameter 3.5 mm. This Corhula is one of the commonest and most characteristic shells in the Collazo shales. Fossil species of this genus already described from the Greater Antilles are the following: Corbula sericea Dall, C. heterogenea Guppy, C. vietck Guppy, C. viminea Guppy, C. dominicensis Gabb, C. cercadica and C. caimitica Manry. From all of these the present species differs. Owing to pressure resulting from the disturbance of the shales, the shells are more or less deformed, and at first it seemed as if two forms might be present, one more triangular, the other more nasute. But this difference seems due rather, to accidents, as the sculpture and general aspect of all are similar, and I have selected as the type a specimen that seems to have suffered least from pres- sure and to represent what appears to have been the normal form. Localities. — Eio Collazo near San Sebastian, Stations 23, 49, 53, 54. Genus Panopea Menard Panopea, species indet. An imperfect internal mold of what appears to represent a very large and extremely obese Panopea was found in the collection. The mold measures 90 mm. in length, 70 mm. in altitude and 76 mm. in diameter. . Locality. — Xear Arecibo, Station 459. Genus Teredo Linnaeus Teredo incrassata (Gabb) Kuphiis incrassatus Gabb, 1873, Trans. Amer. Phil. Soc, XV, p. 246; 1881, Jour. Acad. Nat. Sci. Phil., (2), VIII, p 342, PI. XLIV, figs. 12a-e. Teredo fistula? Guppy, 1876, Quart. Jour. Geol. Soc. London, XXXII, p. 529. Teredo incrassata Dall, 1903, Trans. Wagner Inst. Sci., Ill, pt. 6, p. 1587. Teredo incrassata Maury, 1917, Bull. Amer. Paleont., No. 29, p. 235, PI. XXXIX, fig. 24; Idem, No. 30, p. 25. Kuphus incrassatus Vaughan, 1919, Bull. 103 U. S. Nat. Mus , p. 558. Gabb first described the tubes of this Teredo from Guayubin, Santo Domingo. Later he found it was abundant and widespread in the Miocene of that island, throughout the Samba Hills and also south of Samana Bay. He states that, "it is a very common 46 SCIENTIFIC SURVEY OF PORTO RICO and characteristic fossil of the upper part of the Dominican Mio- cene."' The writers party in Santo Domingo also collected Teredo incrassata in abundance. The tubes were seen loose along the road- side from Los Quemados to Caimito. They were several feet long and resembled the roots of trees. At Caimito they were present in the Teredo limestone and occurred at Cercado in Bluft' 3. Thus this species ran through both the Scoiisia laevigata and Aphera islacolonis formations. Dall lists Teredo irin-assata from Bowden, Jamaica, in the Simp- son and Henderson collection. Its range is now extended to Porto Rico, where Reeds found it in gi-eat quantities and, as the list of localities shows, extremely wide spread. Besides its distribution in the Miocene of these three islands of the Greater Antilles, Teredo incrassata was also identified bv Gabb at Sapote on Rio Reventazon, Costa Rica. It is of interest that although such large numbers of the tubes have been found, no valves have vet been discovered. The shell therefore remains unknown. The descriptions and identifications rest entirely upon the tubes. These are wrinkled with annular growth lines, often somewhat gnarled and may attain several feet in length and an inch and a quarter in diameter. Smaller portions of the tubes are frequently divided into two by an internal septum. Localities. — Rio Collazo near San Sebastian, Stations 24, 25, 33, 35, 36, 38, 39, 44, 45, 46, 48, 49, 51, 52, 54, 57, 58, 59, 61, 63 ; near Lares, Stations 82, 92 ; Rio Collazo, Station 95 ; Seiior Rabell's Ranch, Rio Guajataca, Stations 110, 111. 112; near Aguada, Sta- tions 117, 119, 122, 123; near Quebradillas, Station 126; near Juana Diaz, Stations 160, 163, 184, 189 ; near Ponce, Stations 288, 292, 299, 300; East Shore of Guanica Harbor, Stations 368, 373, 374, 375 ; near Guanica Central, Station 392 ; near Arecibo, Sta- tions 441, 469; near Manati, Stations 474, 477, 478, 479, 480, 481, 482, 483; near Ciales, Stations 495, 498, 500, 502; near Arecibo, Station 509. CLASS SCAPHOPODA Genus Dentalium Tjinnaeus Dentalium diazicum, new species Plate VII, Figure 1 Shell with no longitudinal sculpture, polished and entirely smooth except for faint miscroscopic, annular growth lines, tube curving, tapering con- MAURY, TERTIARY MOLLUSC A FROM PORTO RICO 47 siderably, circular in cross section. Length of fragment 19 mm., diameter of larger end 2.75 mm, of smaller end 1.50 mm. Tliis species is about the size and has the same curvature as apical fragments of Dentalium dissimile Guppy from the Miocene of Jamaica and Santo Domingo but is at once differentiated by its absence of longitudinal sculpture. Dentaliinyi haytense Gabb from Santo Domingo is smooth and circular in cross section, but the tube is nearly straight. Dentalium pyrum Pilsbry and Sharp is ovate in section, while D. prcecursor, another smooth form described by the same authors is oval in cross section. Among other Miocene Antillean Dentalid may be mentioned D. caUiglyptum, D. tryoyii and D. cossmannianum of Pilsbry and Sharp, and B. glaucoterrarum Maury which are all sculptured species. Locality.- — Near Juana Diaz, Station 185. Genus Cadulus Philippi Cadulus poncensis, new species Plate VII, Figure 13 Shell small, curving, not very slender, circular in cross-section, slightly constricted at the larger extremity, evenly convex up to about two milli- meters from the apex when it rather suddenly tapers. Just at the apex the shell seems very slightly swollen. The surface apparently was perfectly smooth. Length of shell 7 mm., greatest diameter 1.50 mm. From a chalky phase of the limestone near Ponce comes a single specimen of a Cadulus. The shell substance appears to have been replaced, but the original form is well preserved. The small Do- minican species, Cadulus phenax Pilsbry and Sharp (Proc. Acad. Xat. Sci. Phila., XLIX, p. 472, Ph XI, figs. 23, 24, 1897) of which we collected hundreds of shells in the bluffs of Rio Mao, is about this length, but is a very much more slender and delicate shell. The Bowden form C. dentalinus Guppy, is characterized by circular rings around the smaller part of the tube. Cadulus denticulus- tigris Maury from Rio Gurabo, Santo Domingo, is about the length of the Ponce shell, but is much more inflated centrally. Locality. — Xear Ponce, Station 283. 48 SCIENTIFIC SURVEY OF PORTO RICO CLASS GASTROPODA Order Ctenobranchiata Genus Calyptrsea Lamarck Calyptrsea centralis (Conrad) Infundibulm centralis Conrad, 1841, Amer. Jour. Sci., XLI, p. 348; 1S45, Foss. Medial Tert., p. 80. PI. XLV, fig. 5. Trochita centralis Emmons, 1858, Geol. N. Car., p. 276, fig. 193. Trochita colUnsii Gabb, 1875, Jour. Acad. Nat. Sci. Phila., (2), VIII, p. 342, PI. XLIV, figs. 11, 11a. Calyptrwa centralis Dall, 1892, Trans. Wagner Inst. Sci., Ill, pt. 2, p. 353. This widespread Miocene species is represented by a single in- ternal mold, but the characters of the pillar are concealed by the matrix. The recent analogue and probable descendant of C. cen- tralis is Calyptrcea candeana d'Orbigny, of which Dall and Simpson report finding a great number of dead shells at Mayaguez. Calyptrcea centralis occurs according to Dall in the Chipola Miocene of Florida and in the Miocene of ]^orth and South Carolina, Virginia, Mary- land and Sapota, Costa Rica. Locality. — Xear Quebradillas, Station 130. Genus Natica Scopoli There are many internal molds of Katicoid shells in the collection. They are evidently of various species but scarcely any can be spe- cifically determined. Natica cf. canrena (Linnaeus) Moercli Nerita camerna (Linnaeus in part) Auct., Mcerch, 1877, Malak. Blatt.. 24. p. 62. Natica canrena Gabb, 1873, Trans. Amer. Phil. Soc, XV, p. 223. Natica canrena Guppy, 1876, Quart. Jour. Geol. Soc, XXXII, p. 518. Natica canrena Dall, 1892, Trans. Wagner Inst. Sci., Ill, pt. 2, pp. 364-365. Natica canrena Brown and Pilsbry, 1912, Proc. Acad. Nat. Sci. Phila.. p. 508. Natica canrena Maury, 1917, Bull. Amer. Paleont, No. 29, p. 134, PI. XXIII, fig. 10; Idem. No. 30, pp. 10 27. Natica canrena Vaughan, 1919, Bull. 103 U. S. Nat. Mus.. p. 561. One specimen agrees well in form with Natica canrena and is probably that species, but since it is an internal mold, it cannot show the characteristic, tangential surface plicge which would defi- nitely determine the identification. Natica canrena is an ancient, MAURY, TERTIARY MOLLUSCA FROM PORTO RICO 49 static form rangiug from the Tertiary to the recent. It is found as a fossil in Trinidad, Santo Domingo, Gatim, Costa Rica, and Florida. Its present range is from the Antilles to Pernambuco, Brazil. Locality. — East shore of Guanica Harbor, Station 373. Section Stigmaulax Moerch Natica (Stigmaulax) sulcata (Bom) Nerita sulcata Born, 1780, Mus. Cses. Vindobonensis, p. 400, PI. XVII, figs. 5, 6. Natica sulcata Guppy, 1866, Quart. Jour. Geol. Soc. London, XXII, p. 290, PI. 18, figs. 14, 15; 1867, Proc. Sci. Assoc. Trinidad, p. 157. Natica sulcata Gabb, 1873, Trans. Amer. Phil. Soc, XV, p. 223. Natica sulcata Guppy, 1874, Geol. Mag. London, p. 437; 1876, Quart. Jour. Geol. Soc. London, XXXII, p. 518. Natica (Stigmaulax) sulcata Ball, 1892, Trans. Wagner Inst. Sci., Ill, pt. 2, p. 366. Natica sulcata Ball and Simpson, 1902, Bull. U. S. Fish Comm., XX, pt. 1, p. 438. Natica {Stigmaulax) sulcata Ball, 1903, Trans. Wagner Inst. Sci., Ill, pt. G, p. 1585. Natica (Stigmaulax) sulcata Maury, 1917, Bull. Amer. Paleont., No. 29, p. 135, PI. XXIII, fig. 13; Idem, No. 30, pp. 11, 22. 24, 27. Natica (Stigmaulax) sulcata Jones, 1918, Jour. Geol. XXVI, p. 742. As early as 1867, Guppy, the pioneer and indefatigable paleon- tologist of the West Indies, had listed this species from the Miocene of Cumana, Venezuela, and from Jamaica and Santo Domingo. Dall has also recorded it from Bowden, and Jones lately found it in the Maissade beds, Eiviere L'Ayaye, Haiti. My Dominican expedition found it in abundance in the Miocene blue clays, the specimens from Bluif 3, Cercado de Mao, being particularly large and fine. Eeeds obtained a single much broken shell from Porto Kico which though so imperfect has the groovings and form char- acteristic of this widespread species. This is one of the few cases where the Miocene shell has lived on unchanged to the present time. It is still found on the Porto Rican shores. Natica sulcata is easily recognized by its ornamentation of uniform groovings which traverse the entire surface, keeping parallel to the margin of the outer lip. Locality.— B.iibelV& Kanch on Rio Guajataea, Station 110. 50 SCIENTIFIC SURVEY OF PORTO RICO Genus Amauropsis Moerch Amauropsis guppyi gurabensis Mauiy Amauropsis guppyi gurabensis Maury, 1917, Bull. Amer. Paleont., No. 29, p. 138, PI. XXIII, fig. 20; Idem. No. 30, p. 24. Amauropsis guppyi gurabensis Jones, 1918, Jour. Geol. XXVI, p. 742. An internal mold in the collection shows the broad form, short, acute spire, and broadly shouldered whorls that characterize this variety' of Gabb's Amauropsis guppyi (Amaura Guppyi Gabb, Trans. Amer. Phil. Soc, XV, p. 224, 1873). The typical form has a more evenly sloping spire and proportionally longer body whorl. My party found the variety, gurabensis rather commonly in the bluffs of Rio Gurabo, Los Quemados, Santo Domingo. Jones later obtained it from the Maissade beds. Riviere L'Ayaye, Haiti. The Porto Rican mold agrees well in form with our Dominican shells. Locality. — Near Aguada, Station 123. Genus Xenophora Fischer de Waldheim Xenophora conchyliophora (Bom) Trochus conchyliophorus Born, 1778, Mus. Caes. Vinci., Index, p. 333. Phorus reclusus Conrad, 1855, Proc. Acad. Nat Sci. Phil., VII, p. 2C2. Phorus agglutinans Gabb, 1873, Trans. Amer. Phil. Soc, XV, p. 241. Xenophora agglutinans De Gregorio, 1890; Mon. Eocene Ala., p. 144. Not of Lamarck. Xenophora conchyliophora Dall, 1892, Trans. Wagner Inst. Sci, III, pt. 2, pp. 3C0-3G2; 1915, Bull. 90, U. S. Nat. Mus., p. 105. PI. XV, figs. 1. 3. Xenophora conchyliophora Dall, 191(), Proc U. S. Nat. Mus., LI, p. 518, PI. LXXXVI, fig. 10. Xenophora conchyliophora Maury, 1917, Bull. Amer. Paleont., No. 29, p. 133, PI. XXIII, fig. 7; Idem. No. 30, pp. 22, 23. The Porto Rican representatives of Xenophora are all internal molds; but almost undoubtedly they are the fillings of shells of X. conchyliophora. This ancient, static species dates from the Ripley Cretaceous of Alabama through Eocene, Oligocene, Miocene and Pliocene times down to the recent fauna. It is living in 14 to 250 fathoms from Ilatteras to Guadeloupe and in the West Indies generally. The Dominican Miocene shells that my party gathered from the blulfs of Kio Gurabo were in all respects like those on the beaches of that island so it is no surprise to find this species in synchronous, Porto Rican deposits. Localities. — Near Quebradillas, Stations 12G, 128, 129. MAURY, TERTIARY MOLLUSCA FROM PORTO RICO 51 Genus Turritella Lamarck Turritella altilira culebrina, new variety Plate VIII, Figure 4 Shell with uniform spirals in the concave central part of the whorls; two raised spiral ridges one near the proximal and one near the distal suture. The lower may have been slightly beaded, the upper is crested with a double row of beads. Grooved area between ridges sculptured v.ith five or six uniform spiral threads. Altitude of most complete specimen 50 mm. greatest diameter (increased by flattening from pressure) 12 mm. There are several Turritellas from the Collazo shales, which though flattened by pressure have retained the shell substance and sculpture. They are remarkably like the Gatun species T. altilira Conrad {T. gabhi Toula). It may seem strange to refer this species to the Gatun shell rather than to Guppy's Turritella tornata def- scribed from Cumana, Venezuela, and listed also from Jamaica. But these shells are so much more like Toula's, and Brown and Pilsbry's figures of the Gatun species than like Guppy's figure of the Cumana type that without a series of the actual fossils this identification seems truer. Localities. — Kio Collazo near San Sebastian, Stations 54, 60, 61, 94. TurriteUa coUazica, new species Plate VIII, Figure 5 Shell large and heavy, flattened by pressure but still retaining its char- acteristic and very striking sculpture. The unique fragment consists of four whorls each with a very heavy and prominent broad swollen band encircling the base just above the proximal suture. This band occupies about one-third of the whorl. The remaining portion is flat and shows traces of having been ornamented with about six coarse spirals. Height of four whorls 65 mm., greatest diameter (increased by flattening of the shell) 35 mm., least diameter 20 mm. This species is at once recognized by the very heavy rounded ridge defining the base of each volution. The species with which it seems to be most closely related is the North American Eocene shell, Turritella mortoni Conrad. I know of no Antillean species at all resembling it. The fossil would seem to be a descendant of Con- rad's shell with the ridge not sharp and carinate as in that species but broad, round and swollen. Locality. — Rio Collazo near San Sebastian, Station 64. 52 SCIENTIFIC SURVEY OF PORTO RICO Turritella poncensis, new species Plate VII, Figure 6 Shell of medium size, tapering rather gradually from the base towards the apex. Whorls flattish, each usually marked by six stronger primary spiral threads between which are a varying number of fine secondary spirals visible only with a lens. Anteriorly the whorls are roundly carinated and project so as to partly overhang the suture immediately beneath them. This overhanging of the basal part of the volutions is the most conspicuous character of the shell and differentiates it at once from the common West Indian Miocene species, Turritella planigyrata Guppy. A fragment of about four whorls measures 28 mm. in length and 14 mm. in greatest diameter. This species seems nearest akin to the larger Gatun shell, Turri- tella mimetes Brown and Pilsbry. Locality. — !N"ear Ponce, Station 285. Turritella guanicensis, new species Plate VII, Figure 7 Shell was fairly large, tapering rather rapidly towards the apex. The sculpture of the two whorls that precede the last volution consists on each of an anterior and a posterior pair of spiral, beaded ridges separated by a narrow groove. Between the whorls is a concave space in the center of which lies the suture. On the last shorl, in addition to the two pairs of beaded, spiral ridges, there is a somewhat smaller, basal carina, just beneath the anterior pair making them appear like three ridges and forming in all five instead of four ridges on the body whorl. On this whorl also there is a threadlike secondary spiral in the groove between the two pairs of beaded ridges. A fragment of about four volutions measures 22 mm. in length and 12 mm. in greatest diameter. On the one hand, this species bears some resemblance to Turritella forresti Brown from Willoughby Bay, Antigua, but that shell is much smaller, about half the diameter of the Porto Rican, and has three major spiral ridges, beaded and paired, and between these three groups of major ridges are clusters of five, finer, secondary spirals. On the other hand, the aforementioned Ponce shell recalls in quite a striking manner the sculpture of the much later Floridian Turritella apicalis tensa Dall, which has also two pairs of beaded ridges to every whorl and seems to be the Pliocene analogue of the Porto Rican species. Locality. — Guanica Harbor, Station 374. MAURY, TERTIARY MOLLUSC A FROM PORTO RICO 53 Genus Cerithium Bruguiore Cerithium russelli arecibense, new variety Plate VII, Figure !) Each whorl ornamented with a <-ow of small, nearly iiniform beads, about eight to a half volution. Beneath this beaded row a series of rounded, not perfectly regular longitudinal ribs, about twelve to a volution. Entire surface marked by fine incised spirals, visible without the lens; on last whorl tending to alternate with secondary, microscopic spirals. Last two whorls measure 15 mm., in height and 11 mm., in greatest diameter. This variety is founded upon a fragmentary external mold of a Cerithium resembling C. russelli Maury, from Los Quemados, Rio Gnrabo, Santo Domingo. It differs notably in its smaller size and details of ornamentation. The sculpture of the Porto Rican shell is rather bolder, the ribs few, stronger and more regular than in the Dominican type. Locality. — Near Arecibo, Station 471. Cerithium utuadicum, new species Plate VII, Figure 8 Whorls, three of which show in the rock imprint, slightly convex and ornamented with arching, rounded, longitudinal ribs, about nine to half a volution, separated by narrower interspaces. A few of the ribs can be traced from suture to suture across the whorl, but the majority are so interrupted posteriorly by an abrupt change of sculpture as to lose their identity. This change consists in the development of a band bearing, regular, uniform, bead-like nodules, twelve or more to half a volution. This beaded row comes out well in the artificial cast. Beneath it is a very fine row of nodules, best seen in the external mold, as it is too delicate to form a good impression. Beneath the row of fine nodules are about seven primary spiral threads, usually alternating with finer, secondary spirals. Height of frag- ment 16 mm., greatest diameter approximately 10 mm. The specimen consists of the fragmentary mold of a richly-sculp- tured, turreted shell which at first glance one would think a Terehra, but on examination under a lens the ornamentation appears more like that of certain unusual Cerithia. This species has a decided general resemblance to the figure of Cerithium coraUicolum Dall from the Upper Flint River beds, Georgia (Proc. U. S. ^at. Mus., LI, PI. LXXXVII, fig. 6, 1917). But the Porto Rican shell is smaller, slenderer, with finer, more numerous ribs, a minor beaded 54 SCIENTIFIC SURVEY OF PORTO RICO row beneath the major, and is a more elegantly sculptured shell than the earlier Georgian form. A recent specimen in the American Museum labelled Vertagus faciatus Bruguiere, Jay Collection, from Ceylon, resembles the fossil in having a nodose band, giving the shell a Terebra-like aspect, but the ribs are lower, straighter, closer and more numerous and the spirals fewer than in the fossil species. Locality. — About midway between Arecibo and Utuado, Station 471. Subgenus Campanile Bayle Section portoricia, new section Shell large, many-whoiied, turreted, very elongate-conic, tapering gradu- ally from a comparatively narrow tase to the apex. Suture very profoundly excavated. Whorls rounded, each shewing distinct traces of a narrow, deep medial groove, perhaps r:p:esenting surface sculpture. The specimens are all in the form of internal molds. The cylin- drical axial perforation represents the space occupied by the dis- solved columella. The interior surface of this perforation has a coating of calcite crystals which obscures to some extent the struc- ture, but there is no indication of any grooves representing in re- verse columella plications. The pillar apparently was smooth. Although so imperfect, the suggestion of a new section for these singular molds seems justified by their unique characters. In the very deep sutures, rounded whorls and curious Pleurotomarian-like groove they differ from such forms as Campanile claytonense Al- drich from the Midwayan Eocene of the Gulf States and also from the great Cerithium giganteum of the Paris basin Eocene. In a superficial way they bear more resemblance to Ilalloysia biplicaia Briart and Cornet from the lower Eocene of Mons, but that has a plicate columella and its affinities are with Nerinea. The presence in North America of large Ceritliiums recalling those of the Paris basin Eocene was first noted by Dall in a letter" to Dr. Fischer regarding the basal Eocene beds of the Gulf States. He remarked: ''Dans ces calcaires du Tertiaire inferieur se mon- trent des fragments d'un Cerithium de tres grande taille, paraissant voisin du C. giganteum de I'Eocene du Bassin de Paris. Or, ce type est trouve pour la premiere fois en Amerique." In 1894, Aklrich 'Published in Bull. Soc. Geol. de France, XVIII, (3) 1890, p. 327. MAURY, TERTIARY MOLLUSCA FROM PORTO RICO 55 described Cerithiuni (Campanile) claytonoise from tiic -Midwavan Eocene of Alabama. This attains 125 mm., and has flattened whorls with the suture not excavated. In a discussion of the Oligocene fauna of Bainbri.dge, Georgia, Dall remarks:'^ "A feature of somewhat unusual interest paleon- tologically is the presence in the upper bed of a relatively large number of species of the Cerithiidae, several of them of unusual size, recalling the analogous group in the Parisian Eocene of France, and not paralleled in any of the other Tertiary horizons of the United States as far as known." These Porto Rican specimens are, however, very much larger than any Cerithiums from the Flint River Oligocene, nor are they like Dall's species. I know of no Xorth American or x\ntillean fossil resembling these Porto Rican molds. I believe they represent a unique fauna having affinities with European Eocene forms, but of a younger geological period. Campanile (Portoricia) laricum, new species Plate VIII, Figures 1, 2 At the base of the Arecibo limestone a number of internal molds were collected for which I have suggested the above described new section Portoricia. The largest fragment consists of six whorls and shows evidences of pressure, resulting in distortion. The six volu- tions of this mold measure 80 mm., in height. Another fragment of three whorls is 33 mm. in height and 45 mm. in greatest diameter. All the fragments appear to have been the same species and all show interrupted traces of the medial groove, which looks like the mark- ings on internal molds of Pleurotomarian shells, but was probably some surface sculpture that grooved the internal wall of the volution. Localities. — ISTear Lares, Stations 57, 58, 59. Genus Strombus Linnaeus Strombus cf. haitensis Sowerby strombus haitensis Sowerby, 1S50, Quart. Jour. Geol. Soc. London, VI, p. 48, PI. IX, fig. 7. Strombus haitensis Guppy, 18C7, Proc. Sci. Assoc. Trinidad, p. 157. Strombus bituberculatus Gabb, 1873, Trans. Anier. Phil., Soc, XV, p. 233. Not of Lamarck. »Proc. U. S. Nat. Mus., LI, 1917, p. 487; Pis. LXXXIII to LXXXVIII. 56 SCIENTIFIC SURVEY OF PORTO RICO Strombus haitensis Guppy, 1874, Geol. Mag. London, p. 438; 1876, Quart. Jour. Geol. Soc. London, XXXH, p. 521. Strombus haitensis Dall, 1903, Trans. Wagner Inst. Sci., IH, pt. 6, p. 1584. Strombus haitensis Maury, 1917. Bull. Amer. Paleont., No. 29, p. 118, PI. XX, fig. 1; Idenu No. 30, pp. 10, 18, 22, 23, 24. Several internal molds probably represent fillings of Strombus haitetisis. When compared with shells of this species collected by the writer from the type region in the Yaqui Valley, Santo Domingo, the molds are seen to correspond to the interior of the shell. They also show traces of the stronger, npper row of spiny prongs. Abso- lute certainty is not possible, but in all probability the molds are those of S. haitensis. First described by Sowerby from Heneken's Dominican shells, this species has been later found in the Yaqui Valley by Gabb and the writer. Gabb referred his specimens to the living Strombus bituberculatus. The differences between the Miocene and the recent species are noted in my descriptions of Dominican fossils. Dall lists S. haitensis from Henderson's and Simpson's collection of Bowden fossils. The species thus seems to have lived on the Miocene shores of the adjacent islands of Santo Domingo, Jamaica and Porto Rico. Localities. — Senor Rabeirs Ranch, Rio Guajataca, Station 112; near Aguada, Station 117. Strombus cf. bifrons Sowerby strombus bifrons Sowerby, 1850, Quart. Jour. Geol. Soc. London, VL p. 48. PI. IX, fig. 9. Strombus bifrons Guppy, 1866, Quart. Jour. Geol. Soc. London, XXII, p. 287; 1867, Proc. Sci. Assoc. Trinidad, p. 157. Strombus pugilis Gabb (In part) 1873, Trans. Amer. Phil. Soc, XV, p. 233. Not of Linnaeus. Strombus bifrons Guppy, 1874, Geol. Mag. London, p. 438. Strombus bifrons Guppy, 1876, Quart. Jour. Geol. Soc. London, XXXII, p. 521. Strombus bifrons Dall, 1890, Trans. Wagner Inst. Sci., Ill, pt. 1, pp. 176, 177; 1903, Idem, pt. 6, p. 1584. Strombus bifrons Maury, 1917, Bull. Amer. Paleont., No. 29, p. 119, PI. XX, figs. 2. 3; Idem, No. 30, pp. 10, 13, 18, 22, 23, 27. In the collection is an internal mokl which probably represents Strombus bifrons Sowerby, When compared with Dominican shells of that species, it corresponds in size and form to the interior; but no positive identification is possible because of the very imperfect preservation of the Porto Rican specimen. The type locality of MAURY, TERTIARY MOLLUSC A FROM PORTO RICO 57 Stronibus hifrons is the Yaqui Valley, Santo Domingo where it was first found by Colonel Heneken and described by Sowerby. Guppy and Dall have recorded its presence in the Bowden beds, Jamaica. This very pretty and graceful shell of the Miocene shores of the Greater Antilles had for its analogues in Florida Strombus aldrichi Dall and S. chipolanus Dall. The Isthmian S. gatunensis Toula resembled it to a less degree as it lacked the shoulder spines. The living representative of this type of Strombus is S. columha of the oriental seas. Locality. — Seiior Rabell's Ranch, Rio Guajataca, Station 110. Strombus cf. proximus Sowerby strombus proximus Sowerby, 1850, Quart. Jour. Geol. Soc. London, VI, p. 48, pi. IX, fig. 8. Strombus proximus Guppy, 1867, Proc. Sci. Assoc. Trinidad, p. 157. Strombus pugilis Gabb (in part) 1873, Trans. Amer. Phil. Soc, XV, p. 233. Not 8. pugilis Linnaeus. Exclude Gabb's other synonyms. Strombus proximus Guppy, 1874, Geol. Mag. London, p. 438; 1876, Quart. Jour. Geol. Soc. London, XXXII, p. 521. Strombus pugilis Dall (in part) 1890, Trans. Wagner Inst. Sci., Ill, pt. 1, p. 177. Not pugilis Linnaeus. Strombus proximus Maury, 1917, Bull. Amer. Paleont. No. 29, p. 119, PI. XX, figs. 4, 5; laem, No. 30, pp. 10, 13, 22, 27. Strombus proximus Jones, 1918, Jour. Geol. XXVI, p. 741. The type of S. proximus was collected by Colonel Heneken in the Yaqui Valley, Santo Domingo, seventy years ago. Twenty years and more later Gabb found it there also, but referred it erroneously to Strombus pugilis. Jones lately reports finding S. proximus in the Maissade beds. Riviere L'Ayaye, Haiti. Several internal molds in the collection are probably of this species as they correspond to the form of the Dominican shells. Positive identification is not possible, but as far as the material at hand permits one to form an opinion, the three very characteristic Dominican Miocene species, S. haitensis, 8. bifrons and S. proximus were also living in Porto Rico. The nearest living analogue of Strombus proximus seems to be >S'. gracilior Sowerby, from Panama. Localities. — Near Ponce, Stations 289, 292. LIBRARY;^ 58 SCIENTIFIC SURyEY OF PORTO RICO Genus Orthaulax Gabb Orthaulax aguadillensis, new species Plate IX, Figure 4 Shell large and heavy, form of spire short and blunt, like that of Orthaulax pugnax. This at once distinguishes the shell from the Dominican species, 0. inornatus Gabb, which is high-spired. A further characteristic of the shell is the evenly rounded form of the shoulder, which in cross-section would be almost perfectly circular. This marks it off very decisively from the Floridian, Chipolan species, Orthaulax gabbi Dall, which is markedly triangular at the shoulder. The spire measures 45 mm., in diameter. A single specimen of this Orthaulax was collected by Reeds at Agiiadilla. It is imperfect, but undoubtedly a typical member of this very important index genus. The shell was submitted to Dall who compared it with the types of the various known species in the National Museum. He replied that "The Orthaulax is nearest to 0. pugnax but as the margin of the spire is gone it is impossible to be certain. I think it is new." One might be criticized for describing so incomplete a specimen as new were this a less rare and less stratigraphically important genus. Moreover, no complete adult shell of either 0. inornatus or 0. pugnax has ever been found. Though heavy and apparently strong the shells seem to go to pieces very easily and usually one tinds only heads, as in this case or fragments of the heavy pillars. Xo other molluscan shells were found associated with this Orthau- lax, but an Echinoderm occurred. It is a little surprising that the nearest ally of this Porto Rican Orthaulax should be not Gabb's 0. inornatus, — from the adjacent island of Santo Domingo and in the Tampa and White Beach beds, Florida, but 0. pugnax. The latter ranges geographically from the Tampa, Florida beds and those of Bainbridge, on the Flint River, Georgia to Cuba, Antigua and the Canal Zone, and geologically from the Middle Oligocene of Antigua to the Upper Oligocene of the Tampa and Flint River formations. Locality. — Aguadilla, Station 3. Genus Cypraea Tjiun.Tus Cyprsea sancti-sebastiani, new species Plate VII, Figure 12 Molds of the usual oval-cylindrical Ci/pnra-shape, the outer lip extending considerably above the spire; exterior characters unknown. Length of MAURY, TERTIARY M0LLU8CA FROM PORTO RICO 59 smaller mold 18 mm., width 11mm., thickness 9 mm.; corresponding meas- urements of the larget- mold, which is figured, 29 x 17 x 12 mm. A number of molds of Cyprcea varying in size were obtained from the Collazo shales. The large specimens are so deformed that it is impossible to say whether they are identical with the less distorted smaller molds or not. Of the smaller specimens, described above, the first is about the size of the Miocene to recent species, Cyprcea spurca Linnaeus, but is less contracted basally and more cylindrical. The other is in size like C. spurcoides Gabb, but less inflated and appears to be specifically the same as the smaller specimen. Localities. — Rio Collazo near San Sebastian, Stations 24, 36. Genus Dolium Lamarck Subgenus Malea Valenciennes Malea camura Guppy Malea camura Guppy, 18C6, Quart. Jour. Geol. Soc. London, XXII, p. 287, PI. XVII, fig. 9; 1867, Proc. Sci. Assoc. Trinidad, pt. 3, p. 158. Malea Hngens Gabb (in part), 1873, Trans. Amtr. Phil. Soc, XV, p. 223. Not M. ringens Swainson. Malea camura Guppy. 1874, Geol. Mag. London, new ser., decade 2, I. p. 439; 187G, Quart. Jour. Geol. Soc, London, XXXII, p. 525. Malea camura Dall, 1903, Trans. Wagner Inst, Sci., Ill, pt. G. p. 1584. Malea camura Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phil., p. 356. Malea camura Maury, 1917, Bull. Amer. Paleont., No. 29, p. 112, PI. XIX, fig. 3; Mem, No. 30, pp. 11, 13, 23. 24. Malea camura Jones, 1918, Jour. Geol. XXVI, p. 741. Malea camura Vaughan, 1919, Bull. 103 U. S. Nat. Mus., p. 561. This species was first described by Guppy from an immature Jamaican shell. Later he identified it from Santo Domingo and Panama. Gabb referred his Dominican specimens to the recent West coast species, Malea ringens Swainson. Dall lists M. camura from the Henderson and Simpson collection of Bowden, Jamaica fossils; Pilsbry and Brown record it from Gatun ; Jones has lately found it in the Maissade beds, Riviere L'Ayaye, Haiti. The writer has obtained it in abundance from Cercado de Mao and Los Que- mados, Santo Domingo. On comparing an external mold and its artificial cast from Porto Rico with Dominican specimens the agreement is almost line for line with some of our younger shells of Malea camura, leaving no doubt of complete identity. The closest ally and probable descend- 60 SCIENTIFIC SURVEY OF PORTO RICO ant of Malea camura is M. ringens Swainson living on the West Coast. Zetek in Los Moluscos de la Republica de Panama, p. 29, 1918, lists the latter shell from Panama. The present distribution of the subgenus Malea is restricted to the West Coast of Central and South America to Polynesia and the Philippines. It has become strictly Pacific, while in Tertiary times it was abundant in the Antillean and Isthmian Atlantic waters. Locality. — ^ear Quebradillas, Station 131. Genus Pyrula Lamarck Pyrula hoveyi, new species Plate VIII, Figure 3 ; Plate IX, Figure 7 Shell fig-shaped, spire unusually high for the genus; entire surface reticu- lated by very close-set, sharply defined longitudinal ridges crossing the spiral sculpture and equalling in thickness the series of primary spirals. The spiral ornamentation consists of delicate, but sharp, primary spirals at intervals of two and a half to three millimeters apart, and between each pair of primaries are three much finer subequal threads. Thus the secondary thread which occupies the center of each interspace is only very slightly more pronounced than the two tertiaries which lie on either side of the secondary. The earliest whorls are not shown in the rock imprint of the shell, but the third from the body whorl shows the point at which the longitudinal ridges suddenly begin. The primary spirals clearly precede them, but how far back towards the apex they extended is unknown. Length of incomplete internal mold 42 mm., greatest width 26 mm. Very likely the shell when adult attained a much larger size. From the above description it is clear that this Porto Rican shell is not of the Antillean and East Coast Pyrula papyratia stock, of which Burnett Smith has described an ancestral Bowden form as Pyrula pihhryi. The P. papyratia race have pronounced secondary spirals, very much stronger than the tertiaries, resulting in a very different type of sculpture from our fossil, and the longitudinal lines are so inconspicuous that the surface is not reticulated. Xor is the Porto Rican fossil like Guppy's Pyrula carbasea from the Caroni series of Trinidad. That species has very characteristic sunken interspaces between the primaries, and appears to be an aberrant form. Two molds of the Pyrula, one being a nearly complete internal mold and the other an impression in the rock, are present showing most of the spire and the upper part of the body whorl. The sculp- MAURY, TERTIARY MOLLUSC A FROM PORTO RICO 61 tiire is perfectly preserved and an artificial cast of the external mold reveals every detail of the exquisitely fine ornamentation of the original. This fossil is closer to the Isthmian species Pyrula micronernatica Brown and Pilsbrv from the Pecten bed in the Culebra Cut, but was larger, with a much higher spire and with very close-set longi- tudinal ridges instead of the wide-spaced axial threads of that spe- cies. Nor is there any knotting at the points of intersection of spirals and axials as noted in P. micronematica. Nevertheless, the kinship of the Porto Kican shell is with this species, because like it, its relationship is to the West Coast and not to the East Coast group of Pyrulas. In the unusual prominence of the spire the Porto Rican fossil resembles recent specimens in the American Museum of Pyrula dussumieri Valenciennes, living in the China seas. In its sculpture it is almost precisely like a specimen of Pyrula reticulata Lamarck from Japan. Its affinities are very clearly with the recent Pacific fauna. The shell is named in honor of Dr. E. O. Hovey. Localities. — Near Arecibo, Stations 468, -i70. Genus Tritonium Link Subgenus Colubraria Schumacher Colubraria juanica, new species Plate IX, Eigures 2, 3 Shell of medium size, broadly fusiform, somewhat flattened on the oral and aboral faces; suture distinct, whorls very slightly convex; outer lip bordered with a heavy, marginal varix, and within bearing about sixteen short, sharp lirae; body whorl with an obsolete varix on the center of the oral side; aperture elliptical; anterior canal short, strongly recurved; entire surface of shell sculptured with a very fine, even reticulation formed by the intersections of longitudinal and spiral ridges. Length of decollate shell 23 mm., greatest width 9 mm. Dall mentions Colubraria lanceolata Menke from the Pliocene marl of the Caloosahatchie, a species which is also living in the Antilles. But that shell is much slenderer than the Porto Rican fossil. Locality. — Near Juana Diaz, Station 185. 62 SCIENTIFIC SURVEY OF PORTO RICO Genus Phos Montfort Phos costatus Gabb Phos costatus Gabb, 1873, Trans. Amer. Phil. Soc. XV, pp. 212, 213. Phos costatus Maury. 1917, Bull. Amer. Paleont., No 29, p. SS, PI. XIY, figs. 13, 14; Idem. No. 30, p. 13. Phos costatus Jones, 1918, Jour. Gcol. XXVI, n. 740. Amoiiii- the fossils is an external mold of a Phos which shows the robust form, deep suture, and strong, rather distant, rounded longitudinal ribs, characteristic of Gabb's Dominican species, Phos costalus. The type, which Gabb never figured, is with the main Gabb collection in the Philadelphia Academy; but the writer in 1916, obtained specimens of the shell from Cercado de Mao, Santo Domingo, and identified them by comparison with metatypes of Gabb's at Cornell University. The Porto Rican rock imprint does not reveal the aperture as it is more a side view of the shell ; but the form, sculpture, and ribbing are so evidently that of this rugged Phos that there seems no doubt of its complete identity with the Dominican species. W. F. Jones reports PJios costaius from ihe ^laissade beds on Rio Blanco, north of Maissade, and also in the same formation on Riviere L'Avave, Haiti. Until then it had never been identified outside of the Dominican Republic. Its Miocene range is now extended to the neighboring island of Porto Rico. Like the following species this Plios is referable to the section Strongylocera Moerch which Dall has suggested might well be re- vived for these and kindred species. Locality. — Xear Quebradillas, Station 129. Phos fasciolatus Dall Pho'i ( Stronglocera) fasciolatus. Dall. 1S97. Proc. U. S. Nat. Mus XIX, No. 1110. p. 311, PI XXVIII. fig. 12. Ffio.s fasciolattts Maury. 1917, Bull. Amer. Paleont.. No. 29. p. 88. PI. XIV, figs. 15, Ifi; Idem. No 30. pp. 13. 27. Phrs fasciolatus Jones. 1918, Jour Geol. XXVl. p. 741. 'J'he type of Phos fasciolatus was collected by Bland at l^otrero, on tJKi Rio Amina, Santo Domingo, and dest-rihed by Dall. The writer obtained this species in abundance at (^aimito, Rio Cana, and it was also present at Cercado, Rio Mao, Santo Domingo, — oci-iii'i-iiiir at both localities in tlie Aplirni ishicoloiiis t'oi-niat ion. At MAURY, TERTIARY MOLLUSCA FROM PORTO RICO 63 Cereado it was associated with FJios costaius, and it is interesting to note that Jones has since found these two species associated in the ]Maissade beds, Riviere L'Avaye, Haiti. Among the Porto Rican fossils is a single external mold of a Phos apparently this species. The impression in the rock is very clear and sharp and represents the dorsal side of the shell. The whorls are convex, suture deep and the imprinted portion of the last whorl shows six even, rounded ribs. The ribs are closer than in Phos costatus and the shell is more regular and less rugged and robust. The characters of the aperture of the Porto Rican shell are not shown by the material in hand, but judging from the dorsal aspect of the mold and its artificial cast, it is the same as the Do- minican shells I referred to Phos fasciolatus Dall. This species is now reported for the first time beyond the limits of Santo Domingo and Haiti. In Porto Rico also it is found asso- ciated with Phos costatus. Locality. — Near Quebradillas, Station 131. Genus Alectrion Montfort External molds of two species of Alectrion from the Quebradillas limestone are perhaps identical with the Dominican Miocene forms, but they are too imperfect for definite determination. Genus Murex Linnaeus Murex messorius Sowerby Murex messorius Sowerby, 1840, Proc. Zool. Soc, p. 137. Murex messorius Reeve, 1845, Conch. Icon., Murex, fig. 90. Murex recurvirostris Gabb, 1873, Trans. Amer. Phil. Soc, XV, p. 201. Not M. recurvirostris Broderip. Murex messorius DaU, 1890, Trans. Wagner Inst. Sci., Ill, pt. 1, p. 139. Murex messorius Dall and Simpson, 1902, Bull. U. S. Fish Comm., XX, pt. 1, p. 407. Murex messorius Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci., Phila., p. 353. Murex messorius Maury, 1917, Bull. Amer. Paleont , No. 29, p. 101, PI. XVI, figs. 1, 2; Idem, No. 30, pp. 13, 21. Murex messorius Zetek, 1918, Los Moluscos de la Republica de Panama, p. 23. Murex messorius Vaughan, 1919, Bull. 103 U. S. Nat. Mus., p. 561. This is a typically static species of wide geographic and geologic range. Abundant in the Antillean Miocene, it is still living on unchanged in the West Indies, and extends from Cedar Keys west 64 SCIEXTIFIC SURVEY OF PORTO RICO to Texas and south to Colon, Panama. Recent shells ditfer in no respect from the fossil. The latter have been identified in the Gatun beds by Brown and Pilsbry ; in the Dominican Miocene bv Gabb (as M. recurvirostris) ; the writer has obtained them from Cercado and Los Quemados, Santo Domingo ; and Jones has listed the species from the Maissade beds, Riviere L'Ayaye, Haiti. The range is now extended eastward to Porto Rico. The Porto Rican representatives are fragmentary imprints of the exterior of the shell ; but the sculp- ture is so perfectly preserved and accords so exactly in every detail with that of Dominican Miocene specimens that the identification is conclusive. Murex messorius has been found by Dall and Simpson living at Mayaguez, Porto Rico. Other recent species of this island's fauna are Murex antillarum Hinds, M. rufus Lamarck, M. hrevifrons Lamarck, M. pomum Gmelin, M. micromeris Dall, M. cellulosus Conrad, and M. inter medius Adams. Localities. — Xear Quebradillas, Stations 130, 134, 135, Genus Marg^nella Lamarck Marginella coniformis Sowerby Marginella coniformis Sowerby, 1850, Quart. Jour. Geol. Soc. London, VI. p. 45. Marginella conformis Guppy, 18G6, Quart. Jour. Geol. Soc. London. XXII, p. 288, PI. XVII, fig. 2. Marginella coniformis Gabb, 1873, Trans. Amer. Phil. Soc , XV, p. 221. Marginella coniformis Guppy, 1876, Quart. Jour. Geol. Soc. London, XXXII, p. 528. Marginella coniformis Guppy and Dall, 1896, Proc. P. S. Nat. Mus., XIX, No. 1110, p. 309. Not Marginella coniformis Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila., p. 348, Pi. XXIV, fig. 12. Marginella coniformis Maury, 1917, Bull. Amer. Paleont., No. 29, p. 70, PI. XI, figs. 5, 5a. Marginella coniformis Vaughan, 1919, Bull. 103 U. S. Nat. Mus., p. 561. Several external molds of a Marginella show the characters of the Santo Domingan ]\rioc'cne species, M. coniformis Sowerby. On comparison with specimens of this species they are seen to agree in size, form, number and direction of the eolumellar plaits, and in the extension of the aperture to the apex of the spire. This species has been erroneously reported from Cumana, Venezu- ela. The specimen Guppy listed from there he later found to be MAURY, TERTIARY MOLLUSCA FROM PORTO RICO 65 M. cincta Kiener, while that from the Caroni Series of Trinidad is a Persicula near obesa. Its occurrence at Gatun also appears very doubtful, because the figure Pilsbry and Brown give of a Gatun specimen is very unlike Guppy's illustration of the Bowden shell, and unlike our specimens from the type locality, which was the Yaqui Valley, Santo Domingo. Moreover the figure of the Gatun shell distinctly shows that its aperture does not continue to the spire, whereas Sowerby says of the type of M. coniformis, "labii extern! margine inflexo, lato, crasso, depresso, ad apicem continuo." Whether Vaughan records this species in his 1919 list from an actual Gatun shell or from the Gatun report of Pilsbry and Brown I do not know. But it would seem that, as far as now known, Marginella coniformis was limited to the three neighboring Greater Antilles, — Santo Domingo, Jamaica, and Porto Rico. Localities. — Seiior Rabell's Ranch on Rio Guajataca, Station 112; near Quebradillas, Stations 131 and 134. Genus Mitra Marty n Mitra henekeni Sowerby Mitra henekeri Sowerby, 1850, Quart. Jour. Geol. Soc. London, VI, p. 46, PI. IX, fig. 5. Mitra henekeni Guppy, 1867, Proc. Sci. Assoc. Trinidad, pt. 3, p. 160. Mitra henekeni Gabb, 1873, Trans. Amer. Phil. Soc, XV, p. 219. Mitra henekeni Guppy, 1874, Geol. Mag. London, new ser., decade 2, I. p. 440. Mitra henekeni Guppy, 1876, Quart. Jour. Geol. Soc. London, XXXII, p. 528. Mitra henekeri Dall, 1903, Trans. Wagner Inst. Sci., Ill, pt. 6, p. 1583. Mitra henekeni Maury, 1917, Bull. Amcr. Paleont., No. 29, p. 74, PI. XII, figs. 5, 5a; Idem, No. 30, pp. 10, 19, 22, 26. Mitra henekeni Jones, 1918, Jour. Geol. XXVI, p. 741. The collection of Porto Rico fossils made by Dr. Reeds affords several incomplete external molds showing the characteristic sculp- ture of this handsome shell which is so common in the Miocene of Santo Domingo. One of the molds shows distinctly the three colu- mella plaits, the posterior two being well defined and sharp, the anterior nearly obsolete as described by Sowerby in 1850. The type was collected in Santo Domingo by Colonel Heneken. Later this species was identified by Guppy in the Caroni Series of Trinidad and the Jamaican Miocene. Dall lists it from Bowden, Jamaica; it was obtained by the writer from Los Quemados, Caimito and Cer- cado de Mao, Santo Domingo; Jones has lately found it in the 66 SCIENTIFIC SURVEY OF PORTO RICO Maissade beds, Riviere L'Avaye, Haiti, and its range is now ex- tended eastward to Porto Rico. Adult examples of Mitra heneJieni show a marked general re- semblance to the figure of M. antille^isis Dall of the living fauna, extending from Cape Lookout to the Island of Grenada. The recent shell is, however, much larger, attaining a length of 80 mm. against about 55 mm. in the fossil, and the living species has four strong columellar plications. The genus Mitra is well represented in the recent Porto Rican fauna and includes Mitra l)arhadensis Gmelin, M. nodulosa Gnielin, M. hanleyi gemmata Sowerby, M. straminea Adams, and M. micro- zonias Lamarck. Localities — iSTear Quebradillas, Stations 125, 130, 133. Genus Lyria Gray Lyria cf. musicina (Heilprin) Voluia musicina Heilprin, 1887, Trans. Wagner Inst. Sri.. I, p. 109, PI. XV, fig. 45. Lyria musicina Dall, 1890, Trans. Wagner Inst. Sci., Ill, pt. 1, p. 85. Lyria musicina Dall, 1915, Bull. 90 U. S. Nat. Mus.. p. 59, PI. IX, figs. 1, 4. A single imperfect internal mold of a Lyria in the collection shows traces of about five broad, longitudinal ribs on half of the last volution and a suggestion of a constricting band below the suture. On breaking away the part of the mold which filled the shell's aperture one sees very distinctly five or six very strong an- terior grooves, the first two oblique, and a group of about the same number of faint, posterior grooves. These grooves show where the mud filling the aperture pressed against the folds on the columella of the shell which has since been wholly dissolved away. The evidence gathered from the mold proves that it is not a filling of the very common Dominican Miocene shell, Lyria pulchella Sowerby. That has far more numerous and narrow ribs and the anterior plaits are not so heavy and are transverse. Obviously, this is a member of the Eocene Lyria costata Sowerby group, which in the American Oligocene to Miocene was followed by the Floridian species, Lyria musicina Heilprin. The last whorl of L. costata is much more elongated in proportion to its width than that of the Porto Rican mold. It could not be that species, but it corresponds very well with the form of Lyria musicina. Very probably the mold is iden- tical with that shell, but an exact identification is not possible. MAURY, TERTIARY MOLLUSCA FROM PORTO RICO 67 Jji/ria musicina is recorded from the Upper Eocene (Jacksonian) of Ocala, Florida. The upper Oligocene of Tampa Bay, and the lower Miocene of the Chipola marls, Florida. With it Dall found Lyria pulchella in the Tampa formation. Now it appears that just as the latter Antillean Miocene species had emigrated to Florida, so the Florida species had established an exchange colony in Porto Ilico. Locality.- — Near Aguada, Station 117. Genus Oliva Martyn Oliva cf. brevispira Gabb Oliva brevispira Gabb, 1873, Trans Amer. Phil. Soc, XV, p. 215. cf. Oliva giraudi Cossmann, 1913, Journ. de Conchyliologie, LXI, p. oG, PI. V, figs. 1, 4-8. Oliva brevispira Maury, 1917, Bull. Amer. Paleont., No. 29, p G8, PI. X, figs 10, 17; Idem, No. 30, pp. 11, 13, 24, 2G. Several internal molds of a medium-sized, very short-spired Olive, on comparison with shells of Oliva brevispira Gabb, are seen to have the same general form that a filling of one of these shells would possess. No positive identification can be made with the material in hand ; but the probabilities strongly suggest the identity of the Porto Kican molds with this Santo Domingan species. Oliva brevispira was obtained in great abundance on the writer's Dominican expedition at Cercado, Los Quemados and Caimito, and was identified by comparisons with Gabb's metatypes at Cornell University. A very similar and seemingly identical form has been described by Cossmann as Oliva giraudi from Martinique. Other- wise this very common Dominican shell has not heretofore been found outside of that island. This contrasts with the wide distribution of the much larger species, Oliva cylindrica described in 1850 by Sowerby from Hene- ken's Dominican collection, and closely allied to the recent Oliva reciicularis Lamarck. Sowerby's species is found in the Isthmian deposits and those of Cumana, Venezuela, and in the insular Tertiary beds of Trinidad (Caroni series), Barbuda and Jamaica (Bowden). Jones has also lately found it in the Maissade beds, Riviere L'Ayaye, Haiti. It has not yet turned up in the Porto Rican collection, but is to be expected in this fauna.- 68 SCIEXTIFIC SURVEY OF PORTO RICO The recent Olives living on the Porto Rican shores are Oliva recticularis Lamarck and Oliva carihceensis Dall and Simpson, Localities. — Near Ponce, Station 285 ; near Gnayanilla, Station 321. Genus Olivella Swainsoii Olivella muticoides (Gabb) Oliva muticoides Gabb, 1873, Trans. Amer. Phil. Soc, XV, p. 215. Olivella mutica variety muticoides Dall, 1890, Trans. "Wagner Inst. Sci., Ill, pt. 1, p. 45. Olivella muticoides Cossmann, 1913, Journ. de Conchyliologie, LXI, p. CO. Olivella muticoides Maury, 1917, Bull. Amer. Paleont., No. 29, p. 68, PI. XI, fig. 1; Idem, No. 30, pp. 24, 26. Olivella muticoides Jones, 1918, Jour. Geol. XXVI, p. 741. In the collection of Porto Rican fossils are a niunber of internal and external molds of a species of Olivella. Fortunately one of the latter is so oriented in the rock as to show the characters of the aperture. An artificial cast of this mold accords perfectly with shells of Olivella muticoides collected on my expedition to Santo Domingo at Los Qiiemados, Cercado de Mao and Caimito and identified by comparisons with Gabb's metatypes at Cornell Uni- versity. The Porto Rican imprints show perhaps a slightly more deeply channelled suture, but in his original description Gabb men- tions this deep channelling as a characteristic. It is carried to an extreme in the form 0. canaliculata Gabb which is a broader variety as Gabb's metatypes show. Olivella muticoides is closely allied and probably ancestral to the recent 0. mutica Say, in ]\Laya- guez Harbor, Porto Rico, and elsewhere in the Antilles and ranging along the mainland from Cape Hatteras to Trinidad and from Florida west along the Gulf coast to Texas. But the fossil shell is slenderer than the recent and shows a series of plications on the columella which are not concealed by the callus. A related fossil species is Olivella duplicata Conrad. Olivella boussaci CossiiKinn, fossil, from Martinique, has somewhat the general aspect of the Dominican shell but the aperture is lower. Jones has lately iden- tified Olivella muticoides in the Maissade beds, Riviere L'Ayaye, Haiti. It is now for the first time recorded outside of the Dominican and Haitian Republics. Localities. — Near Quebradillas, Stations, 12G, i;55, 144: near Ponce, Station 299. MAURY, TERTIARY MOLLUSC A FROM PORTO RICO 69 Genus Cancellaria Jjaniarck Cancellaria Isevescens Guppy Cancellaria Icevescens Guppy, 186G, Quart. Jour. Geol. Soc. London, XXII, p. 289, PI. XVII, fig. 12; 18G7, Proc. Sci. Assoc. Trinidad, p. 157. Cancellaria Iwvescens Gabb, 1873, Trans. Amer. Phil. Soc. XV, p. 239. Cancellaria Icevescens Guppy, 1874, Geol. Mag. London, p. 438; 1876, Quart. Jour. Geol. Soc, XXXII, p. 520. Cancellaria Iwvescens Maury, 1917, Bull. Amer. Paleont., No. 29, p. 64, PI. X, fig. 6. In the fossil Gastropod exhibit in the American Museum are specimens of Cancellaria Iwvescens Guppy, collected by Thomas Bland from the Bowden beds of Plantain Garden, St. Thomas, Jamaica. One of these is so like the figure of Guppy's type, which was also Jamaican, that it might pass for the original. The others show interesting variations in degrees of loss of sculpture on the body whorl. The shells are large and solid, the largest being 40 mm. X 21 mm. while the type was 45 mm. x 28 mm. Gabb identified shells which he collected over forty-five years ago in Santo Domingo. My party did not find this species there, but I figured one of Gabb's shells. It was only 25 mm. in height but otherwise bore considerable resemblance to Guppy's shell. CanceUaria laevescens portoricana, new variety Plate VII, Figure 10 Mold with finely cancellated spire and body whorl smooth, except for a few spirals at the base. It appears to be a dwarfed variety of Cancellaria Icevescens. It resembles Gabb's Dominican specimen of C. kvvescens but is still smaller and is only a third the size of Guppy's type. Locality. — ISTear Quebradillas, Station 130. Cancellaria, species indet. There are several fragmentary internal molds apparently of the later volutions of several species of Cancellaria. The molds show grooves representing in reverse the columella plaits, and the lirations marking the interior of the outer lip of this genus. Localities. — Near Quebradillas, Stations 126, 128. 70 SCIENTIFIC SURVEY OF PORTO RICO Genus Tereljra Biuguiere Terebra cirrus Dall Terebra (Acus) bipartita Sowerby variety cirrus Dall, 1895, Proc. V. S Nat. Mus., XVIII, p. 38. Terebra (Oxymeris) bipartita variety cirrus Dall, 1903, Trans. Wagner Inst. Sci., Ill, pt. 6, PI. LIX, fig. 28. Terebra cirrus Maury, 1917, Bull. Amer. Paleont.. No. 29, p. 25, PI. Ill, fig. IT; Idem, No. 30, pp. 12, 24, 27. The type of Terebra cirrus was described by Dall from Eio Amina, Santo Domingo, and in 1910 specimens were collected on the writer's expedition to that island at Caimito, Los Qnemados and Cercado de ]\Iao, — being in all cases from the Apheni idacolonis formation. In ornamentation Terebra cirrus resembles the Isthmian species T. gatunensis Toula but is easily distinguished from that shell by its biplicate columella which allies it to the Dominican group of Terebra bipartita Sowerby. Terebra gatunensis has a single, nearly obsolete plait on its pillar. A fragmentary imprint of a fossil Terebra was found by Reeds in Porto Rico. An artificial cast of this shows the ornamentation very perfectly and is an excellent match of Dominican specimens of Terebra cirrus. This appears to be the first fossil Terebra recorded from Porto Rico. The scarcity of the genus there con- trasts with the rich variety in the Miocene of the Yaqui Valley on the neighboring island of Santo Domingo where the writer obtained sixteen distinct specific forms. The recent fauna of Porto Rico comprises half a dozen Terebras of which Simpson has given brief descriptions in ^follusca of Porto Rico. Locality. — Near Pcmce, Station 285. Terebra sansebastiana, new species Plate VII, Figure 11 Specimen incomplete. Last whorl shows traces of about 15 longitudinal. nearly straight ribs, wider than their interspaces, surmounted by ribbed band about 2 mm., wide. Main ribs become slightly arched on penultimate whorl. Indistinct traces of spiral sculpture over entire surface. Height of fragment 23 mm., greatest diameter of flattened mold 16 mm., least dia- meter 6 mm. This species is founded upon a distorted internal mold fiom the Collazo shales of what seems to be a Terebra . perhaps allied to MAURY, TERTIARY M0LLU8CA FROM PORTO RICO 71 T. didocata Say. The fact that, this shell is a member of the odd fauna of the Collazo shales seems to be siifRcieiit excuse for naming such an imperfect specimen a new species. Locality. — Rio Collazo, near San Sebastian, Station 54. Genus Drillia Gray Drillia consors (Sowerby) Pleurotoma consors Sowerby, 1850, Quart. Jour. Geol. Soc. London, VI, p. 50. Pleurotoma consors Guppy, 1866, Quart. Jour. Geol. Soc. London, XXH, p. 280; 1867, Proc. Sci. Assoc. Trinidad, p. 159. Turris {Drillia) militaris Gabb, 1873, Trans. Amer. Phil. Soc, XV, p. 207. Not Drillia militaris Hinds, 1843, Proc. Zool. Soc, p. 38. Pleurotoma consors Guppy, 1876, Quart. Jour. Geol. Soc. London, XXXH, p. 527. Pleurotoma sp. aff. PI. alesidota (Dall) var. macilenta Toula, 1911, Jahrb. der k. k. Geol. Reichsanstalt Wien, LXL p. 506, PI. XXX, fig. 11. Not alesidota var. macilenta Dall, 1889. Drillia consors Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila., p 345. Drillia consors Cossmann, 1913, Journ. de Conchyliologie, LXL PP- 20-22, PI. n, figs. 8-14. Drillia consors Maury, 1917, Bull. Amer. Paleont., No. 29, p. 53, PI. VIII, figs. 15, 16; Idem, No. 30, p. 18. Drillia consors Jones, 1918, Jour. Geol. XXVL D- "-il- Drillia consors Vaughan, 1919, Bull. 103 U. S. Nat. Mus., p. 561. Several blocks of limestone contain fragmentary imprints which unquestionably represent the very beautiful and unmistakable form and sculpture of Drillia consors. The type of this fine species was collected seventy years ago by Colonel Heneken in the Yaqui Valley, Santo Domingo, and described by Sowerby ; it is now in the British Museum. Guppy identified this species in 1860, in the Bowden beds of Jamaica, and later it has been recorded by various authors from Gatun, Mindi, and Martinique. Gabb referred his Dominican specimens to the recent West Coast species, Drillia militaris Hinds, l)ut the resemblance is only a very general one. The fossil is very much more like the recent, deep sea, Barbadian shell, Drillia alesi- dota macilenta Dall, which may be its descendant. Toula first called attention to this interesting aflinity. It is not surprising to find that Drillia consors was also living in Miocene times on the Porto Rican shores. Localities.— Senor Rabell's Ranch, Rio Guajataca, Station 112 ; near Quebradillas, Stations 126, 131. 72 SCIENTIFIC SURVEY OF PORTO RICO Drillia diazica, new species Plate IX, Fig-ure 1 Shell elongate, turreted, very slenderly fusiform, the last whorl extending into a very long, straight, anterior canal; aperture narrow; suture linear, wavy; sub-sutural channel smooth to the eye, but with six to eight extremely fine, microscopic spirals, and with delicate but sharp, microscopic, deeply arcuate growth-lines. Beneath the sub-sutural channel the whorls are sculp- tured with sharply defined, regular, evenly spaced, rounded, longitudinal ribs, numbering five to half a volution on the ultimate and the same on the penultimate whorl. On each of the later volutions are four major spiral threads which are strongest on crossing the crests of the ribs. Between every two major spirals are three, very fine, minor spirals, visible only with a lens. The longitudinal ribs terminate abruptly where the last whorl narrows to form the canal, but the two sets of spirals continue to the end of the canal although they become less regular and weaker. Estimated length of shell with tip of spire complete 37 mm , greatest width 11 mm. The affinities of this species are clearly with the two Miocene Dominican forms, Drillia fusifonnis (Gabb) and D. cercadonis Maury, and with the Gatun Miocene form Drillia fusinus Brown and Pilsbry. Drillia fusiformis {Defrancia fusifonnis Gabb, Trans. Amer. Phil. Soc, XV, p. 209, 1873) has only about eight ribs, crossed by three major spirals. Drillia cercadonis has nine ribs, uniform spirals, and a smooth subsutural channel. The an- terior canal of the Porto Ilican fossil is also longer than that of these Dominican shells. The Gatun analogiie, Drillia fusinus has a very long canal, but the ribs are feeble, and appear obsolete on the later whorls. Locality. — Near Juana Diaz, Station 175. Genus Conus Linnaeus Cone-shells are present from several of the Porto Kican horizons, but all are in the form (.f internal molds and usually very imperfect. With the exception of two, it would be unwise to hazard even a comparison. Conus recognitus Gnppy Conus solidus Sowerby, 1850, Quart. Jour. Geol. Soc. London, VI, p. 45. Not C. solidus Sowerby, 1841, Proc. Zool. Soc, Conch. Illust., Cotms No. 76, PI. LVI, fig. 5G. Consus solidus Guppy, 1866, Quart. Jour. Geol. Soc, XXII, p. 287, PI. XVI, fig. 1. Conus recognitus Guppy, 18G7, Proc. Sci. Assoc. Trinidad, p. 171. MAURY, TERTIARY MOLLUSCA FROM PORTO RICO 73 Conus pyriformis Gabb, 1873, Trans. Amer. Phil. Soc, XV, p. 229. Not C. pyriformis Reeve, 1843, Conch. Icon. I, PI. XIII, fig. 70. Conus recognitus Giippy, 187G, Quart. Jour. Geol. Soc. London, XXXII, p. 527. Conus recognitus Dall, 1903, Trans. Wagner Inst. Sci., Ill, pt. 6, p. 1583. Conus recognitus Maury, 1917, Bull. Amer. Paleont. No. 29, p. 45, PI. VII, fig. 9; Idem, No. 30, pp. 12, 21, 26, 27. In the collection is an internal mold of a Conus of the same size as Dominican shells of C. recognitus Guppy, and showing the very characteristic pear-shaped or fig-shaped contours of that species. In all probability the mold is the filling of C. recognitus. My party found this species in the lower, or Aphera horizon in the Yaqui Valley, Santo Domingo. It is also at Bowden. The recent West Coast, Conus pyriformis Reeve is the descendant of migrants through the Isthmus. Locality.— Near Aguada, Station 122. Conus of. tortuosostriatus Toula Conus (Chelyconus) tortuosostriatus Toula, 1911, Jahrb. der k. k. Geol. Reichsanstalt Wien, LXI, p. 508, PI. XXXI, fig. 22. Conus (Hemiconus) tortuososti iatus Cossmann, 1913, Journ. de Conchylio- logie, LXI, p. 40, PL III, figs. 28, 29. Conus tortuosostriatus Maury, 1917, Bull. Amer. Paleont., No. 29, p. 41, PI. VI, fig. 9; Idem, No. 30, pp. 22, 24. Another internal mold from Aguada resembles in form the slender, high-spired, delicate, Isthmian shell, Conns tortuosostriatus Toula, and the Dominican forms that I collected and referred to that species. No positive identification is possible, but the Porto Rican mold is very probably of the same shell. Locality. — Near Aguada, Station 122. Order Opisthobranchia Genus Haminea Leach Haminea granosa (Sowerby) Bulla granosa Sowerby, 1850, Quart. Jour. Geol. Soc. London, VI, p. 51, PI. X, fig. 10. Bulla granosa Guppy, 1867, Proc. Sci. Assoc. Trinidad, pt. 3, p. 155. Bulla granosa Gabb, 1873, Trans. Amer. Phil. Soc. XV, p. 246. Bulla granosa Guppy, 1874, Geol. Mag. London, p. 437; 1876, Quart. Jour. Geol. Soc. London, XXXII, p. 518. Bullaria granosa Maury, 1917, Bull. Amer. Paleont., No. 29, p. 20, PI. Ill, fig. 10; Idem, No. 30, p. 27. Sowerby, in his description of the type of this species, which Heneken had collected in the Yaqui Valley, Santo Domingo, re- t 74 SCIENTIFIC SURVEY OF PORTO RICO marks on its resemblance in form to the recent Bulla hijdatis, since made the geno-type of Haminea Leach. Recent specimens Libelled Haminea hydatis in the American Museum, from the Mediterranean are certainlv verv like in form and size to the fossil species. These recent shells are the 11. hydatis, of most authors, but really //. navicula Da Costa. The true hydatis of Linnanis is oblong-oval and smaller while navicula is ventricose like the fossil species. But the resemblance there ceases. For the living shells are nearly or perfectly smooth while the most striking feature of the fossil is its strongly cancellated sculpture, which be- comes granose at the points of intersection of the subequal spiral and longitudinal ridges. Internal molds representing this species are common in the col- lection under examination. A number correspond perfectly in size and form to Sowerby's figured type. These molds retain the can- cellation of the original surface and some even show the granular points of intersection. There seems no doubt of their complete identity with the Dominican species. These measure about 18 mm. in length ; but there are some larger molds, about 25 mm. in length, which show the cancellated surface and seem to be only larger speci- mens. Possiblv more material will show them worthv of varietal rank. Many authors have quoted Haminea granosa irvin Santo Domingo, but the only definite locality known is Zone I at Caimito, Rio Cana, where they were collected by the IMaury expedition in 1916. The species has not been found heretofore outside of the Dominican Tiepublic, but the present collection shows it was also a common Porto Rican Miocene shell. Recent species of Haminea in Porto Rican waters are //. eler/ans Gray and //. succiiiea (\)nrad. These are both spirally striated, but have no longitudinal sculpture and are quite unlike the fossil species. Localities. — Near Aguada, Station 117, near Quc'i)radillas, Sta- tions 126, 128, 129, 130, 131, 133, 135, 136; near GuayaniUa, Station 321. Haminea quebradillica, new species Plate IX, Figure 5 External mold, ornamentation spiral but not cancellate. spirals sharply incised, lines of uniform strength, almost microscopic, about nine lines to MAURY, TERTIARY M0LLU8CA fJx'OM POh'TO RICO 75 5 mm. These alternate with perfectly flat bands. Occasionally, however, there is a wider band as if one line has been omitted. Altitude of incomplete mold 18 mm., diameter 13 mm. As far as the dorsal aspect of the mold and its artificial cast permits one to judge, this pretty and delicate species must have been most nearly akin to the recent Ilantinea elegans Gray, living in the Antilles and from Florida to Rio de Janeiro. Dall and Simpson record one specimen taken at Mayaguez Harbor, Porto Rico. In this recent species the entire surface is also ornamented with incised lines, but they are not uniform as in the fossil, and very fine lines are mingled with coarser, unequally spaced grooves. The even, delicate uniform lines distinguish the Miocene from the living shell. The other living Porto Rican species of this genus is the smaller, cylindrical shell, H. succiriea Conrad. Of fossil Hamineas a Martinique Miocene species has been de- scribed by Cossmann as Haminea ventripotens (Journ. de Conchy- liologie, LXI, p. 8, PI. I, figs. 8-11, 1913) which he likens in some respects to H. elegans. But the Martinique shell has a per- fectly smooth surface so that its resemblance to H. elegans and to this fossii is only one of general form. It is distingiiished from H. granosa Sowerby by its different sculpture and absence of raised cords. Locality. — Near Quebradillas, Station 125. Genus BuUaria Rafinesque BuUaria paupercula Sowerby Bulla paupercula Sowerby, 1850, Quart. Jour. Geol. Soc. London, VI, p. 52. Bulla paupercula Guppy, 1867, Proc. Sci. Assoc. Trinidad, pt. 3, p. 155. Bulla paupercula Gabb, 1873, Trans. Amer. Phil. Soc. XV, p. 246. Bulla paupercula Guppy, 1874. Geol. Mag. London, p. 437; 1876, Quart. Jour. Geol. Soc. London, XXXII, p. 518. Bulla paupercula Dall, 1890, Trans. Wagner Inst. Sci., Ill, pt. 1, p. 18. Bulla striata Dall, 1903, Trans. Wagner Inst. Sci., Ill, pt. 6, p. 1583. BuUaria paupercula Maury, 1917, Bull. Amer. Paleont., No. 29, pp. 18-19, PI. Ill, fig. 8; Mem, No. 30, pp. 12. 24, 26, 27. BuUaria paupercula Jones, 1918, Jour. Geol. XXVI, p. 741. From Ponce comes an internal mold of a BuUaria slightly larger than shells of B. paupercula Sowerby from the Yaqui Valley Mio- cene, Santo Domingo. In its form and size it is almost exactly like specimens in the American Museum labelled B. occidentalis Adams from the Pleistocene of North Creek, Florida ; but its spire is much 76 SCIENTIFIC SURVEY OF PORTO RICO more deeply sunken. In this very deeply involute spire it resembles B. paupercula and althoiigii a trifle larger, it is really akin to it rather than to the recent Antillean B. occidentalis. The Miocene species, B. paupercula and B. vendreyesiana Guppy, the recent B. occidentalis and the Pliocene to recent B. amygdala Dillwyn are all members of the group of B. striata Bruguiere and are often recorded as the latter species. The true Bidlaria striata is now restricted to the Mediterranean and Southern Europe, but during the Pliocene it had a much wider geographic range, as it has been found by Dall in deposits of that age in Florida and has also been recorded from synchronous beds in West Africa. This species has a characteristically somewhat hump-backed form due to its tapering at the upper end. Bulla amygdala is the recent American analogue of striata but the body is more cylindrical. It is living throughout the West Indies. Typical specimens are much larger, heavier and more solid than the Miocene ancestral shell, B. paupercula. The latter species is abundant along the bluffs of the upper Rio Cana and in general in the Aphera islacolonis formation, Santo Domingo. Dall lists Bidla '.striata from Bowden, Jamaica, which is probably the same as these. Jones has recorded B. paupercula from the j\Iaissade beds, Riviere L'Ayaye, Haiti. Its Miocene range is now extended to Porto Rico. Locality. — N"ear Ponce, Station 299. Genus Scaphander Montfort Section Bucconia Dall Bucconia reedsi, new species Plate IX, Figure 6 Internal mold distinctly and regularly spirally striated. On the dorsal side the striations are crossed by longitudinal ridges, more or less uniform and equidistant, but weaker than the spirals. Mold somewhat flattened in median dorsal region. Length of incomplete mold ;54 ram., estimated com- plete length approximately 40 mm., greatest width 28 mm. The species which seems most like this fossil is Scaphander (Bucconia) grandis Aldrich from the Jacksonian of J^unker Ildl, Louisiana, and the Ocala beds of Central Florida. This mold especially resembles Dall's figure'' of the Ocala shell and is less like Alrich's illustration of the type^'\ Bucconia grandis attained a "Trans. Wagner Inst. Sci., 1890, III, pt. 1, I'l. X, t'l?. <<. '"Geol. Survey Ala., 1886, Bull. No. 1, p. 35, PI. Ill, fig. 1- MAURY, TERTIARY M0LLU8CA FROM PORTO RICO 77 length of 66 mm. against about 40 mm. in the Porto Rican shell. Moreover, both Aldrich's and Ball's figures represent grandis with very close-set transverse striae, about 10 to a distance of 5 mm., while this mold shows only 6 to an equal distance. The ornamentation of the dorsal side suggests the sculpture of Haminea granosa Sowerby, but that is more decidedly cancellated and the shell typically is only one-third the size of the form now described. The imperfect state of preservation renders it impossible to de- termine positively the generic position of this shell ; but it is tenta- tively referred to Ball's section Bucconia, which he describes as a thin-shelled Sabatia without the body-callus. This harmonizes with the globose form, thin shell, and distinct spiral striation which dis- tinguish the Porto Rican species. As far as I am aware, this shell is wholly unique among Antillean fossils. Its unusual size and spiral sculpture at once differentiate it. The species is named in honor of Dr. Chester A. Reeds by whom it was discovered in 1915. There is a very rare living deep sea species, dredged by the Challenger at 500 fathoms, off the Philippines, and described by Watson as Scaphmider niveus. This is somewhat smaller and has more distant and weaker spirals than the Porto Rican fossil, but in its Bullaria-like form and large size it bears more resemblance to our species than any recent shell I know. Locality. — ISTear Ponce, Station 285. Fig. 1. Fig. 2. Fig. 3. Fig. 4. Fig. 5. Fig. 6. Fig. 7. Plate I Ostrea sansehastiana, new species; height 66 mm. Area (Scapharca) aguadica, new species; length 17 mm. Area, {Scapharca) guajatica, new species ; length 7 mm. Area (Scapharca) collazica, new species; length 28 mm. Area (Scapharca) sansehastiana, new species; length 18 mm. Ostrea herheyi, new species; height 36 mm. Ostrea collazica, new species; flat valve; height 79 mm. Scientific Survey of Porto Rico and the Virgin Islands VOLTTME III, Plate I Plate II Fig. 1. Ostrea collazica, new species; convex valve of a large in- dividual; width 95 mm. Scientific Survey of Pokto Kico and the Virgin Islands Volume III, Plate II Plate III Fig. 1, Peden sansehastianus, new species; height of shell 35 mm. Fig. 2. Pecten mesenticus, new species; height of shell 35 mm. Fig. 3. Amusium {Propeamusium) hollicki, new species; convex valve; height 12 mm. Fig. 4. Amusium {Propeam,usium) hollicki; flat valve of another individual; height 16 mm. Scientific Survey of Porto Rico and the Viu«in Islands Volume III, Plate III Plate IV Fig. 1. Pecten guajatacus, new species ; operculate valve ; approxi- mate length 45 mm. Fig. 2. Pecten guajatacus, new species; convex valve; height 55 mm. Fig. 3. Pecten sansebastianus laresianuSj new variety; height 31 mm. Fig. 4. Pecten guanicus, new species; height 24 mm. Fig. 5. Pecten camuycensis, new species; height 25 mm. Fig. 6. Crassatellites juanadiazus, new species; length 28 mm. Fig. 7. Crassatellites juanadiazu^, new species; viewed from an- terior end; height 16 mm. Scientific Survey of Porto Rico and the Virgin Islands Volume III, Plate IV Plate V Fig. 1. Spondylus lucasi, new species; height approximately 33 mm. Fig. 2. Thracia (Cyathodonta) reedsi, new species; height 20 mm. Fig. 3. Cuspidaria islahispanioloe Maury, length 8 mm. Fig. 4. Cuspidaria juaTiadiaza, new species; length 10 mm. Scientific Suuvey of Poirro Uico and the Vircin Islands Volume III, I'late V Plate VI Fig. 1. Tellina disparoides, new species; length 25 mm. Fig. 2. dementia rahelli, new species; normal form; height 39 mm., length 40 mm. Fig. 3. dementia rahelli, new species; crushed into triangular form by pressure and made more ventricose; height 43 mm., length 33 mm. Fig. 4. Cardium (T rig onio cardia) sandi-sehastiana, new STpecies; length 12 mm. Venericardia juncalensis, new species; height 29 mm. Venericardia collazica, new species ; length 34 mm. Venericardia rahelli, new species; length 20 mm. Phacoides (Here) quehradillicus, new species; height 21 mm. Callocardia riocollazica, new species; length 28 mm. Corhula collazica, new species ; right valve ; length 13 mm. Corhula collazica, new species; another individual; left valve 12.5 mm. Fig. 5. Fig. 6. Fig. 7. Fig. 8. Fig. 9. Fig. 10. Fig. 11. Scientific Survey of Porto Rico and the Viroin Islands Volume III, Plate VI Plate VII Fig. 1. Dentaliuni diazlcum, new species; length 19 mm. Lucinisca hoveyi, new species; from artificial cast of ex- ternal mold; length 11 nun. Tagelus (Mesopleura) huhhardi, new species; internal mold; length 35 mm. Chione quehradillensis, new species; from artificial cast of external mold; length 21 mm. Chione quebradilletisis guajatica, new variety; from arti- ficial cast of external mold; length 20 mm. Turritella poncensis, new species ; from artificial cast of external mold; height of fragment 28 mm. Turritella guanicensis, new species; height of fragment 22 mm. Cerithium utuadicum, new species; height of fragment 16 mm. Cerithium russelli arecihense, new variety; from arti- ficial cast of external mold; greatest diameter 11 mm. Cancellaria Icevescens portoricana, new variet}' ; from arti- ficial cast of external mold; greatest diameter 11 mm. Terehra sansehastiana, new species; height of fragment 22 mm. Cyprcea sancti-sehastiani, new species; height 29 mm. Cadulus poncensisj new species; length 7 mm. Fig. 2. Fig. 3. Fig. 4. Fig. 5. Fig. 6, Fig. 7, Fig. 8, Fig. 9, Fig. 10, Fig. 11, Fig. Fig. 12 13, Scientific Survey of Porto Rico and the Virgin Islands Vor-UMK III, Pi-ate VII Plate YIII Fig. 1. Campanile (Portoricia) laricum, new species; internal mold; height 33 mm., greatest diameter 45 mm. Eig. 2. Campanile (Portoricia) laricum, new species; largest in- ternal mold, somewhat flattened by pressure: height 80 mm. Fig. 3. Pyrula lioveyi, new species ; incomplete internal mold of a young shell; height 42 mm. Fig. 4. Turritella altilira culehrina, new variety; drawn from the actual shell with fine details of sculpture restored from its external mold; length of fragment 39 mm. Fig. 5. Turritella collazica, new species ; incomplete internal mold ; height 65 mm. Scientific Survey of Pouto Rico and the Viucin Islands VoiiUME III, Plate VIII Plate IX Fig. 1. Drillia diazica, new species; height of decollate shell 33 mm. Fig. 2. Colubraria juanica, new species; aboral aspect of shell; height 23 mm. Fig. 3. Colubraria juanica, new species; oral aspect of the same shell; height 23 mm. Fig. 4. Orthaulax aguadilllensis, new species; view showing the extreme shortness of the spire and the circular outline of the shell at the shoulder. Unique specimen, mineral- ized; greatest diameter 45 mm. Fig. 5. Haminea quehradillica, new species; from artificial cast of external mold; height 18 mm. Fig. 6. Scaphander (Bucconia) reedsi, new species; incomplete internal mold; greatest diameter 28 mm.; estimated length when complete 40 mm. Fig. 7. Pyrula hoveyi, new species ; fragment of spire and of body whorl. From an artificial cost of an external mold; height 23 mm. SciKN'TiFrc SuRVKV HF I'liiiid Kico Axn THE ViKciN Islands VOLUMK III, I'LATK IX NEW YORK ACADEMY OF SCIENCES SCIENTIFIC SURVEY OF Porto Rico and the Virgin Islands VOLUME Ill-Part 2 Tertiary Mollusca from the Lares District, Porto Rico — Bela Hubbard NEW YORK: Published by the Academy 1920 TERTIARY MOLLUSC A FROM THE LARES DISTRICT, PORTO RICO By Bela Hubbard INTRODUCTION The material here described was collected in making a study of the geology of the Lares District in the summer of 1916, under the auspices of the New York Academy of Sciences and the Insular Government of Porto Rico. The present paper is a part of the report on the geology of the Lares District, the completion of which has been delayed two years by the war. As the section dealing with the general geology and stratigraphy has not yet been completed, the present report, dealing with the Tertiary Palaeontology, is pub- lished in advance. It Avill be noted under the descriptions of the species, that full use has been made of the material collected by Dr. C. P. Berkey in 1914, by Dr. D. R. Semmes in 1915, and by Dr. A. K. Lobeck in 1916, and in some cases their specimens have been used for illustration. No attempt will be made here to describe the stratigraphy, but the following tabulation of the subdivisions* of the Younger Series of the north coast is given in explanation of the horizons referred to under the descriptions of the species. Younger Series San Juan formation (Berkey, 1915) Pleistocene to Recent. UNCONFORMITY I Quebradillas limestone (Berkey, 1915) Arecibo Group I Upper Oligoceue (Bowden) \ Los Puertos limestone (Hubbard, 1917) (Arecibo formation j Upper Oligoceue of Berkey, Pepino ^ Cibao Limestone (Hubbard, 1917) formation of Hill I Middle Oligoceue and Vaughan) I Lares formation (Hubbard, 1917) \ Middle Oligoceue San Sebastian shale (Berkey, 1915) Middle Oligocene unconformity Older Series (Berkey^ 1915) Cretaceous *These subdivisions were made and described by the writer in a short paper presented before the New York Academy of Sciences, Section of Geology and Mineralogy, m February, 1917. 80 scientific survey of porto rico Acknowledgments Acknowledgments are made to Seiior Xarciso Rabell, of San Sebastian for aid in collecting fossils and for the interest which he took in the field work ; to C. J. Maury for the opportunity of com- paring some of the material with hers; to Miss A. L. Hepburn, Columbia University Library, for valuable assistance in securing the many necessary but scattered palaeontological references ; and to Dr. A. W. Grabau for advice and assistance in describing many of the species. FOSSIL LOCALITIES- Field Numbers. 1. Lares road, north side, near K. 38 San Sebastian shale. 7. On the cart road 1.75 miles north-east of Lares. Lares limestone. 8. About 0.5 mile north of No. 7. Lares limestone. 10. About 1 mile north-east of No. 8. Lares limestone. 23. About 5.5 miles north of Lares, and about 2 miles north of Central SoUer. Echinoid zone of Cibao limestone. 24. About 6.5 miles north of Lares, on cart road to Camuy. at southern boundary of the belt of pepino hills. Base of Los Puertos limestone. 41. On the cart road to Camuy, 0.25 mile south of the village of Cienega, near the Central Alianza. Quebradillas limestone. 42. On the cart road to Camuy, 0.25 mile north of the Central Alianza. Quebradillas limestone. 52(a). On the cart road to Collazo 1.75 miles south of Quebradillas, and a short distance north-west of the village of San Antonio. Quebradillas limestone. 59. Valley of the Guajataca River, 1 mile south of Planas. Los Puertos limestone. 61. On the upland surface near the valley, and 0.3 mile south of No. 59. Los Puertos limestone. 62. About O.G mile south of No. 61, on the lowland just south of the belt of pepino hills. Cibao limestone. 64. About 0.3 mile south of No. 62, on cart road to Collazo. Cibao limestone. 65. On the cart road 2 miles north-east of Collazo. Lares limestone. 72. Lares road at K. 2S.G. San Sebastian shale. 74. Lares road at K. 29.3. San Sebastian shale. 75. Lares road at K. 30.4. San Sebastian shale. 76. Lares road at K. 32.9. San Sebastian shale. 83. Collazo Falls section, at top of second falls below the bridge. San Sebastian shale. HUBBARD, TERTIARY MOLLUSC A FROM LARES DISTRICT 81 84. Collazo Falls section, at base of second falls below the bridge. San Sebastian shale. 86. Collazo Falls section, at top of fourth falls below the bridge. San Sebastian shale. 89. Collazo Falls section, below the sixth falls below the bridge, in the lowest horizon exposed in the section. San Sebastian shale. 92. Lares road at K. 28.8, in ledge 12 feet above the road. San Sebastian shale. 100. Talus at base of the cuesta north of Lares. Lares limestone. 118. On the upland surface 0.25 mile north-west of Lares plaza. San Sebastian shale. 120. About 180 yards north-west of No. 118. San Sebastian shale. 126. Lares road, near K. 37. San Sebastian shale. 140. Base of the cuesta on west side of the Camuy River valley at the fork of the cart road to Utuado and the new road to Arecibo San Sebastian shale. 142, Cuesta at No. 140, in the limestone cliff overlying the basal shale. Lares limestone. 158. On cart road to Camuy, about 500 yards north-west of K. 19.5 of the new Arecibo-lares road. Cibao limestone. 161. About 500 yards north-west of No. 158, on the same cart road. Cibao limestone. 162. About GOO yards north-west of No. 161, on the same cart road. Echinoid zone, Cibao limestone 165. Near the point of emergence of the subterranean Camuy River. Cibao limestone. 167. A short distance north of No. 165, on the west side of the Camuy j^jygj. Cibao limestone. 174. About 0.5 mile north of No. 167, west side of the Camuy River on cart road to Camuy. Los Puertos limestone. 176. About 0.75 mile north of No. 174, on the same road. Los Puertos limestone. 179. About 0.4 mile north of No. 176, on the same road. Los Puertos limestone. 182. About 0.6 mile northwest of No. 179, and about 0.5 mile south of the ford where the cart road crosses to the east side of the Camuy River. Los Puertos limestone. 191. On the automobile road between Camuy and Quebradillas, near K. 40.5. Quebradillas limestone. 192. On the same automobile road, near K. 40. Quebradillas limestone. 198. On the automobile road west of Quebradillas, near K. 28.7. Quebradillas limestone. 204. Sea cliffs at the east mouth of the American Railroad tunnel at the mouth of the Guajataca River, north coast. Quebradillas limestone. S2 SCIEXTIFIC SURVEY OF PORTO RICO 20.J. In the railroad cut SO yards west of the railroad bridge over the Guajataca River, and 450 yards east from the atitomobile road grade crossing. Qtiebradillas limestone. 209. Guajataca River valley, west of the village of San Antonio. Quebradillas limestone. 21G. About 1.7.5 miles south of K. 14.7 (Isabela-Aguadilla automobile road), on cart road near Arenales bajos. Quebradillas limestone. 217. Nearly 0.5 mile south of No. 216. Quebradillas limestone. 220. Near Arenales altos, about 2.5 miles south of No. 217, and 150 yards east of the cart road. Los Puertos limestone. 225. On the cart road near Robles, 25 feet below the summit of the cuesta. Top of the Lares limestone. 226. Zone about 90 feet below No. 225. Lares limestone. 227. A 40 foot exposure of marl on the cart road south of No. 226, and 250 feet below the summit of the cuesta. Lares limestone. 229. About 1 mile north of K. 17.5 (Lares road), on cart road near Hato arriba. Base of Lares limestone. 2:J0. A short distance south of No. 229, and about 50 feet lower in elevation. Top of San Sebastian shale. 231. About 100 yards south of No. 230, at a slightly lower elevation. Coral zone of upper San Sebastian shale. 232. About 200 yards south of No. 231, and 1,350 yards north of K. 17.5 (Lares road), at an approximate elevation of ISO feet above the road. A five foot exposure of marl marking the dementia dariena zone. San Sebastian shale. 232(b). A fifteen foot exposure immediately overlying No. 232, and remarkable for the abundance of TiirriteJla tornata. San Sebastian shale. 233. On a small secondary cuesta a short distance south of No. 232. San Sebastian shale. 234. South of No. 233 on the same cuesta. San Sebastian shale. 235. South of No. 234, and 1,050 yards north of K. 17.5 (Lares road) at an elevation of 180 feet above the road. San Sebastian shale. 236. South of No. 235, and 900 yards north of K. 17.5. San Sebastian shale. 237. South of No. 236, and 820 yards north of K. 17.5, at an elevation of 100 feet above the roatl. San Sebastian shale. 244. On the cart road to Camuy, 1 mile north-e;ist of Lares, in limestone quarry near summit of the cuesta. Lares limestone. 254-5-6. Talus from the cuesta north of Lares. Taken from the road alon.s the Guajataca River, about 0.75 mile north of Lares plaza. Lares limestone. 261. Collazo Falls section, at base of second falls helow the bridge, and overlying No. 84. San Sebastian shale, 262. Collazo Falls section, at toi) of third falls below ihe bridge. San Seltastian shale. HUBBARD, TERTIARY MOLLUSCA FROM LARES DlHTHlCT b3 263. Collazo Falls section, half way between top and bottom of the fourth falls below the bridge. San Sebastian shale. 264. Collazo Falls section, immediately underlying No. 263. San Sebastian shale. 2G5. Collazo Falls section, at base of the fourth falls below the bridge. San Sebastian shale. 267. Collazo Falls section, near the base of the section, overlying No. 89. San Sebastian shale. 268. Collazo Falls section, in the red sandstone at the sixth falls below the bridge, and overlying No. 267. San Sebastian shale. 280. About 880 yards north of Moca on cart road. Lares formation, shale facias. 281. About 1,100 yards north of Moca on cart road. Lares formation, shale facies. 282. About 1,150 yards north of Moca on cart road. Lares formation, shale facies. 287. About 1.25 miles north on Moca on cart road. Top of the Cibao limestone. 313. Sea cliff 0.75 mile northeast of the Point Borinquen Light House, from a stratum 55 feet above sea level. Quebradillas limestone. 314. The same cliff, in a stratum 10 feet above sea level. Quebradillas limestone. 317. About 1.25 miles north of K. 2 (Aquadilla-Isabela automobile road), on cart road, 350 yards north of the American Railroad grade crossing. Quebradillas limestone. 318. A short distance south of No. 317, in railroad cut, about 200 yards north- east of grade crossing. Quebradillas limestone. 319. In quarry east of cart road, and about 0.6 mile north of K. 2 (Aguadilla- Isabela road). Quebradillas limestone. 324. Small limestone outlier about 300 yards south of K. 1.7 (Lares road). Lares limestone. 330. Southwest of bridge over Culebrines River near K. 2 (Aguadilla-Aguada automobile road). Lares limestone. 331. Aguadilla-Aguada road, south of sugar railroad crossing near K. 2. Lares limestone. 332. Same road, about 300 yards southwest of No. 331. Lares limestone. 333. On automobile road to Rincon, 1,150 yards west of Aguada. Exposure on north side of road. Lares limestone. 334. Same road, 100 yards west of No. 333. Lares limestone. 335. Same road, 290 yards west of No. 334. Lares limestone. 343. Same road, 640 yards northeast of bridge over the Rio Grande. Lares limestone. 344. Same road, south of No. 343, and 460 yards north of bridge over the Rio Grande. Lares limestone. 84 SCIENTIFIC SURVEY OF PORTO RICO 370. Exposure in sea cliff on south side of the American Railroad track at Point Jiguero, 930 yards northeast of the lighthouse. The San Juan formation may be seen here, plastered on the Tertiary limestone. Lares limestone. 616. Uppermost stratum in quarry 0.5 mile south-east of Auguada, on south side of road, and northeast of the American Railroad grade crossing. Lares limestone. 619. Same as No. 616, but from lowest exposure in the quarry. Lares limestone. 621. Exposure on south side of railroad tracks, about 580 yards south-east of the Aguada railroad station. Lares limestone. 664. East side of Camuy River valley, 2.25 miles southeast of Camuy. Quebradillas limestone. 670. East side of Camuy River valley, 1 mile southeast of No. 664. Quebradillas limestone. 671. Same locality as No. 670, but from a stratum a few feet above the latter. Quebradillas limestone. 678, East side of Camuy River valley, about 1.5 miles due south of No. 671, and about 0.75 mile north of No. 182. Quebradillas limestone. 749(a). On cart road, about 0.5 mile north of bridge at Collazo. Lares limestone. 757. One mile northeast of Lares, on the cuesta, in quarry near No. 244. Lares limestone. 758. Same locality as No. 757, but at a slightly higher horizon. Lares limestone. 800. About 100 yards south-east of No. 24. Top of Cibao limestone. 801. In railroad cut at east end of railroad bridge over the Guajataca River, north coast. This locality is about 50 yards east of No. 205. Quebradillas limestone. The following- localities are recorded from field notes of Dr. C. P. Berkey. 10. North of the military road in the vicinity of Carolina. Quebradillas limestone. 77. Military road 1.5 kilometers west of Ponce. Upper Ponce formation. (Corresponds with the Quebradillas limestone.) 78. Three kilometers west of Ponce. Upper Ponce formation. 95. Near K. 28 Lares road. San Sebastian shale. 96. A short distance west of Lares on the Lares road. San Sebastian shale. 98. In the railroad cut near the bridge over the Guajataca River, north coast. Quebradillas limestone. 176. Vicinity of Moca. Lares formation, shale facies. HUBBARD, TERTIARY MOLLUSCA FROM LARES DISTRICT 85 syst™atic description of species Genus Atrina Gray Atrina rabelli, new species Plate X, Figure 1 Margins of shell are arcuate strongly folded upward, particularly the antei-ior and posterior portions; central convex zone with flat lateral slopes, the cross-section of the valve being broadly V-shaped; surface marked by small, regular concentric ripples which become obsolete on the lateral mar- gins; central convex zone marked by two low radial ribs which die out near the ventral margin. Length 71 mm., semi-diameter 15 mm., height 100 mm. A single external mold was found showing only the ventral half of the shell. It is a large, rapidly widening form of the type of A. chipolana Dall, but is distinct from any form yet described. This shell is named in honor of Senor Rabell, geologist, of San Sebastian, who has taken an active interest in the geological work of the IST. Y. Academy of Sciences and rendered every possible assistance to the geologists in their field work. Localitij. — 204. Horizon . — Quebradillas limestone. Genus Pecten Miiller Pecten (Pecten) laresense, ne^v species Plate X, Figures 2, 3 Shell small of the concavo-convex type; right valve extremely convex, left valve shallow concave; shell equilateral; ears small and strongly ribbed radially; right valve with twenty-seven square smooth ribs and nearly flat interspaces of equal width, marked by fine, sharp, raised concentric threads which rise on the sides of the ribs; left valve with similar ribs and inter- spaces but with ribs medially grooved distally; ears of left valve with imbri- cate radial threads; anterior ear of left valve concave and correspondingly convex in the right; byssal notch small, ctenolium apparently lacking; sub- margins on both valves absent, the ribs extending to the base of ear; interior grooved and Urate ventrally; cardinal crura obscure and irregular; small amorphous pair of auricular crura; provinculum distinct; adductor scars markedly posterior. A right valve measures in length 20 mm., height 17 mm., diameter 8.5 mm.; left valve measures in length 20 mm., height 17 mm., concave diameter 2 mm. This shell is distinct from any other Pecten in the Porto Rican fauna. 86 SCIENTIFIC SURVEY OF PORTO RICO Type localities.— 2?>'[, 232, 233. Other localities.— 2S2 (b), 126, 118, 236, 1. Hoi'izon.—San Sebastian shale. (abundant and widely dis- tributed.) Pecten (Euvola) reliquus Brown and Pilsbry variety portoricoensis, new variety Plate X, Figure 4 Only right valves were found, whicli agree closely with the speci- mens of P. reliquus in the (^)lumbia University Gatun collections. The Porto Rican shell differs in having fewer, higher ribs with narrow interspaces, and in having the lateral slopes faintly ribbed instead of smooth as in the Gatun shell. The most perfect right valve has fourteen strong rounded ribs with several faint ones on each lateral slope. Length 35 mm., height 34 mm., diameter 3 mm. Fragments were collected showing a size of at least 50 mm. The left valves seem to have been easily destroyed, as none were found in the Panama collections. Ti/pe locality. — 165. Other localities.— ICA, 198. Range. — Cibao limestone to (^uebradillas limestone. Pecten (Nodipecten) nodosus Linne Plate X, Figure 5 Pecten nodosus Linne, Syst. Nat., No. 1G4, 1758. Pecten. magnificus Gabb, Trans. Amer. Phil. Soc, XV, p. 25G, 1S73 (not of Sowerby). Pecten {Nodipecten) nodosus Dall, Trans. Wagner Inst. Sci., Ill, pt. 4, p. 717, 1898. Pecten (Nodipecten) nodosus Maury. Bull. Amer. Paleont., V, No. 29, p. ISO, 1917. The specimens show three prominent strongly nodose ribs, one in center of valve and two lateral. These alternate with two lower, sub- jiodose ribs marked* by a median and two lateral threads. The subnodose ribs break np into scaly tliroads adjacent to the snb- margins. The most perfect sjxMMnieii meiisnves in length I'U nnn.. HUBBARD, TERTIARY M0LLU8CA FROM LARES DISTRICT 87 height :-^S iiiin., scini-diauu'icr 7 iinn., but fragments were found showing a height of 40 nun. Tyjoe locality. — 198. Other locality. — 42. Horizon. — QiiebradilUis limestone (not abmidant). Pecten (Chlamys) collazoensis, new species « Plate XI, Figure 1 Shell small, slightly convex; surface with twenty-one to twenty-two rounded ribs and narrower rounded interspaces; ribs ornamented with fine, thin, erect scales extending the whole width of the rib, and on either side at juncture with the interspace, a row of minute, densely crowded, bead-like spinules, which are much smaller and more densely crowded than the scales on tops of the ribs. Submargins and ears sculptured with a continuous series of squamose radial threads; anterior ear (of right valve) longer than the posterior ear, and with deep byssal notch. Length 11.5 mm., height 12. .5 mm., semi-diameter 2 mm. j^o well preserved left valves were found. This shell is apparently the progenitor of P. 'portoricoensis. Localities.— Collazo Falls, 232, 233 ? Horizon. — Lower? San Sebastian shale, (some doubtful specimens were found in the upper part of the San Sebastian shale.) Pecten (Chlamys) portoricoensis, new species Plate XI, Figures 2, 3 The typical form is a moderate sized shell (length 24.5 mm., height 26.5 mm., diameter 9 mm), equivalve, sub-equilateral; ears moderately large, un- equal (anterior larger) ; byssal notch large but not deep; ctenolium incon- spicuous; single pair of cardinal crura; valves grooved within, and scalloped and lirate at ventral margin; surface with twenty-six to twenty-seven rounded ribs and slightly narrower, flatly rounded interspaces; ribs ornamented with one to three rows of fine spinose scales which in the young shells show a tendency toward the beaded ornamentation of P. collazoensis. This feature usually persists longer next the submargins than centrally. Following this P. collazoensis stage, the ribs and interspaces become smoother, the latter relatively broader; a medium thread or keel appears on each rib, and on this keel the small scales become narrower, more spinose and more widely spaced. When the shall is half grown, lateral keels, similarly spinose, appear low down on the sides of the ribs. These correspond in position with the lateral beads of the P. collazoensis stage but there is always a gap between 88 SCIENTIFIC SURVEY OF PORTO RICO the two where the sides of the ribs are smooth. Finally, in the adult, an interstitial thread, ornamented like the keels, appears in each interspace. In the last stage of some adult specimens an additional pair of spinose lateral keels appears one on each side of the median keel. On the lateral slopes next the submargins the ribs break up into densely squamose threads, which, decreasing in size, continue in unbroken series across the submargins and the ears. The right anterior ear differs from the others in having a strongly rugose area adjacent to the submargin, and above this the radial threads are strong and coarsely squamose. Camptonectes striation is very plain on un- worn specimens. This is an extremely variable shell, apparently related to P. tJie- tides Sowerby. It is the most abundant Peden in the Tertiary of north side of the island and numerous specimens show intergrada- tions between the different varieties, thus fortunately preventing the error of making two or more species. The typical forms are es- pecially interesting in that they show the species to have been de- rived from P. (Chlamys) collazoensis, which was not found in hori- zons above the San Sebastian shale. P. portoricoensis ranges from the San Sebastian shale through the Lares limestotie, and has been found by Dr. Berkey in the Ponce limestone on the south side of the island. This shell is obviously derived from P. collazoensis, and both might conceivably be included under one specific name. The latter, how- ever, retains its typical ornamentation throughout, without the slightest tendency toward a change, whereas in P. portoricoensis the P. collazoensis stage has become so accelerated that it is unnoticeable in some specimens and absolutely lacking in variety reticulatis. Type localities.— 234:, 235, 343. Other localities.— 232, 118, 261, 1, 332, 76, and Collazo Falls, 244. Stratigrapliic range. — Throughout the San Sebastian shale, and in the lower Lares limestone also found by Dr. Berkey in the Juana Diaz shale. Pecten (Chlamys) portoricoensis, new species variety reticulatis Plate XI, Figure 4 Shell, while smaller than the type, is similar in every respect, except that In the young the ribs are crossed by fine, sharp, densely but evenly spaced HUBBARD, TERTIARY M0LLU8CA FROM LARES DISTRICT 89 concentric lamellae, which thicken slightly on crossing the ribs, the whole producing a reticulate ornamentation. This feature is most noticeable cen- trally, since adjacent to the submargins are the typical scales, which do not appear until later on the ribs of the central portion of the shell. The reticu- late ornamentation persists until the shell has reached one-quarter or one- third adult size, and is then replaced by the typical spinose scale. The lateral spinose keels and the interstitial threads do not appear until the shell has reached maturity. Average size specimen measure in length 1G.5 mm., height 18 mm., diameter 6 mm. The largest specimen measures 24 mm. in height. Type localities. — 343, 619. Other locality. — 74. Range — San Sebastian shale and Lares formation. Pecten (Chlamys) portoricoensis, new species variety grandis Plate XI, Figures 5, 6 This shell differs from the type and from variety reticulatis chiefly in its much greater size, and, except in gerontic individuals, in its relatively flatter valve. Unfortunately the specimens are all more or less worn in the umbonal region, but the reticulatis orna- mentation seems to predominate in the young. Some specimens, however, show in the young shell in P. collazoensis ornamentation, especially near the submargins. In some individuals the interstitial thread does not appear until the last stages of growth. In others it appears early but is not spinose until the adult stage is reached. In still others the ribs remain unornamented until the adult stage, though this may be due partly to wear. In the largest shells there are five threads on the ribs in the last stages of growth, and two additional threads sometimes appear in the interspaces. The camp- tonectes striation is obscured by the growth lines, which are es- pecially strong in this variety. A large flat-valved specimen measures in length 36 mm., height 39 mm., semi-diameter 8.5 mm. In this specimen the ribs are low and broad. A gerontic shell from the San Sebastian shale measures, in length 30 mm., height 32 mm., semi-diameter 8.5 mm. These geron- tic shells show a perodic swelling and contraction of the ribs and are 90 SCIENTIFIC SURVEY OF PORTO RICO notably convex. Superficially, they look like another species, es- pecially Avhen the ribs are worn smooth. Type localilies. — 7G, and K. 29.2 Lares road. OtJier localities.— 158, 229, 203, 72, 344, 233, 263 (bottom 3rd falls), 757, 165?, Collazo Falls, and vicinity of Collazo (float), 140, 1, 261, 244. Range. — San Sebastian shale. Lares limestone, Cibao limestone. Pecten (Chlamys) grabaui, new species Plate XIT, Figure 1 Shell transversely sub-ovate, equivalve, moderately convex with the greatest convexity toward the umbones; length 27 mm., height 26.5 mm., diameter 11.5 mm.; umbonal third of shell smooth, remainder with twenty- three low rounded ribs and slightly narrower shallow rounded interspaces; following the smooth stage appear faint regular widely spaced raised con- centric threads, strongest on the ribs, which become more distinct as the ribs increase ventrally. These threads then become irregular, break up into three rows of fine sharp scales on the ribs. The ribs become threaded and grooved distally, the scales giving a squamose character to the threads. There are as many as five of these longitudinal threads on the distal portions of some ribs. Likewise, in the interspaces an interstitial thread appears which is similarly squamose. Thus in the final stage of growth there is a typical Chlamys ornamentation. On the lateral slopes the ribs break up into .squamose radial threads, which continue across the submargins and ears: ears unequal, the anterior ears being longer and more strongly sculptured; byssal notch deep; ctenolium fine; double pair of cardinal crura; auricular crura low, indistinct; camptonectes striation strong. In first going over the specimens it seemed as though three or four species were represented, but intermediate forms made it neces- sary to group them as variations of an exceedingly variable species, similar to P. (Chlamys) indecisus Dall, a form from the Vicksburg Oligocene. Like the latter, this shell shows variations which arc intermediate lietween a true Chlamys and an Amusium. As the type of the species, that form is chosen which best illustrates the greatest number of characters common to the group. Type localities. — 343, 619. Horizon. — Lares formal ion. HUBBARD, TERTIARY MOLLUSC A FL'OM LARES DISTRICT 91 Pecten (Chlamys) grabaui, new species variety aguadensis Plate XIT, Figure 2 Shell moderately convex in umbonal region, nearly flat ventrally; surface nearly smooth, but with obsolete ribs visible chiefly as orange colored rays in the epidermis; camptonectes striation distinct; concentric lamellae, char- acteristic of the group, visible only under a lens. They are frequently worn off, leaving microscopic impressed lines in their place; submargins radially striate; ears rather strongly radially sculptured as in the other varieties; interior of shell strongly Urate, the lirae being paired and corresponding with the juncture of rib and interspace on the exterior. The interior is otherwise the same as in the other varieties: ctenolium distinct; length 18.5 mm., height 18.5 mm., semi-diameter 3.5 mm. This interesting variety is a typical Chlamys in shai)0 but has many of the characters of Amusium. It seems to be the primitive member of this group, since the other varieties pass through the smooth aguadensis stage more or less early in growth and develop the ribbed stage. Localities.— Q19, 333, 331. Horizon. — Lares formation. Pecten (Chlamys) grabaui, new species variety hatoensis Plate XII, Figure 3 Ribs appear very early, and become high and rounded; concentric lamellae regular and widely spaced, but becoming irregular near ventral margin where the ribs are radially grooved and interstitial thread appears in each interspace; submargins radially striate; ears strongly sculptured; ctenolium distinct; crura as in the type form; lirae obsolete except distally. Length 21.5 mm., height 21 mm., diameter 8.5 riim. Some specimens do not. reach the high ribbed stage, but ribs are rather low and rounded throughout, with the concentric lamellae crowded ventrally and showing a tendency to alternate in strength. Another specimen from a higher horizon and which may be a muta- tion from this variety, has high, strong ribs, in which the irregularity of the lamellae, grooving of ribs, and interstitial threads appear much earlier in the growth of the shell, and in the interspaces shows as many as thee interstitial threads in the final adult stage. 92 SCIENTIFIC SURVEY OF PORTO RICO Type localities.— dS2, 225. Other localities.— Md, 229, 333, 287, 167?, 62. Range. — Lares formation to Cibao limestone. Pecten (Chlamys) grabaui, new species variety gnayabensis Plate XII, Figure 4 Shell marked by smooth submargins. The specimens can be arranged into a series, as in variety hatoensis, varying from low to high, strong, rounded ribs, and with concentric lamellae varying from simple and distinctly spaced, to irregular and crowded. As in variety hatoensis. the ribs appear early, covering the entire shell except the prodissoconch. Interior of the shell faintly Urate throughout. Shell otherwise like the type form of the group. The largest specimen measures in length 17 mm., height 17.5 mm., semi- diameter 4.5 mm. Type localities.— Q2, 332. Other locality.— 2S2. Range. — Lares formation to Cibao limestone. The probable relationship of the group is shown in the following table. Table 1 Pecten (Chlamys) grabaui, new species Horizon Cibao limestone Lares formation San Seba.stian shale Group with smooth submargins variety (juaynbensis (shell ribbed) variety aguafictusis (.shell smooth) Group with striate submargins variety hatoen.tis (shell ribbed) P. grabaui (type form) (partially ribbed) HUBBARD, TERTIARY MOLLUSCA FROM LARES DISTRICT 93 Pecten (Chlamys) hodgii, new species Plate XII, Figure 5 Shell sub-ovate, sub-equilateral, strongly convex in umbonal region, slightly convex in ventral region; ears sub-equal, deep byssal notch and strong ctenolium; surface with twenty rounded ribs and deep, rounded, narrower interspaces; both ribs and interspaces grooved with fine radial threads which are densely spinose; ears and the rather broad submargins marked by sub- spinose radial threads; inner margin strongly scalloped and interior grooved. In the early stages of growth the ribs are not radically threaded, and the ornamentation consists of closely spaced, minute, sharp trans- verse scales on the ribs which join with raised concentric lamellae in the interspaces. When the shell has reached one third full size, the radial threads appear and the scales break up into minute densely spaced spines, which, ventrally, become elevated, curved, and U- shaped with concave side toward the beaks. Throughout most of the early stages the interspaces are devoid of threads and spinules, which appear much later than they do on the ribs. The concentric lamellae become obsolete ventrally. This shell is apparently closely related to an unlabelled specimen in the Kemp collection from Gatun, which seems to be a new species. In the latter the transverse scale stage is retained longer, the con- centric lamellae are noticeable throughout, and the spinules are not long or conspicuous as in the Porto Rican shell. If they are related, the Gatun shell would seem to be the more primitive of the two. It may be related to P. oxygonum canalis and P. oxygonwm optimum. of Brown and Pilsbry and to P. gahhi Dall. Dr. Maury h-as de- scribed a similar and apparently related shell P. camuycencis from the Quebradillas limestone. One complete right valve and some frag- ments were found. This shell is named in honor of Dr. Edwin T. Hodge, who studied the geology of the Coamo District, Porto Eico. ' Locality. — 314. / Horizon. — Quebradillas limestone (rare). Pecten (Aequipecten) lobecki, new species v*^^^***^''' Plate XIII, Pigure 1 "*«-^.^-- Shell sub-orbicular, equilateral, moderately convex, ears large, smooth, apparently sub-equal (portion of anterior ear concealed); shell sculptured with fourteen strong rounded ribs and sub-equal rounded interspaces; con- 94 SCIENTIFIC SURVEY OF PORTO RICO centric sculpture of fine raised lamellae, strong in the interspaces, obsolete or. tops of the ribs; fine concentric striae in the interspaces between the lamellae, and from one to three radial threads in each interspace, which with the con- centric lines produce a cancellate structure. The ribs are nearly smooth apparently because of wear, but occasionally show faint radial threads distally; submargins narrow, smooth, not depressed. Length 25 mm., height 24 mm., semi-diameter 4 mm. A single left valve was found which resembled in some respects P. cercadica Maury. Locality. — 619. Horizon. — Lares limestone. Pecten (Plagioctenium) rabelli, new species Plate XII, Fioure 6 Shell suboi-bicular, slightly oblique in adult, sub-equivalve (right valve slightly more convex); ears small, subequal; shell with thirteen to fourteen strong, smooth, rounded ribs, with slightly narrower, smooth rounded inter- spaces; right anterior ear with faint radial theads, other ears smooth; sub- margins narrow, smooth, depressed, ctenolium distinct. The surface of the valves presents no very marked feature to the unaided eye, but under a strong lens, very minute, closely, evenly spaced and remarkably regular concentric lines are visible, crossing the entire shell, including the ears. In the largest specimens a gerontic character appears in the periodic swelling and con- tracting of the ribs. These gerontic shells are more convex than the others. Length 36 mm., height 34 mm., semi-diameter 8 mm. A smaller specimen has a semi-diameter of 5 mm. Localities. — K. 29.2-29.3 Lares road, K. 25 Ponce-Adjuntas road, Juana Diaz, Yauco. Horizon. — San Sebastian shale, Juana Diaz shale. Pecten (Plag^octenium) cercadica ^laniy Plate XIII, Figures 2, 3 Pecten cercadica Maury, Bull. Amer. Paleont., V, No. 29, p. 188, PI. 34, fig. 11, 1917. Shell bilaterally symmetrical, sub-equivalve (right valve of slightly greater convexity); submargins broad, smooth except for faint growth lines; ears large, sub-equal, nearly smooth, triangular in outline except the right anterior, which is marked by deep byssal notch and byssal sulcus below, and surface of ear with five imbricated radial threads; ctenolium absent or obscure; other ears marked by very faint radial threads or striation, strongest HUBBARD, TERTIARY MOLLUSCA FROM LARES DISTRICT 95 near base of ear; surfaces of valves similar and marked by fourteen to seven- teen strong rounded ribs and slightly narrower interspaces, the whole crossed by tine close growth lines, periodically incised so as to give the effect of obscure beading of the ribs, a feature not evident on worn specimens; valves grooved within, but lirate only near ventral margins; single pair of strong cardinal crui'a. Average size; — length 21 mm., height 21 mm., semi-diameter 3.5 mm.; largest specimen; — length 2G mm., height 27 mm., semi-diameter 4.5 mm. This shell resembles P. (Plagiocienium) andersoni Arnold of the Pacific Coast Miocene, but is smaller. The latter species differs in having the left valve more convex than the right. The shell here described appears to correspond to Maury's description which was based on a left valve only. Type localities.— 619, 333, 324. Other localities.— SSI, 227 (top 30 feet). Horizon. — Lares limestone. Pecten (Plagioctenium) borinquenense, new species Plate XIV, Figures 1, 2 Shell sub-orbicular, sub-equivalve, slightly oblique, sub-marginal slopes slightly concave (left valve slightly less); sculpture of the two valves dis- similar. Right valve with sixteen to nineteen strong flat ribs of T-rail cross- section, wider than interspaces, especially in the unworn specimens where the lateral overhang obscures the borders of the interspaces; interspaces rounded and sculptured with raised concentric threads, very regular and evenly spaced (one-fifth mm. apart in ventral part of shell); tops of ribs marked by obscure median keel; concentric threads cross the ribs and are bowed up in the foi-m of lamellae on crossing the median keel. The ribs are usually worn smooth showing the lamellae on top as a series of V-shaped striations; ears sub-equal, small; hinge line less than half the length of shell, sub-margins nearly smooth; ctenolium obscure and marked by deep byssal groove bordering the line of teeth; byssal notch small; adductor scar large, central; left valve with seventeen to twenty-one strong, high, rounded ribs and sub-equal rounded interspaces; both ribs and interspaces crossed by fine raised concentric threads, somewhat irregular and more distantly spaced than in the right valve, and worn or obscure on crossing the ribs; ears sub-equal and sculptured by radial grooves and incremental lines; sub- margins as in the right valve. Interior of both valves smooth or faintly fluted, but strongly fluted and lirate around the ventral margins; greatest convexity in the umbonal region. The ribs of both valves are rounded in the nepionic stage. Adult shell measures in length 32 mm., height 31 mm., diameter of right valve 10 mm. 9G SCIENTIFIC SURVEY OF PORTO RICO This shell has ribs similar to P. diegensis Dall of the Pacitic Coast Pleistocene, but is otherwise a different shell. Maury's P. caimitica, a single left valve from the Santo Domingo Miocene is similar to the left valve of this shell. Locality. — 313. Horizon. — Upper Qiiebradillas limestone. Genus Amusium Bolten Amusium papyraceum Gabb Pleuronectia papyracea Gabb, Trans. Amer. Phil. Soc, V, p. 257, 1S73. Amusium papyraceum Dall, Trans. Wagner Inst. Sci., Ill, pt. 4, p. 718, 1898; Idem, pt. 6, p. 1586, 1903. Amusium papyraceurn Maury, Bull. Amer. Paleont., V, No. 29, p. 190, PI. 26, fig. 22, 1917. Found only in fragments, indicating a size of 35 by 36 mm. Localities. — 118, (talus), 261. Horizon. — San Sebastian shale. Amusium mauryi, new species Plate XIV, Figure 3 Shell small, thick, moderately convex, entirely smooth except for very fine growth lines; ears sub-equal, large, sub-triangular; juncture of ears with shell slightly depressed, but not otherwise marked off. The interior of the valve is inaccessible, but there seem to be no internal ribs or else very obscure ones. They do not show on the exterior, and there are no radial rays or striations of any sort. The shell is unusually solid. It is probably related to A. luna Brown and Pilsbry, and to A. mortoni Ravenel, but differs from these in its small size and relatively much larger ears. Length 19 mm., height 19.5 mm., semi-diameter 3 mm. A single left valve, which was at first included with the specimens of A. papyraceum. Dr. Maury, in looking over the specimens, pointed out the mistake. Locality. — Collazo Falls. Horizon. — San Sebastian shale. Amusium (Propeamusium) hollicki Maniy Locality. — 75. Horizon. — San Sebastian shale. HUBBARD, TERTIARY MOLLUSCA FROM LARES DISTRICT 97 Genus Spondylus Linne Spondylus bostrychites? Guppy Spondylus bifrons Sowerby, Quart. Jour. Geol. Soc, VI, p. 53, 1849; not of Goldfuss, Petref., II, p. 99, pi. 106, figs. lOa-c, 1835. Spondylus bostrychites Guppy, Proc. Sci. Assoc. Trinidad, p. 176, 1867. Spondylus bostrychites Gabb, Trans. Amer. Phil. Soc, XV, p. 257, 1873. Spondylus bostrychites Dall, Trans. Wagner Inst. Sci., Ill, pt. 4, p. 758, 1898; pt. 6, p. 1586, 1903. Spondylus bostrychites Maury, Bull. Amer. Paleont., V, No. 29, p. 190, pi. 32, fig. 4, 1917. A small internal mold of a Spondylus, which, because of its small hinge area and sub-equal valves, is identified doubtfully with the above species. Length 22 mm., height 26 mm., diameter 15 mm. Localities.— 1Q2, 23?, 126. Range. — San Sebastian shale to Cibao limestone. Spondylus gumanomocon Brown and Pilsbry Spondylus americanus Gabb, Trans. Amer. Phil. Soc, V, p. 257, 1873; not of Lamarck. Spondylus gumanomocon Brown & Pilsbry, Proc. Acad. Nat. Sci. Phila., p. 514, 1912 (footnote). Spondylus gumanomocon Maury, Bull. Amer. Paleont., V, No. 29, p. 191, 1917. An internal mold and a fragment of the external mold were found of a shell which is like 8. gumanomocon but somewhat smaller. It may be a small variety or the young shell. Lower valve strongly convex, beak long, erect; surface with strong ribs 5 mm. apart, ap- parently spinose, and alternating with three or four small obscure ribs. The ribs are slightly Avavy; upper valve orbicular, slightly convex ; ribs known only from the internal mold but seem to be like those of the upper valve except that they are finer ; hinge line about 24 mm. in length. The internal mold shows the beak of the lower valve to be considerably excavated. Length 55 mm., height of lower valve 66 mm., of upper valve 54 nnu., diameter 29 mm. From the proportions of the internal mold it will be seen that this shell is less elongate than S. gumanomocon as described from Santo Domingo. The upper valve is much flatter than the Gatun species, S. scotti Brown & Pilsbry, but the shell is about the same size as the latter and is similarly sculptured. Localities. — 140, 162. Range. — San Sebastian shale to Cibao limestone. 98 SCIENTIFIC SURVEY OF PORTO RICO Genus Ostrea (L) Lamarck Ostrea antiguensis Brown Plate XV, Figure 1 Ostrea antiguensis Brown, Proc. Acad. Nat. Sci. Phil., p. C14, PI. 19, fig. 7; PI. 20, figs. 1, 5, 6, 1913. These shells were at first regarded as a variety of 0. hmtensis, but on comparison, were found to be the same as Dr. Maury's Porto Rican specimens, identified bv Dr. Pilsbry as 0. antiguensis, ac- cordingly that determination is accepted, although it is diihcult to believe that the reflexed character of the upper valve is a specific character. All of the specimens here described -were taken from a zone of yellow chalk in the upper Quebradillas exposed in sea cliffs near Pt. Borinquen. The abundance of these large oysters is remarkable. They reach an enormous size and thickness, but the largest specimen collected measures in leugth 140 mm., height loO mm., diameter (the two valves together) 110 mm. These forms are obviously geron- itc individuals. The lower valves are strongly curved upward ventrally toward the center, but are reflexed (or curved down) on the lateral margins. Likewise, the upper valves are curved down on the lateral margins, and strongly reflexed at the ventral margin, which results in a strong interlockiug of the two valves. The u]->per valves are almost as thick as the lower ones. The upper valves reach their maximum thickness between the center and the ventral margin, and in both valves the animal has excavated its body cavity directly below the beaks by resorption of the prismatic layers at this point with increased deposition farther on toward the ventral mar- gin. Thus as seen in a longitudinal section of the shell these layers resemble the stratification in a migrating sand dune. The excavated body cavity occupies but a small portion of the interior of the shell. Other specimens were found which, although almost as large as those above described, are apparently not gerontic individuals. The reflexed character of the upper valve is slightly or altogether absent, and the shell does not attain a great thickness. The spinose ribs (about 9 in number) and rounded furrows are well shoMTi on these specimens, whereas they are obsolete or extremely irregular on the gerontic forms. The adductor scar is high and nearly central in the HUBBARD, TERTIARY MOLLUSCA FROM LARES DISTRICT 99 thin shells, but well toward the anterior margin in the gcrontic shells. It is deeply impressed in the latter. The resilifer is re- markably small for shells of this size. Localities. — 314; San Jnan District (Semmes). Horizon. — Upper Quebradillas limestone. Also reported from the "Guanica shaly limestone" by Dr. Manry, Ostrea haitensis So\verby 0. haitensis Sowerby, Quart. Jour. Geol. Soc. London, VI, p. 53, 1850. 0. haytensis Gabb, Trans. Amer. Phil. Soc, XV, p. 257, 1873. O. veatchii Gabb, Pal. Cal., II, p. 34, PI. 11, fig. 59; PI. 17, fig. 21, 1869. 0. hermanni Conrad, Proc. Acad. Nat. Scl. Phila., V, p. 267, 1853. 0. vespertina Conrad, Pac. R. R. Rept., V, p. 325, PI. 5, figs. 36-38, 1855; Gabb, Pal. Cal., II, p. 107, 1869. 0. virginica Guppy, Quart. Jour. Geol. Soc, XXII, p. 577, 1866. Not of Gmelin. 0. virginica var. californica Marcou, Geol. No. Am., 1858. 0. haitensis Guppy, Quart. Jour. Geol. Soc. London, XXXII, p. 532, 1876. 0. haitensis Dall, Trans. Wagner Inst. Sci., Ill, pt. 4, p. 685, 1898. Idem, pt. 6, p. 1586, 1903. 0. haitensis Maury, Bull. Amer. Paleont., V, No. 29, p. 182, PI. 31, figs. 1, 2, 1917. The shells are rather thin, orbicular in outline, and strongly pli- cate with seven or eight sharply angular ribs, very high centrally at the ventral margin, and sharply angiilar furrows between. The ribs are not noticeably spinose ; resilifer shelf -like, due to excavated beak; adductor situated in central-posterior portion of vale. The shells do not attain a large size, the largest specimen measuring in length 72 mm., height 74 mm., diameter of lower valve about 30 mm. Localities.— 225, 800. Range. — Lares limestone, Cibao limestone. Ostrea haitensis Sowerby, variety? Plate XV, Figures 2, 3 Some small shells were found in which the lower valves are sculptured like 0. haitensis, with spinose ribs, frequently angular, while the upper valves are nearly smooth. The lateral inner mar- 100 SCIEXTIFIC SURVEY OF PORTO RICO gins adjacent to the beaks are finely creniilate, a feature not slio\\ii in the specimens of 0. Jiaitensis. Largest specimen measures in length oQ mm., height 43 mm., diameter lower valve 7 mm. A smaller more orbicular lower valve measures in length 22 mm., height 25 mm., diameter 6.5 mm. These little shells may be the young of 0. haitensis, or a variety. Locality. — 62. Horizon. — Cibao limestone. Ostrea virginica Gmelin 0. virgiana of Lister and others. O.virginica Gmelin, Syst. Nat., p. 3336, 1792; Dilwyn, Descr. Cat., 1, p. 277, 1817; Lam., Anim. s. Vert., VI, p. 207, 1819. O.edulis Akerly, Am. Monthly Mag., II, p. 296, 1818 (not Linn6). 0. virginiana Sowerby, Genera, Ostrea, f. 2, 1822. 0. borealis Lamarck, Anim. s. Vert., VI, p. 204, 1819. 0. canandenis Lamarck, op. cit., p. 207, 1819. 0. triangularis Holmes, Proc. Elliot Soc, I, p. 29, 1856. 0. fundata Holmes, Post-Pl. Foss. S. Car., p. 11, PI. 2, fig. 10, 1858. 0. semicylindrica Say, Jour. Acad. Nat. Sci. Phila. (1), II, p. 258, 1822. 0. virginica Gabb, Trans. Amer. Phil. Soc, XV, p. 257, 1873. 0. virginica Guppy, Quart. Jour. Geol. Soc. London, XXXII, p. 532, 1876. 0. virginica Dall, Trans. Wagner Inst. Sci., Ill, pt. 4, p. 687, 1898. O. virginica Maury, Bull. Amer. Paleont., V, No. 29, p. 184, 1917. The specimens show the usual great variation, but comparison with recent specimens of 0. virginica shows no constant differences. The Porto Rican shells vary from elongate, subtriangular to ovate: beak of lower valve usually pointed, straight or slightly curved laterally; resilifer elongate in the large shells; lower valves are frequently of extreme convexity, while the upper valves are gently convex or flat, but very irregular; beak of lower valve excavated. An average seized lower valve measures in length 50 mm., height 90 mm., diameter 33 mm. Some are much broader, and a few- specimens reach a much greater size and greater thickness. Color usually bluish, mottled with white. Type locality.— ^2.%% Other localities.— 2S1, 65, 227 (top 30 ft), 7 to 8 (on road-flat). Horizon. — Upper lares limestone (very characteristic). HUBBARD, TERTIARY MOLLUSCA FROM LARES DISTRICT 101 Ostrea sellseformis Conrad variety portoricoensis, new variety Plate XIII, Figures 4, 5, 6 Shell of moderate size, pear-shaped to elongate oval, never attaining very great thickness; valves dissimilar; lower valve with numerous radial ribs imbricated by raised laminae; resilifer deep; margins of ligament area marked by narrow groove which continues for some distance along the inner margins adjacent to the beak, and is denticulate (or marked by row of small pits) throughout; remainder of inner margin fluted by the ribs, or, in some specimens, smooth; adductor scar large, oval, nearly central; beak more or less curved laterally; upper valve smaller, convex to nearly flat; surface nearly smooth or marked usually by even, widely spaced, slightly raised concentric lamellae; beak curved laterally but less pointed than in lower valve, and with shorter ligament area; resilifer moderately impressed or even raised and convex in some individuals; lateral grooves indistinct or absent; inner margin crenulate adajacent to beaks, but smooth ventrally; adductor scar smaller than in lower valve and uniformly pear-shaped in all specimens. An average sized shell measures, lower valve, in length 62 mm., height 70 mm., upper valve, in length 42 mm., height 60 mm. Diameter (both valves) 19 mm. The upper valves of the young shells are more C(mvex. A typical young upper valve measures in length 23 mm., height 33 mm., diameter 7 mm. Type locaIiUe&.—S4:S, 333, 176, 59, 800. Other localities.— 619, 332, 64. Eange. — Lares limestone, Cibao limestone, Los Puertos limestone. Ostrea collazica Maury Shell sub-elongate to sub-orbicular, heavy, attaining great thickness in large individuals; valves dissimilar; lower valve convex, with numerous small irregular radial plications which divaricate from a median or slightly posterior zone; plications finely imbricated by thin, slightly raised laminae; beak strongly recurved laterally; ligamental area extremely long in large specimens; resilium deep but becoming shallow in gerontic individuals; margins of ligamental area marked off by narrow crenulated groove; inner margins near the beak finely crenulated; upper valve flat, with beak similarly recurved, usually more closely gyrate; surface smooth except for the thin laminae; interior similar to lower valve except for an amorphous, tooth-like process, located on the inner margin just anterior to the ligamental area, which is transversely denticulate or marked by cross grooves. In some speci- mens the margins of both valves are similarly marked near the beaks; adductor scars of both valves large and situated slightly posterior. Average size adult;— length 75 mm., height 85 mm. 102 SCIENTIFIC SURVEY OF PORTO RICO Fragments indicate that the shell reaches a considerably larger !size, in which the lower valve is gi-eatly thickened. Two of these large lower valves were fonnd in wliich the tendency of the beak to curve laterally has apparently been almost lost, the beak being long and pointed. An unusual feature of this shell is the small amount of prismatic structure. Some of the specimens are made up almost entirely of nacreous laminae t)f micaceous thinness. This shell is of the 0. sellceformis divaricata type, and the young individuals might readily be taken for a variety of 0. sellceformis. The full grown flat upper valves with their closely gyrate beak bear a strong resemblance to 0. vaughni Dall of the Tampa Silex beds. Tyfye localities. — 7 to 8- (float), Lares road just east of Collazo bridge, K. 29.2 to 29.3 Lares road. Other localities.— 100, 220, 231, 233, 267, 89, 232(b), 236. Range. — San Sebastian shale to Lares limestone. Characteristic of the San Sebastian shale, and occuring in the lowest fossilifcrous zone of the latter. Ostrea cahobasensis Pilsbry and Brown variety portoricana, new variety Plate XVI, Figure 1 ; Plate XVII, Figure 1 Shells reach a vei-y large size, and are characterized by their elongate form and long, pointed, and usually straight beak. The young shells are much less elongate in form. The body cavity tends to remain the same size during growth, most of the addition in height being in the beaks. The resilifer is shallow in the lower valve, and convex in the upper valve, and in both valves it is bordered by a broad, prominent ridge on each side. The latter are marked off from the lamellae of the lateral slopes by a deep impressed zone, in which there is no trace of dentition or crenulation. The beaks may be twisted or straight, and there is considerable variation in the outline of the shells due to the accidents of growth. The shells apparently grew in crowded colonies. The lower valve is convex, the upper slightly convex, or nearly flat in the younger shells. The beak of the lower valve always pro- jects far beyond the beak of the upper valve, and in the young, is deeply excavated within, below the resilifer. The position of the adductor scar is variable, being rather high in the young, but travels ventrally with the growth of the shell, until, in the largest individuals, it is located at the ventral one-third or one-fourth of the shell. The large shells have an hour glass-shaped body cavity, consisting of a deep round cavity immediately below the resilifer, a contraction centrally, and a large crescent-shaped ventral cavity which begins opposite the adductor scar, and swings around below the HUBBARD, TERTIARY MOLLUSCA FROM LARES DISTRICT 103 scar, and close to the ventral margin. This is a fairly constant fea- ture of the larger shells, knd is strongly developed in some of the young or half grown shells. The surface of both valves is marked only by concentric lamellae, and shows no signs of radial fluting of ribs. The lamellae are, however, occasionally quite irregular. This shell, while apparently answering the description of the Haitian species, shows differences of varietal rank. The largest specimen measures in length 100 mm., height 270 mm., diameter (both valves together) 95 mm. Localities. — West of Guaiiica Mill in basal limestone a few feet above tlie contact with the Older Series (Lobeck) Fragments of a similar shell Avere found in the Lares limestone, bnt their identity with the Gnanica shell is by no means certain. Horizons. — Ponce limestone, Lares limestone (donbtful). Genus Lithophaga Bolteii Lithophaga nigra d'Orbigiiy Lithodomus niger d'Orbigny, De la Sagra, Hist. Polit. y Nat. Isla de Cuba, pt. 2, V, p. 351, 1845; Atlas, VIII, PL 28, figs. 10, 11, 1855 (Spanish Ed.). Modiola caribwa Philippi, Abbild. u. Beschr., Ill, p. 20, PI. 2, fig. 5, 1847. Modiola antillarum Philippi, op. cit. p. 20, PI. 2, fig. 4, 1847; Zeitscher., p. 116 (not of d'Orbigny); young shell. Mytilus lithophagus Gibbes, S. Car. Cat., p. 22, 1848; not of Linne. Lithophagiis nigra Morch, Cat. Yoldi, II, p. 56, 1853. LitJiophagus antilla7~um Reeve, Conch. Icon., X, PI. 2, fig. 7, 1857. Lithophaga nigra Dall, Trans. Wagner Inst. Sci., Ill, pt. 4, p. 799, 1898; Bull. 90, U. S. Nat. Mus., p. 129, 1915. Lithophaga nigra Maury, Bull. Amer. Paleont, V., No. 29, p. 194, 1917. This shell is known from the Tampa Silex beds, and continues to the recent fauna of the Antilles. The recent shell of Cuba as described by d'Orbigny, has a length of 50 mm. The Porto Rican fossils differ only in size, the largest measuring in length 30 mm., height 9 mm., diameter 8 mm. The occurrence is chiefly in the form of internal molds, but fragTuents of the shell show the vertical striae which end abruptly at a transverse oblique line extending from the beaks to the posterior ventral margin. Locality. — 225. Horizon. — Upper Lares limestone. 104 SCIENTIFIC SURVEY OF PORTO RICO Genus Leda Schumacher Leda peltella Dall Plate XVII, Figure fi Leda acuta Gabb, Trans. Amer. Phil. Soc, XV, p. 255, 1873. Not L. acuta Conrad, 1832, nor Sowerby, 1837. Leda peltella Dall, Trans. Wagner Inst. Sci., Ill, pt. 4, p. 579, PI. 32, fig. 5; Idem, pt. 6, p. 158G, 1903. Leda peltella Maury, Bull. Amer. Paleont., V, No. 29, p. 161, PI. 2G, fig. 9, 1917. One of the most abundant fossils in the Quebradillas limestone. The posterior sulcation is always broad and distinct ; anterior sulcation narrow and often indistinct. Length 11 mm., heigh 6.5 mm., semi- diameter 3 mm. No specimens were found less than 9 mm. in length. Type locality. — 204. Other locality. — 660. Horizon. — Quebradillas limestone. Genus Area Linne Area yaquensis Maury Area yaquensis Maury, Bull. Amer. Paleont., V, No. 29, p. 164, PI. 30, fig. 14, 1917. Several specimens M-ere found of a very small Area which closely resembles Maury's A. vaquensis. a somewhat doubtful species from the Santo Domingan Oligocene. The Porto Rican shells show no distinctive characters, and are placed with A. yaquensis because they correspond more closely to this than to any other species. The specimens are very numerous and all about the same size, hence it is unlikely that they are yoimg individuals. Length 8 mm., height 4.5 mm. Type locality. — 204. Other locality. — 225. Range. — Lares limestone to Quebradillas limestone. HUBBARD, TERTIARY MOLLUSC A FROM LARES DISTRICT lO.") Area dariensis Brown and Pilsbiy Plate XV, Figure 5 Area dariensis Brown and Pilsbry, Proc. Acad. Nat. Sci. Phila., p. 362, PI. 22, fig. 10, 1911. Umbo prominent, at anterior third of shell; surface with thirty-three flat ribs and sub-equal flat interspaces, the whole crossed by fine raised concentric lines, strongest on ribs and giving to the latter a nodular appearance; ribs broaden anteriorly and posteriorly, the ten posterior and the five or six anterior ribs medially grooved. Length 26 mm., height 16.5 mm., semi- diameter 5.5 mm. This Area is smaller than the Gatun species, and may be a variety of it. It also resembles the shell described by Tonla as Area cf. consohrina Moore. Localities.— CoWsizo Falls, 261?, 232(b). Horizon. — San Sebastian shale. Area (Scapharea) cf. donacia Dall Plate XV, Figures 6, 7 Area (Scapharea) donacia Dall, Trans. Wagner Inst. Sci., Ill, pt. 4, p. 649, PI. 33, fig. 13, 1898. The size and sculpture answers DalFs description of this species, especially the raised concentric threads in the interspaces of the left valve, a feature lacking on the right valve. It differs from the Bowden form in the number of ribs (27), and in having a shallow median depression, strongest in the left valve. Length 8.5 mm'., height 6.5 mm. A smaller shell measures in length 6.75 mm., height 4.5 mm. Locality. — 204. Horizon. — Quebradillas limestone. Area, species indet. Plate XVII, Figures 2, 3 Shell small, very convex; beaks at anterior third; anterior and posterior ends equally rounded; hinge area broad, especially anteriorly, with beaks widely separate; surface with twenty-five broad, quadrate, rather low ribs and equally wide, shallow rounded interspaces; central ribs medially grooved distally; hinge margin three-fourths the length of shell and slightly raised 106 SCIENTIFIC SURVEY OF PORTO RICO above hinge area; hinge area transversely grooved or striate posteriorly; teeth fine, numerous; interior of valves unknown. Average size measures in length 19 mm., height 10.5 mm., diameter 12 mm. Largest specimen meas- ures in length 25 mm., height 13.6 mm., diameter 16 mm. Tyije locality. — 230. Other- localities.~rdZ, 232(b). Horizon. — Upper San Sebastian shale. Area (Scapharca) collazica Maury The material collected by the writer consists only of internal molds. An average sized specimen measures in length 16.5 mm., height 9.5 mm., diameter Y.5 mm. Localities.— ZZ^, G19, 64, 232(b). Range. — Upper San Sebastian shale to Cibao limestone. Very abundant. Subgenus Barbatia Gray Barbatia (Acar) reticulata Gmelin Area reticulata Gmelin, Syst. Nat., VI, p. 3311, 1792. Area reticulata Sheldon, Palaeontographica Americana, I. p. 20, PI. 4, figs. 8-12, 1916. Barbatia (Acar) reticulata Maury, Bull. Amer. Paleont., V, No. 29, p. 166, PI. 30, fig. 16, 1917. A single fragment was found, sufficiently complete to identify. Length indeterminate, height 6 mm. Locality. — 204. Horizon. — Quebradillas limestone. Barbatia cf. bonaczyi Gabb Plate XIV, Figure 4 Barbatia bonaczyi Gabb, Trans. Amer. Phil. Soc, XV, p. 254, 1873. Area umhonata Dall, Trans. Wagner Inst. Sci., Ill, pt. 4, p. 620, PI. 38, figs. 4, 4a, 1898. Not of Lamarck. Barbatia cf. bonaczyi Maury, Bull. Amer. Paleont., V, No. 29, p. 165, PI. 30, fiK. 15. 1917. A single external mold of a right valve resembling B. bonaczyi Gabb. but smaller. It is nearer the size of B. cf. bonaczyi described HUBBARD, TERTIARY MOLLUSCA FROM LARES DISTRICT 107 by Maury, but the seul])ture is coarser. It is very ]ir()l)ably a variety, but the speciiuen is too poor for complete description. The ribs are crowded centrally and sparser at the ends of the valve. Those on the posterior slope lose their bearded character and become scaly. Valve with shallow median sulcation. Length about 12 mm., height 6 mm. Locality. — 204. Horizon. — Quebradillas limestone. Genus Glycimeris Da Costa Glycimeris portoricoensis, new species Plate XV, Figure B SheU sub-orbicular; beaks not prominent; surface with about thirty strong sub-angular to rounded ribs and sub-angular interspaces, the whole crossed by fine raised concentric threads, regular and evenly spaced. The ribs near the ventral margin broaden rapidly at the expense of the interspaces, and become round in cross-section. Interior of shell unknown. Largest specimen measures in length 19 mm., height 17 mm., semi-diameter 4 mm. This shell resembles G. acuticostata Sowerby, but has broader, more rounded ribs. Localities.— ^0^, 182, 318. Eange. — Upper Los Puertos limestone to Quebradillas limestone (typical of the latter). Glycimeris coUazoensis, new species Plate XV, Figures 4, 5 Shell sub-orbicular, beaks not prominent; hinge area short; teeth numer- ous, in uninterrupted series; surface with thirty-four low angular ribs and angular interspaces; each rib with fine keel or thread on the apex, and with the appearance of a groove on either side of this median thread; inner margin scalloped ventrally; length 15 mm., height 16 mm., semi-diameter, 4 mm. This shell resembles G. gatunensis Toula, but the latter is smaller and more transversely elongate. It also resembles G. portoricoensis of the Quebradillas limestone, of which it may posssibly be the pr(^ genitor. G. portoricoensis differs chiefly in its larger size, more rounded ribs, and strong concentric growth lines. Locality. — 261. Horizon. — Lower San Sebastian shale. 108 SCIENTIFIC SURVEY OF PORTO RICO Genus Venericardia Lamarck Venericardia scabricostata? Guppy Cardita scabricostata Guppj% Quart. Jour. Geol. Soc. London, XXII, p. 293, PI. 18, fig. 10, 1866. Cardita scabricostata Gabb, Trans. Amer. Phil. Soc, XV, p. 252, 1873. Cardita scabricostata Guppy, Quart. Jour. Geol. Soc. London, XXXII, p. 531, 1876. Venericardia scabricostata Dall„ Trans. Wagner, Inst. Sci., Ill, pt. G, pp. 1428, 1586, 1903. Venericardia scabricostata Maury, Bull. Amer. Paleont., V., No. 29, p. 198, PI. 33, fig. 1, 1917. Shell with prominent beaks; 18 to 20 high nodose ribs and nar- rower, V-shaped interspaces which are crossed by strongly marked growth lines; lunule broad, short, and distinct. Length about 4.5 mm., height 4 mm., semi-diameter 1.5 mm. The specimens foimd are much smaller than V. scabricostata, and yet seem to be mature shells; they may be a variety. Locality. — 225. Horizon. — Lares limestone. Venericardia cerrogordensis Maury, new variety Venericardia cerrogordensis Maury, Bull. Amer. Paleont., V, No. 29, p. 199, PI. 33, fig. 3, 1917. A single fragment of anterior half of a left valve, which seems to be a variety of this species, somewhat larger than the Santo Domingo shell. About fourteen strong ribs with strongly beaded keel; inter- spaces wider than ribs, shallow, with wavy growth lines and showing an obscure crenulated radial thread on either side at base of rib. Length about 27 mm., height about 23 mm., semi-diameter, etc., 5.5 mm. This may be a distinct species but the specimen is so incomplete that it has been identified with the nearest form.. The specimen was found in a piece of road metal at K. 26.5 Lares road and its horizon is unknown. The rock is a white limestone like the Quobradillas and contains a mold of Chione cf. woodivardi Guppy. HUBBARD, TERTIARY M0LLU8GA FROM LARES DISTRICT 109 Genus Chama (Linne) Bruguiere Chama involuta Guppy Chama involuta Guppy, Geol. Mag. London, decade 2, I, p. 436, PI. 17, figs. 5a-c, 1874; Quart. Jour. Geol. Soc. London, XXXH, p. 531, 1876. Chama involuta Dall, Trans. Wagner Inst. Sci., Ill, pt. 6, pp. 1398, 1586, 1903. Chama involuta Maury, Bull. Amer. Paleont., V, No. 29, p. 199, PI. 33, figs. 4, 5, 6, 1917. The Porto Rican specimens are unusually large, but apparently of this species. The largest specimen is fragmental, but shows a height of 52 mm. Judging from the specimens, it attaches by either valve. The beak is strongly prosogyrate, making over one complete turn. The lamellae are strongly fluted, almost spinose on one specimen. Localities. — 204, 98(Berkey). Horizon. — Quebradillas limestone. Chama portoricana, new species Plate XVII, Figure 5 Beak closely prosogyrate; surface with extremely crowded, thin, high lamellae, which on the largest specimen rise 8 mm. above the shell surface. These lamellae are quite regular, being gently fluted over the body of the shell, but spinose at the margin in the adult; shell marked by rounded um- bonal ridge; inner margin finely crenulate; cardinal area pustulose; strong bifid and crenulate cardinal tooth in the right valve, with crenulate lateral; shell attaches by either valve, but the cardinal tooth remains in the right valve in either case; adductor scars large; pallial line distinctly impressed. Length 48 mm., height 35 mm., diameter of convex valve about 12 mm. A smaller attached valve measures length 22 mm., height 18 mm., diameter 9 mm. This shell differs from C. congregaioides Maury in its more regular and higher lamellae, and in its more elongate form. Locality. — 204. Horizon. — Quebradillas limestone. Genus Lucina (Bruguiere) Lamarck Lucina coUazoensis, new species Plate XVIII, Figures 1, 2 Shell transversely ovate, convex, produced posteriorly, beaks being slightly forward of center and frequently so close together that they practically touch; 110 SCIENTIFIC SURVEY OF PORTO RICO hinge line long and straight, nearly nine-tenths the length of the shell. The largest specimen, somewhat distorted so as to increase the convexity, meas- ures in length 65 mm., height 40 mm., diameter 45 mm. A smaller shell, not distorted, measures in length 39 mm., height 31 mm., diameter 25 mm. This is a ponderous shell of the type of L. corpulenta Dall, but unusual for its elongate form and long straight hinge line. It oc- curs chiefly as internal molds of both valves intact, but the form is often distorted by pressure. Fraginents of external mold show the surface to be nearly smooth, marked only by fine growth lines, peri- odically incised at irregular intervals. There is an undescribed shell in the Kemp collection from the lower Gatun beds which is apparently this species, or very close to it. Localities. — 74, 92, 86, Collazo (float). Horizon. — San Sebastian shale (abundant at Collazo Falls). Lucina cf. chrysostoma (MeiTSchen) Philippi Plate XVIII, Figure 3 Tellina crysostoma Meuschen, Mus. Gevers., p. 482, 17S7 (typographical error). Lucina chrysostoma Philippi, Abb. und Beschr. neue Conchyl., II, p. 206, PI. 1, fig. 3, 1847. Venus edentiaa Chemnitz, Conch. Cab., VII, PI. xl, figs. 427-9, 1784; Gmelin, Syst. Nat., VI, p. 3286, 1792; not of Linne, Syst. Nat., ed. 10, p. 689, 1758. Anodontia alia Link, Beschr. Rostock Samml., p. 56, 1807. Lucina edentula Reeve, Conch. Icon., Lucina, PI. ii, fig. 9, 1850; Heilprin, Trans. Wagner Inst., I, pp. 102-3, 1886. Loripes chrysostoma Arango, Fauna Mal. Cubana, p. 257, 1878. Lucina chrysostoma Dall, Trans. Wagner Inst., Ill, pt. 6, p. 1354, 1903. Loripes edentula Gabb, Trans. Amer. Phil. Soc, XV, p. 251, 1873. Lucina chrysostoma Maury, Bull. Amer. Paleont., V, No. 29, p. 202, PI. 35, fig. 2, 1917. Resembles L. clirysostoma of the Caloosahatchie Pliocene, but the largest and most convex specimens resemble the recent L. globosa Forskol. Only internal molds have been found, which show lunule impression, and posterior sulcation close to dorsal margin. The specimens range in length 39 mm., height 36 mm., diameter 22 mm. to length 135 mm., height 115 mm., diameter 95 mm. Localities.— 205, 174. 10, 78, 98, 176? (Berkcy). Mona Island, south side of Vieques Island, Guanica, and Ponce (Lobeek). HUBBARD, TERTIARY M0LLU8CA FROM LARES DISTRICT 111 Horizons. — Los Puertos, Quebradillas limestones, and Ponce for- mation. These large shells are very abundant as internal molds in the Tertiary formations of the north and south coasts. Genus Phacoides Blainville Phacoides (Miltha) riocanensis Maury Phacoides (Miltha) riocanensis Maury, Bull. Amer. Paleont., V, No. 29, p. 205, PI. 35, fig. 7, 1917. The Porto Rican shells are the same size as those of the Santo Domingan Oligocene. There seem to be no important differences. The former shells have a slight differentiation in the concentric lamellae, two lamellae of secondary strength alternating with each primary one. A shallow median sulcus is a noticeable feature near the ventral margin, but does not extend to the beak. Length about 4:7 mm., height about 45 mm., semi-diameter 7.5 mm. Localities.— 335, 204, 220, 225, 318, 31Y. Bange. — Lares limestone, Cibao limestone, Los Puertos limestone, Quebradillas limestone. This species is widespread but not abundant. Phacoides (Miltha), species indet. Plate XVIII, Figure 4 Shell of moderate size, convex, slightly inequalateral; hinge area narrow, short, beaks incurved and nearly in contact; anterior margin broadly rounded, posterior margin roundly truncate, basal margin nearly straight; posterior sulcus close to margin; concentric sculpture of fine, even, raised lamellae, about 1/3 mm. apart at ventral margin, and with fine growth lines in the spaces between. Height of shell about 24 mm., length about 34 mm., semi- diameter 7 mm. Only a single fragment was found which was too incomplete for an accurate description, but differed from any other species. This shell resembles P. (Miltha) cf. smWnvoodwardi Maury, a species from the same horizon, but is more elongate than the latter, and has the posterior sulcus much closer to the margin. Locality. — 204. Horizon. — Quebradillas limestone. 112 SCIENTIFIC SURVEY OF PORTO RICO Phacoides (Miltha), species indet. Plate XVIIT, Figure 5 Shell small, orbicular, moderately convex. Sculpture of fine concentric lamellae like P. Jiillsboroense; deep radial sulcus crossing the posterior one-quarter of shell and extending to ventral margin. Fine concentric striae between the lamellae are visible under a lens. Length 12 mm., height 12.5 mm., semi-diameter 3.5 mm. Only one specimen was found, the external mold of a left valve. It may possibly be tbe young of a large species like P. riocanensis. Locality. — 52(a). Horizon. — Quebradillas limestone. Phacoides (Pseudomiltha) laresensis, new species Plate XVIII, Figure 6 Shell sub-orbicular, nearly equilateral, moderately convex; surface with concentric lamellae of the type of P. hilWboroense, but very regular instead of wavy as in the latter; concentric striae between the lamellae; lunule narrow, deeply impressed; escutcheon large, marked off by a narrow sulcus which broadens at the posterior margin; concentric sculpture flexed sharply upward over the escutcheon, where it becomes coarser; surface of shell marked by faint radial striae; cardinal area edentulate; interior of shell unknown. The shell is thin, and in most of the specimens i^ worn partially or wholly away, leaving only the internal mold. Length 36 ram., height 34 mm., diameter 18 mm. Localities.— 2Q2, Collazo Falls, 232(b) ? Horizon. — San Sebastian shale. Phacoides (Lucinisca) calhounensis Dall Plate X, Figure 6 Phacoides {Lucinisca) calhounensis Dall, Trans. Wagner Inst. Sci., Ill, pt. 6, p. 1371, PI. 52, fig. 16, 1903. Phacoides calhounsis Maury, Bull. Amer. Paleont., V, No. 29, p. 204, 1917. (Typographical error.) These shells are variable but have the same general characters as the Florida form. The majority of the specimens are the same size as the latter, but occasionally attain a considerably greater size. The greater prominence and sparcity of the concentric ridges in the um- HUBBARD, TERTIARY M0LLU8GA FROM LARES DISTRICT 113 bonal region is a noticeable feature. The radial ribs are finer and somewhat crowded on the posterior slope and to a lesser extent on the anterior slope. Dall does not mention this character but it may be observed in his illustration. The intersections of the radials wdth the concentric ribs have a tendency to form spinose nodes, especially in the adult stage near the lateral margins. This gives the shell a superficial resemblance to P. hispaniolana Maury, but the sculpture of the latter is much finer. The Porto Rican shell has a large and distinct lunule. An average size specimen measures in length 10 mm., height 9.5 mm., semi-diameter 2.5 mm. The largest specimen measures in length 13 mm., height 12 mm., semi-diameter 4 mm. Type locality. — 204. Other localities. — 52(a), 77(Berkey). Horizon. — Quebradillas limestone (abundant) ; Ponce limestone. Genus Divaricella Von Martens Divaricella prevaricata Guppy Divaricella prevaricata Gxx^^j, Proc. U. S. Nat. Mus., XIX, No. 1110, p. 327, PI. 30, fig. 4, 1896. Divaricella prevaricata Dall, Trans. Wagner Inst. Sci., Ill, pt. 6. pp. 1389, 1587, 1903. Divaricella prevaricata Maury, Bull. Amer. Paleont., V., No. 29, p. 207, PI. 35, fig. 10, 1917. The divaricate lines are finer than those of the Santo Dominiro shell. The single specimen found measures in length 7 mm., height 5.75 mm. Locality. — 204. Horizon. — Quebradillas limestone. Genus Codakia Scopoli Codakia (Jagonia) magnoliana Dall variety borinquenense, new variety Plate XVII, Figure 4 A single right valve "was collected. It is slightly smaller than the shell described from the Korth Carolina Miocene, and the concentric sculpture is more pronounced. The radials are slightly waw as in 114 SCIENTIFIC SURVEY OF PORTO RICO Codakia orbicularis, and increase by intercalation. They vary in strength, parts of the shell being marked by ribs finer and more crowded than in adjacent portions of the surface. The shell is noticeably inequilateral. The hmule is long and narrow. Length 10 mm., height 8 mm., semi-diameter 4 mm. Localities. — 204, 671. Horizon. — Qnebradillas limestone. Genus Cardium (Liniie) Lamarck Cardium (Lsevicardium) ef. serratum Tinne Cardium serratum Linne, Syst. Nat., ed. 19, p. 680, 1758. Cardiuyn lavigatum Lamarck, Auim. s. Vert., VI, pt. 1, p. 11, 1819 (not of Born, Mus. Vind. Test., p. 47, 1780; nor of Linne, Syst. Nat., X, p. 680, 1758). Cardium citrinum Wood, Gen. Conch., p. 223, PI. 54, fig. 3, 1815. Liocardium pictumRa.\ene\, Proc. Acad. Nat. Sci. Pliila., for 1861, p. 44, 1862. Cardium hiatus "Meuschen" fide Krebs, W. I. Cat. Stiells, p. 115, 1864. Cardium lineatum Krebs, op. cit., not of Gmelin, 1792. Cardium pristis Valenciennes, fide Krebs, op. cit. Cardium ovipi(tame7i Reeve, Conch. Icon., Cardium, PI. 7, fig. 36, 1844. Cardium venustum Gabb, Trans. Amer. Phil. Soc, XV, p. 251, 1873. Cardium serratum Ball, Proc. U. S. Nat. Mus., XIX, No. 1110, p. 327, 1896; not serratum of Pennant, 1778). Cardium serratum Bb-U, (in part), Trans. Wagner Inst. Sci., Ill, pt. 5, p. 1110, 1900. Cardium serratum Brown and Pilsbry, Proc. Acad. Nat. Sci. Phila., p. 367, 1911. Cardium serratum Maury, Bull. Amer. Paleont., V, No. 29, p. 212, PI. 36, fig. 8, 1917. Numerous internal molds apparently of this species, and of about the same size as those of the Santo Domingo Oligocene were found. They average about 20 mm. in height. Frngments of external mold show numerous fine thread-like radial ribs. Some specimens show a slight radial sulcation anteriorly. Localities.— 204:, 225, 333?, 64. i?an^e.— Lares limestone to Qucbradillus limestone. Cardium (Trachy cardium) muricoides, new species Plate XIX, Figure 1 Shell sub-ovate, convex; surface with about 36 spinose ribs and narrower, rounded, transversely striate interspaces; Interior of valves grooved and scalloped at ventral margins. All ribs except the three anterior ones have a HUBBARD, TERTIARY MOLLUSGA FROM LARES DISTRICT 115 lateral keel on each edge at the top. The three anterior ribs have only one such keel, placed on the anterior side. The first ten ribs (including the three single keeled ones) are ornamnted with longitudinally ovate, long, erect, or slightly hooked spines along the anterior keel, but spreading out over the center of the rib. All the remaining ribs have the spines on the posterior side. From the eleventh rib to the eighteenth inclusive, the spines are like those of the ten anterior ribs, but commencing with the nineteenth rib the spines posteriorly become narrower, blade-like, set longitudinally on the rib, and curved posteriorly over the adjacent interspace. All the ribs are fiat topped, but the posterior ribs have the tops sloping posteriorly. Length 30 mm., height 32.5 mm., semi-diameter 11 mm. This shell is twice as large as C. howdenense Dall. The orna- mentation especially of the rib is nearer to that of C. muricatum than to that of any other species. It is apparently related to C. (Trachycardium) gatunensis Tonla, and is one of the early members of the C. muricatum group. Type locality. — 179. Other localities. — 225, 24?, and (doubtfully) near Guanica Central (Lobeck). Range. — Lares limestone, Cibao limestone, Los Puertos limestone. Cardium (Trachycardium) cinderellse Maury variety alter natum, new variety Plate XIX, Figures 2, 3, 4 Shell equivalve, slightly inequilateral; beaks high, strongly incurved; hinge line rather long; surface with about thirty-six broad, flat ribs with narrow interspaces; ribs appear medially grooved when worn smooth. When perfect, they show a single row of high, vaulted, inverted V-shaped spinose scales, which cover the width of the rib, and are somewhat unevenly spaced, though not crowded. There is a strong tendency toward alteration of ribs with large and ribs with small spines. This is more marked in some speci- mens than in others. It is a feature not mentioned in Maury's description of the Santo Domingo shell, but shows in the illustration of the latter. The interspaces are faintly grooved radially, and strongly transversely striate; valves internally grooved, and inner margins strongly scalloped ventrally. Length 29 mm., height 33 mm., diameter 26 mm. The sculpture is like G. cinderellce Maury, but the shell is larger, shorter, and more convex. It may possibly deserve a specific name, but the specimens are too fragmental to warrant this. Localities.— 225, 226, 280, 281, 331. Horizon. — Lares formation. 116 SCIENTIFIC SURVEY OF PORTO RICO Cardium (Trigonocardia) sambaicum Maury variety portoricoensis, new variety Plate XIX, Figures 5, 6 N^o complete shells were found, but fragments large enough to show the outline of the shell. They seem to be quite variable. There are some specimens in which the ribs are worn smooth as in Maury's illustration from Santo Domingo. Some specimens show the ribs slightly wider than the interspaces, but usually the reverse is true, especially in the ventral portion of the shell. Fine raised concentric threads cross the interspaces and cross the ribs between the nodes. These threads are much finer and more crowded where thev cross the ribs. In the interspaces they are very irregular, and frequently end short at the center of the interspace. These terminated threads have a tendency to alternate from opposite sides of the interspace as though shearing or faulting had displaced them. The nodes on the ribs are high and prominent. The shells are smaller and less convex than the Santo Domingo form, but the number of ribs, trun- cation and outline of the shell are about the same. Length 13 mm., height 13 mm., semi-diameter 5 mm. Locality. — 204. Horizon. — Quebradillas limestone. Cardium, new species (aff. C. sambaicum Maui*y) Plate XIX, Figure 7 Fragment indicating a shell of the size and general outline of C. sambaicum Maury, but with a somewhat different sculpture of the ribs. Shell with about 25 strong ribs, high and rounded on top, and slightly narrower U-shaped interspaces. The whole is crossed by fine, close concentric threads, quite irregular, and of which every second, third or fourtli one on crossing the ribs expand into broad semi-cone shaped scales the pieces of which are toward the beak. These scales are irregularly spaced, and are especially crowded near HUBBARD, TERTIARY M0LLU8CA FROM LARES DISTRICT 117 the ventral margin. Where the ribs are worn, these scales appear as thin arched lamellae. Length of shell abont 20 mm. Locality. — 204. Horizon. — Qnebradillas limestone. Cardium (Trigonocardia) haitense Soweiby variety cercadicum Maur}' Plate XIX, Figure 8 Cardium haitense Sowerby, Quart. Jour. Geol. Soc. London, VI, p. 52, PI. 10, fig. 11, 1849 (in part). Cardium {Trigonocardia) haitense variety cercadicum Maury, Bull. Amer. Paleont., V., No. 29, p. 212, PI. 36, fig. 6, 1917. This shell corresponds closely to Maury's variety cercadicum of Santo Domingo. The truncation is sharp; ribs with inverted V- shaped nodules ; ribs on the truncation have a few scattered spinose nodules; ribs flat, those anterior to the truncation having the flat tops sloping anteriorly, thus giving the ribs an asymmetric V-shaped cross-section. There are nine ribs on the truncation, and fourteen on the main body of the shell. Concentric threads in the interspaces, which, in the anterior part of the shell occasionally cross the ribs. The nodules on the ribs are rather low and indistinct except on the rib which marks the truncation, where they are prominent. Length 10 mm., height 11 mm., semi-diameter, etc., 5 mm. Localities.— 204:, 52(a), 192, 77? (Berkey). Horizon. — Qnebradillas limestone (characteristic), and in the Ponce limestone. Cardium (Trigonocardia) haitense Sowerby variety areciboense, new variety Plate XIX, Figure 9 Very close to C. haitense cercadicum Maury, but shorter and more sharply truncate, and with fewer, coarser ribs which are more de- 118 SCIENTIFIC SURVEY OF PORTO RICO cidedly angular. The ribs on the truncation near the dorsal margin have regularly spaced V-shaped spines, pointing sharply upward. The nodules and concentric threads are otherwise exactly like those of variety cercadicum. There are six ribs on the truncation and eleven on the remainder of the shell. Length 10 mm., height 14 mm., semi-diameter 6 mm. Locality. — 204. Also Berkey 77. Horizon. — Abundant in the Quebradillas limestone and occurring also in the Ponce limestone. Genus dementia Gray Clementia dariena Conrad Plate XIX, Figures 10, 11, 12 Meretrix dariena Conrad, Pac. R. R. Rep., V, p. 328, PI. G. fig. 55, lS5tJ. Clementia dariena Gabb, Jour. Acad. Nat. Sci. Phila. (2) VIII, p. 344, PI. 44, fig. 16, 16a, 1881. Clementia dariena Dall, Trans. Wag. Inst. Sci., Ill, pt. 6, p. 1235, 1903. Clementia dariena Toula, Jahrb. der K. K. Geol. Reichanst. Wien, LVIII, p. 725, PI. 27, figs. 9, 10, 1908. Clementia dariena Brown & Pilsbry, Proc. Acad. Nat. Sci. Phila., p. 371, PI. 28, fig. 1, 1911. The typical Porto Rican shells are smaller than those of the Panama region and Costa Rica, but otherwise show no marked dif- ferences. The Porto Pico shells are not dwarfed forms, but merely arrested in growth, the average adult being an exact replica of a Gatun shell when the latter is two-thirds grown. This might lead to the supposition that the Porto Rican analogue is a primitive form of the Gatun shell, except for the fact that the former occasionally reaches a size as great as the average Gatun shell and is exactly simliar, even to the loss of ribs near the ventral margin. To em- phasize this point, an average specimen from Dr. Kemp's Gatun collection (in the Columbia University museum) is illustrated here together with one of the large Porto Rican shells, for comparison. A comparison of the Porto Rican shells with available sources is shown in the following table. HUBBARD, TERTIARY MOLLUSC A FROM LARES DISTRICT 119 Table 2 Source Length mm. Height ] mm. ( Diameter Number mm. of prominent ribs . Breadth 3f ventral ribless zone mm. (meas. vertical) Gabb, illustration. Sapote, Costa Rica 57 47.5 33.5 26-35 10 Brown & Pilsbry, illustra- tion. Gatun 65 63 ? 26? IS Toula, illustration. Gatun. . 67 56 ? 28 19 Toula, description. Gatun. . 47 38 ? •? ? Kemp collection, specimen. Upper Gatun 63 61.5 36 33 23 Kemp collection, specimen. Ijower Gatun 51 43 25.5 31 7 do 48 44 24.5 30 10 do (young shell) 27 25? 14 22? Porto Rico, San Sebastian shale at CoUazo Falls .... 48 44.5 26 27 13 Porto Rico, San Sebastian calcareous shale at Robles. 41 34 19 25-30 0-5 do 39 31 23 do 38 33.5 20 do 38 35.5 18.5 do 37.5 33 19.5 do 35 32.5 20 do 35 32.5 17.5 do 35 33 17 do ? 34 20 do ? 33 15 do ? 32 20 do ? 29 17.5 - do ? 28 19 120 SCIENTIFIC SURVEY OF PORTO RICO It will be noticed from this table that there is considerable varia- tion in size and proportions among the specimens. This is due in part to distortion by pressure as regards the proportions. If the average of these measurements is taken and the height and diameter expressed in terms of the length as unity, we have the following- relationship : Length Gabb's illustration, S a p o t e, Costa Rica 1 Kemp's large specimen from the upper Gatun 1 Kemp's smaller specimens from the lower Gatun... 1 Large Porto Rico specimen from the Collazo district 1 Smaller Porto Rico specimens from the Robles district 1 Height 0.83 Diameter 0.59 0.98 0.88 0.93 0.87 0.57 0.51 0.54 0.51 Average length mm. 57 63 42 48 37 It will be noticed that in the larger specimens there is a relatively greater increase in the height and diameter, due to the slight change in shape of the shell with growth. The measurements from Gabb's illustration are of doubtful value in this comparison. The uniformity of size and number of ribs is a feature brought out in the table. The Porto Rican shells have about the same number of ribs as the Gatun shells, and in the ribbed stage are about the same size. The Gatun shells usually, and the Porto Eican shells occa- sionally grow beyond this stage with loss of ribs, an ontogerontic character. The Gatun shells were longer lived, apparently living under more favorable conditions than those of the upper San Sebas- tian shale in Porto Rico. Localities. — 232(b), 233, Collazo Falls, 83, 84, Juana Diaz. Horizon.— San Sebastian shale, Juana Diaz shale. Genus Cytherea Bolten Cytherea (Cytherea) berkeyi, new species Plate XX, Figures 1, 2, 3 Shell large, convex, very inequilateral; right valve with two posterior bifid and one anterior simple cardinal teeth; left valve with one middle bifid and anterior and posterior simple cardinals; left anterior papiUiform small HUBBARD, TERTIARY MOLLUSCA FROM LARES DISTRICT 121 out distinct; adductor scars and pallial line strongly impressed, the latter granulose; pallial sinus large, broad, rounded, ascending; inner margins minutely crenulate; small auxiliary adductor scars, situated close to dorsal margin between beaks and principal anterior adductor scars, and nearer to the latter; obscure, irregular impressed line in interior of each valve, descends vertically from anterior end of beak and dies out midway between beak and ventral margin; surface of valves marked by numerous small radial ribs, somewhat wavy, evenly spaced, and fairly uniform in size, but becoming coarser over the posterior third of the shell, where they alternate in strength, with occasionally two or three small ribs alternating with one lagrer one; the whole cut by fine, low, sharp, raised concentric lamellae, evenly spaced near the beaks, but gradually increasing to 2, 3, or 4 mm. apart toward the ventral margin, where they suddenly become irregularly spaced and crowded, apparently a gerontic character. These concentric lamellae are delicately fluted by the radials in the umbonal region. Minute growth lines cross the sculpture between the lamellae. The concentric sculpture is nowhere as prominent as the radial which increases in coarseness ventrally. The largest specimen, an internal mold collected by Dr. Berkey, measures in length 85 mm., height 68 mm., diameter 48 mm. A large shell of the same type as C. tarquinia Dall and also resem- bling C. lacerata Hanlej, a recent shell from China, and C. antiqua King, recent, from Chile and Patagonia. It differs from all of these in having the radial sculpture predominate over the concentric sculpture. Localities.- — 318, 41, 78, 98 (Berkey). Bayamon and Ponce (Lobeck). Horizon. — Quebradillas limestone Ponce limestone. Grenus Pitaria Romer Pitaria (Hyphantosoma) carbasea Giippy Plate XVI, Figure 2 Cytherea (Circe) carhasea Guppy, Quart. Jour. Geol. Soc, XXII, p. 292, PI. 18, fig. 13, 1866. Callista carbasea Gabb, Trans. Amer. Phil. Soc, XV., p. 250, 1873. Pitaria (Hyphantosoma) carbasea Dall, Trans. Wagner Inst. Sci., Ill, pt. 6, p. 1266, 1903. This shell occurs abundantly as internal molds, but only one frag- ment of external mold was found showing the delicate sculpture. The pallial sinus is sub-angular, deep, reaching nearly to the middle of the valve; interior of shell smooth; exterior surface marked by fine, somewhat wavy or irregular concentric lamellae crossed by fine L^LI ^R AR 122 SCIENTIFIC SURVEY OF PORTO RICO divaricate lines, strongest anteriorly, and forming two zigzag zones near the middle of the valve. These divaricate lines, when viewed under a lens are seen to consist of rows of pits in the lamellae; limnle large, demarked by fine sharp impressed line, and crossed faintly by -the concentric lamellae, thus being rendered indistinct. The shells vary considerably in size, the largest internal mold measuring in length 32 mm., height 26 mm., diameter 16 mm. Type localities. — 64, 225. Other localities.— Q19, 233, 343, 232(b). Range. — San Sebastian shale, Lares limestone, and Cibao lime- stone. Abundant in the Cibao limestone. Genus Chione Megerle von Miihlfeldt Chione woodward! Guppy Plate XIII, Figure Y Venus Woodwardi Guppy, Quart. Jour. Geol. Soc. London, XXII, p. 292, PI. 18, fig. 1, 1866; Idem. XXXII, p. 530, 1876. Chione Woodwardi Dall, Trans. Wagner Inst. Sci., Ill, pt. 6, p. 1291, 1903. Chione Woodtoardi Maury, Bull. Amer. Paleont., V, No. 29, p. 218, PI. 37, fig. G, 1917. This is one of the most abundant fossils in the Quebradillas lime- stone. The shells show no differences from those described from Jamaica and Santo Domingo. Localities— 204:, 217, 52(a), 670, 179, 225. Xear Bayamon (Semmes and Lobeck). ^N'ear Guanica Central (Lobeck). Vieques Island (Lobeck). 77 (Berkey). Culenbrinas Point, Ponce, and K. 2.4 Ponce- Ad juntas road (Mitchell). Horizons. — Lares limestone ?, Quebradillas limestone, Ponce lime- stone. Chione (Lirophora) hendersoni Dall Venus paphia Guppy, Quart. Jour. Geol. Soc. London, XXII, p. 292, 1866. Not of Linn6. Chione paphia Gabb, Trans. Amer. Phil. Soc, XV, p. 249, 1873. Venus paphia Guppy, Geol. Mag., decade 2, I, p. 442, 1874, etc., ex parte.; Quart. Jour. Geol. Soc, XXXII, p. 530, 1876. HUBBARD, TERTIARY MOLLUSCA FROM LARES DISTRICT 123 Chione (Lirophora) hendersoni Dall, Trans. Wagner Inst. Sci., Ill, pt. G, p. 1295, PI. 55, fig. 22, 1903. Chione (Lirophora) hendersoni Maury, Bull. Amer. Paleont, V, No. 29, p. 219, PI. 37, fig. 8, 1917. The Porto Rican shells are apparently closer to the Santo Domingo than to the Bowden shells. The specimens show the obscure crenula- tion at base of ribs, and the sudden thickening of the ribs over the anterior portion of shell. There are eleven or twelve large, widely spaced ribs. They are obsolete in the umbonal region. Interior of shell has deeply impressed pallial line and adductor scars; pallial sinus short and angular. An average specimen measures in length 31 mm., height 22 mm., diameter 14.5 mm. The largest measures in length 36 mm., height 27 mm., semi-diameter 8 mm. Localities.— 179, 331, 343, 64, 619 ?. Range. — Lares limestone to Los Puertos limestone. Genus Tellina (Linne) Lamarck Tellina strophoidea, new species Plate XX, Figures 4, 5 Shell moderately elongate, sub-equilateral; beaks inconspicuous; surface with numerous elevated, sharp concentric lamellae, which, on adult specimens, frequently merge on the posterior part of shell into fewer, coarser lamellae; numerous radial striae can be seen under a lens in transverse light. Ven- trally they are interrupted by the lamellae, but toward the beaks they become more distinct and continuous, so that on the nuclear portion of the shell they are nearly as strong as the concentric sculpture, resulting in an extremely fine cancellate ornamentation; posterior slope with two radial folds and a moderately deep sulcus between. The upper fold is narrow-er and less pro- nounced; posterior dorsal margin marked by accentuation of the lamellae into a slight ridge. In the adult shells the lowest fold is, in the right valve, double, with a slight sulcus between. This is only noticeable at the distal end of the fold. The lamellae bend sharply upward on crossing the lower fold; lunule extremely narrow; interior of shell unknown. Length 48 mm., height 27 mm., semi-diameter 6 mm. This shell is nearest to T. strophia Dall, which it resembles in sculpture, but is considerably less elongate. Localities.— 204., 216, 98(Berkey). Range. — Throughout the Quebradillas limestone. 124 SCIENTIFIC SURVEY OF PORTO RICO Tellina portoricoensis, new species Plate XX, Fignre 6 Shell resembling T. strophoidea but with much finer concentric sculpture. Shell sub-equilateral, slightly convex; beaks low; surface with fine, sharp, hardly raised concentric lamellae, at first uniform and evenly spaced, but ventrally becoming slightly irregular, with every fifth or sixth one more prominent than the others; low, inconspicuous double fold on the umbonal ridge, becoming almost obsolete distally, and over which the concentric lamellae are flexed sharply upward in crossing; fine obscure growth lines between the lamellae; faint radial rays seen only in oblique light; interior of shell unknown. Length 42 mm., height 23 mm., semi-diameter 5 mm. Locality. — 204. Horizon. — Quebradillas limestone. Tellina (Scissula) grabaui, new species Plate XVI, Figures 3, 4 Shell small, elongate, slightly convex, sub-equilateral; anterior rounded, posterior roundly truncate; surface with fine growth lines, periodically in- cised, replaced on the umbonal ridge by widely spaced, sharp concentric lamellae, which cross a faint sulcus above the umbonal ridge and, decreasing in size and strength, continue to the dorsal margin; growth lines crossed by faint, wavy oblique lines, becoming more distinct as they approach the umbonal ridge, where they end suddenly, in contact with the concentric lamel- lae. These oblique lines begin near the anterior margin and slope downward toward the posterior. Interior of shell unknown. Length 20 mm., height 11 mm., semi-diameter 2.5 mm. This shell is close to T. (Scissula) lampra Uall, of the Chipola Oligocene. It is also close to T. rowlandi Toula, a species from the Gatun formation. T. carihea d'Orbigny seems to be the living rep- resentative of these shells. T. (Scissula) cercadica Maiiry, a shell from the Santo Domingo Oligocene, is a similar but less elongate form. Locality. — 204. Horizon. — Quebradillas limestone. Tellina, species aff. T. (Angulus) atossa Dall Plate XI, Figure 7 A single left valve of a Tellina of uncertain identity. Shell moderately convex, showing prominent lateral teeth ; surface with HUBBARD, TERTIARY M0LLU8CA FROM LARES DISTRICT 125 fine sharp concentric lamellae, strongest near the ventral margin, and obscure growth lines between the lamellae. The latter are ac- centuated on the anterior slope; shallow radial sulcus located next to the posterior dorsal margin. Interior of shell unknown. In out- line and general appearance, it resembles T. (Angulus) atossa of the Tampa Silex beds. Length about 20 mm,, height 14 mm., semi- diameter 4 mm. Locality. — 204. Horizon. — Quebradillas limestone. Genus Metis H. and A. Adams Metis trinitaria Dall Plate X, Figure 7 Metis trinitaria Dall, Trans. Wagner Inst. Sci., Ill, pt. 5, p. 1041, PI. 46, fig. 24, 1900. cf. TelUna Uplicata Guppy, Quart. Jour. Geol. Soc, XXXII, p. 530, 1876. Not of Conrad. Metis trinitaria Maury, Bull. Amer. Paleont., V, p. 226, 1917. The specimens differ somewhat from Dall's description, being more nearly equilateral, and the pallial sinuses of the two valves being subequal and of equal height. The shell seems to be quite variable, however. Ventral margin arcuate, with broad median sinus in the right valve: sharp, rounded posterior fold with a second obscure fold higher up; surface marked bv growth lines and faint radial striations, the latter fading out toward the lateral margins. There is some variation in the size and proportions as shown by the following measurements of internal molds: Length 52 mm., Height 44 mm., Diam. 23 mm. Length 54 mm.. Height 44 mm., Diam. 19 mm. Length 48 ram., Height 39 mm., Diam. 16 mm. Some internal molds of this shell, collected in Porto Rico in 1915 by Dr. D. R. Semmes, were shown to Dr. Dall, who identified them as M. trinitaria. This species is reported from Trinidad and Cuba, and doubtfully from Santo Domingo. Specimens from Sombrero, 126 SCIENTIFIC SURVEY OF PORTO RICO W. I., in the Columbia University collections seem to be of this species, but are considerably larger than the Porto Rican shells. Localities.— 204:, 678, 61, 319, 801. 98(Berkey). San Juan district (Semmes). Culebrinas Point (Mitchell). Horizon. — This shell is the chief horizon marker of the Quebra- dillas limestone along the north coast. The internal molds are found in great numbers in many localities. It occurs rarely in the Los Puertos limestone, and Ponce limestone. Genus Solen Linne Solen (Plectosolen) collazoensis, new species Plate XI, Figure 8 Shell produced anteriorly, with beaks 7.5 mm. behind the anterior margin; both ends roundly truncate; height slightly less in middle of shell than at the ends; beaks inconspicuous; surface marked only by growth lines, strengthened on anterior slope, and a narrow furrow extending from the beaks to the anterior-ventral corner. Length 42 mm., height 11 mm., diameter 6.5 mm. This is a small straight shell of the section Plectosolen, having a strong resemblance to the Eocene forms of the Paris Basin, such as S. plagiaulax Cossmann, and S. laversinensis Lef. and Wat. It is probably of the type of S. ohliquus Sowerby. It also resembles n small S. ohliquus Spengler, but is a true Plectosolen. The description is based on a single specimen from the black shale at Collazo Falls. The specimen is badly fractured, though nearly complete. Localities. — 261, Juana Diaz, Yauco. Horizon. — San Sebastian shale, Juana Diaz shale. Genus Psammosolen Risso Psammosolen sancti-dominici Maury Psammosolen sancii-dominici Maury, Bull. Amer. Paleont., V, No. 29, p. 228, PI. 37, fig. 13, 1917. Only two fragments were found, but the marking is distinct and characteristic, and leaves no doubt as to the identity. One specimen seems to be about the same size as the Santo Domingo specimens HUBBARD, TERTIARY M0LLU8CA FROM LARES DISTRICT 127 described by Maury. The other specimen is smaller. The broad median sulcation is barely noticeable. Locality. — 204. Horizon. — Quebradillas limestone. Genus Semele Schumacher Semele, species indet. Shell equivalve, nearly equilateral; beaks low; ends of shell rounded; anterior and posterior dorsal margins make angle of 125°; umbonal ridge low, obscure; escutcheon long and narrow; lunule short, and same width as the escutcheon; surface sculpture of fine, even, sharp concentric lamellae; about fourteen to every 10 mm.; no radial striation visible. Length 35 mm., height 30 mm., diameter 14 mm. This may be a new species, but the specimens are poor and do not permit an exact description, being known only from internal molds and a few fragments of external mold. The sculpture is finer than that of ^S'. claytoni Maury. The shell resembles *S. chipolana Dall, but is more elongate. Locality. — 225. Horizon. — Lares limestone. Genus Thrachia Blainville Subgenus Cyathodonta Conrad Cyathodonta reedsi Maury Shell equivalve, sub-equilateral, moderately convex; anterior dorsal slope slowly descending, anterior rounded, posterior roundly truncate; sculpture of about twenty-five rounded to sub-angular concentric ripples, markedly wavy or irregular, and owing to the thinness of the shell, equally strong in the internal molds. The ripples have a strong tendency to become oblique on the anterior third of the shell; umbonal ridge low, rounded, beyond which the ripples die out, leaving the posterior dorsal smooth except for growth lines. A right valve shows a low obscure radial fold about one-third the distance from the umbonal ridge to the dorsal margin; beaks rather high and strongly incurved. Gerontic individuals show a complete loss of con- centric ripples in the final stages of growth. Adult shell measures in length 27 mm., height 19 mm., semi-diameter 6 mm. This shell is apparently nearest to C. gatunensis Toula but is more 128 SCIENTIFIC SURVEY OF PORTO RICO elongate, more nearly equilateral, and the smooth posterior slope higher and more produced. It is not as convex as the Gatun shell. I'ype Locality. — 204, Other locality.— 226 I liange. — Lares ? limestone to Quebradillas limestone. Genus Corbula Lamarck Corbula collazica Maury Localities. — Collazo Falls, and doubtfully at 120 and 225. Horizons. — Typical of the San Sebastian shale, and "occurs doubt- fully as high as the Lares limestone. Genus Teredo Linne Teredo incrassata Gabb Kuphus incrassatus Gabb, Trans. Amer. Phil. Soc, XV, p. 246, 1873; Jour, Acad. Nat. Sci. Phila., new ser., VIII, p. 342, PI. 44, fig. 12a-e. Teredo fistula? Guppy, Quart. Jour. Geo!. Soc, XXXII, p. 529, 1876. Teredo incrassata Dall, Trans. Wagner Inst. Sci., Ill, pt. 6, p. 1587, 1903. Teredo incrassata Maury, Bull. Amer. Paleont., V, No. 29, p. 235, PI. 39, fig. 24, 1917. The tubes vary in diameter, but average 20 mm. They attain considerable length, but are usually found broken in short pieces. Localities.— 8G, 10, Collazo Falls, 758, 162, 182, 179, 332, 619, 621, 616, 225, 23, 335, 330, 324, 74, 370, 749(a), 334, Horizons. — San Sebastian shale, Lares limestone, Cibao limestone, Los Puertos limestone, Quebradillas limestone. Extremely abundant. Outside the Lares District, this species has been found by Lobeck on Vieques Island (south and east sides), and in basal Tertiary beds near Yauco, P. R, It has been found by Berkey at localities 10, and 78, and by Mitchell in the Ponce formation and Juana Diaz shale. Genus Dentalum Linne Dentalium, species indet. Shell long, slender, nearly straight, slowly tapering, circular in cross- section; shell thick, the aperture being only about one-half the diameter of the tube; surface polished, and marked with numerous fine, very regular. HUBBARD, TERTIARY MOLLUSGA FROM LARES DISTRICT 129 longitudinal threads, visible only under a hand lens and transverse light. The threads appear, under a lens, to he minutely beaded, but under a micro- scope, this appears to be the result of fine raised concentric growth lines, strongest in the interspaces between the longitudinal threads. There are about one hundred longitudinal threads on a tube of 1 mm. diameter. The shell structure is strongly fibrous, the fibrous striations appearing on the longitudinal threads as seen under the microscope. The apex is not shown in any of the specimens. The largest fragment, with a diameter of 1 mm. measures 13 mm. in length. The tapeiing of the tube is hardly discernible in this length. Many fragments were found in the lower black shale of a minute Dentalium, the largest of which is only about 1 mm. in diameter, and the majority of which average less than I/2 mm. Locality. — 261. Horizon. — San Sebastian shale. Genus Calliostoma Swainson Calliostoma portoricoensis, new species Plate XXI, Figure 1 Whorls seven, including a rather high protoconch of two smooth, round whorls; remaining whorls flat; suture narrow but deeply impressed; surface of whorls with six spirals and strongly marked oblique growth lines which cut the spirals, giving them a wavy appearance. The five uppermost spirals are small, sharply raised, with wider concave interspaces. The two spirals immediately below the suture are somewhat smaller and show a tendency to group together. The sixth spiral, immediately above the suture, is very much larger, broad, and bluntly rounded, and forms the peripheral keel; base with eight prominent, broad, flat spirals and fine raised oblique growth lines crossing the spirals and grooves between. Aperture and umbilical area not visible in the specimen. Length 11 mm., diameter 9 mm. A sino-le specimen M^as found of a Calliostoma remarkable for its elongate conical base (more than two-thirds as long as the spire). Locality. — 204. Horizon. — Quebradillas limestone. Genns Turbo Linne Turbo fetkii, new species Plate XXI, Figure 2 Shell small; whorls six; the first two flat topped, strongly keeled; the remaining whorls sub-round, but with slight angulation and broad flat 130 SCIENTIFIC SURVEY OF PORTO RICO shoulder, nearly one-third the axial width of the whorl; suture deeply channelled; rounded spirals of two to three different sizes, increasing by intercalation, with total of about twenty on the body whorl, and of these five are very prominent; entire surface crossed by oblique incremental lines, strongest between the spirals; spirals and incremental lines visible in the sutural channel. Height 13 mm., diameter 11 mm. The al)ove description is based on a single specimen, in which the anterior portion of the body whorl is missing. The shell is remark- able for its extremely deep, squarely channelled suture. The spirals are smooth or Avith microscopic creuulation produced by the crossing of the growth lines. The spiral anterior to the suture is never promi- nent. This shell is like T. rliectogrammicus Dall, of the Florida Pliocene, but differs from the latter chiefly in its smaller size and its deeply channelled suture. Its suture is like that of T. .^penglerianus Gmelen, a recent shell from the West Indies, but the hitter is much larger and has more spirals. This shell is named in honor of Dr. C. R. Fetke, who has w^orked on the Geology of the eastern end of Porto Eico. Locality. — 313. Horizon. — Quebradillas limestone. Turbo dominicensis Gabb variety quebradillensis, new variety Shell larger than the Santo Domingo variety, T. dominicensis laloi Maury, the internal mold measuring in height (with tip of spire missing) 37 mm., maximum diameter 33 mm. Like the Santo Domingo variety, the final whorl is angulated, as presumably are the earlier whorls, though this cannot be ascertained from the specimen. It is more like Gabb's type, however, in having the majority of the spirals creuulate. On the final whorl the spiral anterior to the suture is more strongly crenulate than the others. Anterior to this, on the shoulder, are four minor crenulate spirals, the two central ones being slightly more prominent than the other two. At the shoulder angulation is a very strong, plain spiral, which projects sharply upward, above the surface of the shoulder. The remainder of the whorl is orna- mented with eight or more strong sinujle spirals, each alternating with a single minor crenulate spiral. The basal portion of the whorl is not pre- served. From the small i)ortion visible in the specimen, it seems that the ornamentation of the earlier whorls is similar, but with fewer spirals. The outer lip is Urate for a short distance, there being thirteen lirae and a pos- terior notch below the shoulder angulation. This shell was found among the material collected by Dr. Lobeck in the Quebradillas limestone near Bayamon. It consists of a single HUBBARD, TERTIARY MOLLUSCA FROM LARES DISTRICT 131 internal mold and a portion of the external mold of the same speci- men, which shows the sculpture of the final whorl and a small por- tion of the spire. A few internal molds, possibly of the same species were found in the Quebradillas limestone of the Lares district. Locality. — Kear Bayamon (Lobeck), 204? Horizon. — Quebradillas limestone. Genus Liotia Gray Liotia (Arene) coronata Dall variety portoricoensis, new variety Plate XXI, Figure 3 Shell sman, five whorls, including flat, smooth naticoid protoconch of two whorls. Two spiral keels appear on the third (or first post-nuclear) whorl. A third spiral appears on the fourth whorl just above the suture; suture deeply channelled; shoulder of third and fourth whorls marked by fine beaded spiral ridge immediately below the suture. Fine growth lines crossed the shoulder and produce a slight beading of the uppermost spiral keel; final whorl smooth except for strongly incised growth lines, which are stronger than the almost obsolete spirals. The final whorl tends to lose the coiling habit, becoming almost free from the preceding whorl. Only a single specimen was found, and the aperture and base of the shell are unknown. Height 5 mm., diameter 5 mm. The first four whorls of this shell are quite similar to Dall's Tampa species. In the Porto Rican shell the longitudinal wrinkles, characteristic of the Tampa shell, become obsolete on the fourth whorl, but reappear strongly on the last whorl. The Porto Rican variety seems to have been derived from the Tampa species, and is an interesting example of a phylogerontic form. Locality.— 2Q4:. Horizon. — Quebradillas limestone. Genus Xenophora Fischer de Waldheim Xenophora conchy liophora (Born) TrocJiiis conchyliophorus Born, Mus. Caes. Vind. p. 333, 1778. Trochus agglutinans Lamarck, Anim. sans Vert., VII, XIV. Phorus agglutinans Gabb, Trans. Amer. Phil. Soc, XV, p. 241, 1873, 132 SCIENTIFIC SURVEY OF PORTO RICO Xenophora conchyliophora Dall, Trans. Wagner Inst. Sci., Ill, pt. 2, pp. 360- 362, 1892; Bull. 90, U. S. Nat. Mus., p. 105, PI. 15, figs. 1, 3, 1915. Xenophora conchyUophora Maury, Bull. Amer. Paleont., V, No. 29, p. 133, PI. 23, fig. 7, 1917. The Porto Kican specimens show no differences from the typical forms of this species. Localities.— W^, 225, 24. Range. — Lares limestone to Quebradillas limestone. Very abnn- dant in the Quebradillas. Genus Neritina Lamarck Neritina (chipolana var.?) collazoensis, new species Plate XXI, Lig-nre 4 Shell small, fragile, polished, with low spire, and apparently three whorls. A single specimen was obtained, in which the spire has been crushed and the aperture is not visible. The outline of the shell and the surface mark- ings are like N. chipolana Dall, but with the markings more crowded and numerous, and all being discontinuous. These lines show, however, the same arrangement in groups of three or four, with zig-zags and curves. The curves are frequently convex in opposite directions in adjacent lines, leaving circular areas between the lines, a feature which is also characteristic of the Chipola species, judging from Ball's illustration (Wagner Inst., Ill, pt. 4, PI. 23, fig. 19, 1898). The lines are very narrow, and a dark reddish purple in color. The shell is slightly smaller than the Chipolan form, the height being 4 mm. and the maximum diameter 4.5 mm. In places where the epi- dermis has been chipped off, exceedingly fine and very regular growth lines may be seen with a lens. This shell was obtained from a stratum of lignitic shale in the Collazo Falls section, and is associated with fossil leaves and minnte, thin-shelled bivalves, apparently fresh water forms. This stratum is intercalated in marine beds, and is of interest in showing the con- ditions existing in this locality during deposition of the San Sebas- tian shale. Locality. — 261. Horizon. — San Sebastian shale. Genus Hipponyx Defrance Hipponyx portoricoensis, new species Plate XXT, Figures 5, 6 Shell small, obliquely conical; apex at or near posterior extremity, often overhanging, and slightly recurved; surface with irregular bifurcating radial HUBBARD, TERTIARY MOLLUSCA FROM LARES DISTRICT 133 ribs and less prominent fluted concentric lamellae; Interior with faint but large horse-shoe shaped scar. This shell is very variable, but in all specimens the apex is blunt and rounded. Some specimens show hardly anv recurvature to the apex, while in others this feature is marked, and in all cases it is to be left. A strongly recurved specimen measures in diameter of aperture about 7 mm., height of apex above base 4 mm. A low flat shell without curvature measures in diameter 10 mm., height of apex 3 mm. The largest specimen measures in diameter 11 mm., height of apex 6.5 mm. Localities. — 204, 220, 23 ?. Range. — Los Puertos limestone to Quebradillas limestone. Genus Calyptraea Lamarck Calyptrsea cf. centralis Conrad Infundibulum centralis Conrad, Am, Jour. Sci., XLI, p. 348, 1841; Med. Tert, p. 80, PI. 45, fig. 5, 1845. Trochita centralis Emmons, Geol. N. C, p. 276, fig. 193, 1858. InfundiMiluvi concentricum H. C. Lea, Trans. Amer. Phil. Soc, (2), IX, p. 249, PI. 35, fig. 39, 1845 (young shell). Grochita collinsii Gabb, Jour. Acad. Nat. Sci. Phila., (2), VIII, p. 342, PI. 44, figs. 11, 11a, 1875. Infundibulum candeanum d'Orbigny, Moll. Cuba, II, p. 190, PI. 24, figs. 28, 29, 1842. Galerus candeanus Dall, Hemphill's Shells, p. 335, 1883. Galerus parvulus Dunker, Jahrb. Deut. Mai. Ges., p. 244, 1875. Calyptrea candeana Dall, Blake Rept., p. 284, 1889. Calyptrea centralis Dall, Trans. Wagner Inst. Sci., Ill, pt. 2, p. 353, 1892. A single external mold was found which is apparently this species. Diameter of base 8 mm., height of apex 4 mm. Locality. — 204. Horizon. — Quebradillas limestone. Genus Crucibulum Schumacher Crucibulum auricula Gmelin variety portoricoensis, new variety Plate XXI, Figures 7, 8 Cup typical of C. auricula Gmelin. Surface with fine. Irregular, dichoto- mus radial lines, crossed by fine wavy growth lines. Form varies from 134 SCIENTIFIC SURVEY OF PORTO RICO high, acute apex to low apex with broad base. Cup uniform in size and shape in all the specimens. A shell with low apex measures in diameter of base 20 mm. height of apex 7 mm. Localities. — 204. (also Berkey 98). Horizon. — Quebradillas limestone (fairly abundant). Crucibulum auricula? variety Plate XXI, Figure 9 This is probably one of the many variations of C. auricula. Only a few fragmental specimens were obtained, and the interior is un- known. Apex sub-central ; coarse radial ribs, somewhat irregular, bifur- cating and trifurcating. Diameter 14 mm., height of apex 6 mm. Locality. — 204. Horizoji. — Quebradillas limestone. Crucibulum (Dispotsea) collazum, new species Plate XXI, Figure 10 Shell small, smooth; apex eccentric, sharp, strongly gyrate in the normal direction; basal margin sub-circular, with large rounded sinus on the side below the apex. On either side of the sinus the margin tlaies, or flattens * out, and is thickened, being analogous to the thickened outer lip or varix of coiled gastropods; cup large, deep, and about one-half free from the shell wall. Most of the free half has been broken away in the specimen illustrated, but when perfect, it has a tortuous, or S-shaped curvature, and overhangs the apex cavity, being suggestive of the internal appendage of a Calyptrcea. Diameter of aperture 7 mm., height of apex 4 mm. Two specimens were collected, both showing the marginal sinus, which is therefore assumed not to be an accident of growth. The shell illustrated appears to be an adult specimen. The other shell is smaller, with a correspondingly smaller marginal sinus. Localities. — 84, 261. Horizon. — San Sebastian shale. Genus Natica Scopoli Natica canrena? (Linne) Moercli Neritacanrena (Linng in part) Auct., Moerch, Malak. Blatt., XXIV, p. 62. 1877. Natica canrena Gabb, Trans. Amer. Phil. Soc, XV., p. 223, 1873. HUBBARD, TERTIARY MOLLUSCA FROM LARES DISTRICT 135 Natica canrena Guppy, Quart. Jour. Geol. Soc, XXXII, p. 518, 1876. Natica canrena Dall, Trans. Wagner Inst. Sci., Ill, pt. 2, pp. 364-365, 1892. Natica canrena Brown and Pilsbry, Proc. Acad. Nat. Sci. Phila., p. 508, 1912. Natica canrena Maury, Bull. Amer. Paleont., V, No. 29, p. 134, Pi. 23, fig. 10, 1917. A small Natica was found which is probably the young of N. canrena. The specimen shows on the last whorl the faint oblique striations, which start at the suture. Diameter 7 mm. Several in- ternal molds, possibly of this species, were found in the Cibao lime- stone at locality 22. The largest measures in height 22 mm., diameter 25 mm. Locality. — 204. Horizon. — Quebradillas limestone. Natica (AmpuUina?) collazoensis, ne\v species Plate XXI, Figures 11, 12 Shell large, moderately high spired; suture deeply impressed; whorls six to eight; umbilicus partly covered by callus, or entirely open in some speci- mens, in which the callus may have been broken away. Some specimens show the last whorl strongly flattened below the suture, while in others the whorl is evenly rounded. This is probably a sex difference; shell thin; sur- face with numerous fine, impressed spiral lines, and much finer, almost microscopic growth line threads, periodically incised, marking a resting stage in the growth of the shell. The growth lines are remarkable for their even- ness. Largest specimen measures in height 6G mm., diameter, 52 mm. The shell in size and general appearance resembles Ampullina amphora Heilprin, and is probably related to the large Eocene forms. It has some resemblance to Xatica species indet. Brown & Pilsbry (Proc. Acad. Xat. Sci. Phila., p. 360, PL 25, figs. 1, 2, 1911), a Gatun shell. • Localities.— 262, 265, 86 (base of falls), Collazo Falls, 264 (bot- tom 3rd falls), 83, 235. Horizon. — San Sebastian shale, (very abundant). Genus Epitonium Bolten Epitonium (Cirsotrema) collazoensis, new species Plate XXII, Figures 1, 2 Shell with three and one-half whorls measures in height 25 mm., maximum diameter 13 mm.; whorls rounded; suture distinct, but not deep; whorls with about twenty-four slightly oblique, moderately strong longitudinal ribs 136 SCIENTIFIC SURVEY OF PORTO RICO and two opposite rows of sub-continuous varices, the last varix being at the outer lip; longitudinal ribs fluted by another series of oblique ribs of equal strength, descending to the left at about 45°, continuing from suture to suture, and being more or less interrupted by the longitudinal series of ribs. At base of body whorl, both series of ribs terminate against a strong, rugose or rope-like spiral, which encircles the umbilical area, and is interrupted only by the varices. Over the entire surface of the shell is a fine microscopic cloth- like texture composed of three intersecting series of threads, one series vertical, another parallel with the 45° ribs, and the third series descending to the right at about 45°. The three series are named in order of their relative strength, the vertical series being strongest, and locally obscuring the other two series. Aperture circular. In general appearance the shell is typical of the high spired forms of this group. A single decollate specimen was found of an Epitonium which is remarkable for its microscopic texture, which is somewhat of the nature of that of E. fextuvestitum Maury, but the shell is quite different from the latter. Apparently the nearest relatives of this shell are to be found in the Paris Basin beds. Two such forms, ^ery similar, though smaller, are Cirsotrema subspinosum (Grat.) of the Aquitanien, and C. hourgeoisi de Boury, of the Helvetian. This relationship is of considerable significance in considering the age of the San Sebastian shale. Locality. — K. 29,2-3 Lares road. Horizon. — San Sebastian shale. Genus Turritella Lamarck Turritella tornata Guppy variety portoricoensis, new variety Plate XXT, Figures 13, 14 Shell corresponds with Guppy's description, except that in the last few whorls the anterior keel is double, and there are frequently as many as four spiral threads in the median concavity. There are also from one to two threads on the shoulder and likewise on the slope below the lower keel. The base is spirally threaded and marked by accentuated growth lines. The beading of the spirals is lacking or obscure, due probably to the poor state of preservation of the specimens. All specimens are decollate. The largest has a maximum diameter of 14 mm. and height (4 whorls) of 2G mm.: the average sized specimen has a niaxinmni diameter of 10 mm. This species is extremely abundant in the San Sebastian shale. Xearly all specimens are ])oorly preserved casts or internal molds, and the ornamentation is difficult to make out. HUBBARD. TERTIARY MOLLUSCA FROM LARES DISTRICT 137 l\jpe locality.— 2S2(h). Other localities.— 225, 331, 333, 343, Collazo Falls, 226, 262, S3, 233. Range. — San Sebastian shale to Lares limestone. Turritella planigyrata Guppy T. planigyrata Guppy, Proc. Sci. Assoc. Trinidad, p. 169, 1867; Geol. Mag., p. 408, PI. 18, fig. 5, 1874. T. planigyrata Gabb, Trans. Amer. Plail. Soc, XV, p. 240, 1873. T. planigyrata Guppy, Quart. Jour. Geol. Soc, XXXII, p. 519, 1876. T. planigyrata Maury, Bull. Amer. Paleont., V, No. 29, p. 129, PI. 22, fig. 14, 1917. A fragment comprising the last three whorls. Height 14 mm., maximum diameter 8.5 mm. The specimen agrees with the de- scriptions by Guppy and Gabb, and there seems to be no doubt of tho identit}'. Locality. — 234. Horizon. — San Sebastian shale. Turritella mitchelli, new species Plate XXII, Figure o A single decollate specimen with seven and one-half whorls. Shell large, high spired (apical angle 5°); whorls flat and continuous; suture indistinct or invisible; whorls marked by three prominent crenulated spirals and minor, minutely beaded spiral threads; the whole surface covered by minute spiral striae and longitudinal growth lines, the latter producing the crenu- lation in the spirals. In front of each suture is a flat area, less than one-third the axial width of the whorl, marked by two to three obscure spiral threads; next in order come a single primary spiral, followed by two fairly prominent secondary spirals in the median portion of the whorl. The anterior third of the whorl is marked by the two most prominent primary spirals with a narrow channel between them, giving the effect of a doubled spiral band. The anterior member of this pair is slightly larger than the other on the last two whorls. Of the microscopic sculpture above mentioned, the soiral striae are the most prominent on the primary ridges, and obsolete in the interspaces, whereas the reverse is true of the growth lines, which results in the strongest beading being on the minor spirals in the interspaces. The base is suleated below the anterior primary spiral, and is moderately convex anterior to this sulcus. The surface of the base is marked only by the spiral striae and growth lines, which here are markedly wavy or vermicular. The 138 SCIENTIFIC SURVEY OF PORTO RICO suture is distinct on the last two whorls, but on the spire it is invisible, being concealed by the anterior primary spiral, which overhangs it. Height 59 mm,, maximum diameter 22 mm. As the protoconch and upper portion of the spire are missing, it is difficult to trace the relationship of this shell ^^^th other species. The succession and arrangement of spirals is similar to T. halensis Dall, a shell from the Flint River Oligocene. The finer sculpture and beading resemble T. altilira Conrad, of the Gatun formation, and T. calostemma Pilsbry & Brown, of the Oligocene of Haiti. This shell is undoubtedly a member of the T. tornata — T. altilira group, and is probably closest to T. calostemma. Berkey found a similar, and apparently related shell from the basal Tertiary near Juana Diaz. It differs in having four equally strong major spirals. This shell is named in honor of Dr. G. J. Mitchell. Locality. — Collazo. Horizon. — San Sebastian shale. Turritella portoricoensis, new species Plate XXI, Figure 15 Shell small; spire acute; about ten rounded whorls with about fifteen fine spiral threads with sub-equal intervals; suture distinct, but not channelled until the last three whorls; whorls at first evenly rounded with spiral threads of equal strength, but later becoming angulated by accentuation of spiral threads into a keel at periphery of whorl and another at anterior end of whorl, overhanging the suture; first three whorls loosely coiled; proto- conch missing on all specimens. The angulation begins at about the seventh whorl. A specimen with nine whorls measures in height 13 mm., maximum diameter 4 mm. The yoimg of this shell has the whorls of 2\ pleheia Say, and in the adult, the whorls are like those of T. cequistriata Conrad, in some respects. The later whorls also resemble T. subannulata acro- pora Dall, but are not quite as angular as in the latter species. The nearest Antillean forms are T. planigyrata Guppy, of Santo Domingo, and T. gatunensis Conrad, of Panama. The Porto Rican shell is probably a less specialized member of this group. Locality. — 204. Horizon. — Quebradillas limestone. HUBBARD, TERTIARY MOLLUSCA FROM LARES DISTRICT 139 Turritella berkeyi, new species Plate XXII, Figure 4 Shell extremely high spired, slender; whorls twenty -one; earliest whorls simple, with two keels of equal size; anterior and posterior slopes of whorls straight and of equal width; simple rounded median depression between the two keels; last ten whorls with the posterior keel becoming double and the shoulder of the whorl becoming gently convex, features which are quite promi- nent in the adult stage; beginning with the eighth or ninth whorl an obscure spiral appears on the anterior extermity of the whorl just above the suture and partially overlapped by the succeeding whorl; surface throughout is lacking in ornamentation. Height 73 mm., maximum diameter 11 mm. This shell may possibly be a primitive form of T. perattennuata Heilprin. It resembles T. alcida Dall, b\it is very much more slender. Other somewhat similar forms are T. altilirata Conrad, and T. variabilis variety alticosta Conrad. Locality. — 204. Horizon. — Quebradillas limestone. Genus Petaloconchus Lea Petaloconchus domingensis Sowerby Petaloconchus domingensis Sowerby, Quart. Jour. Geol. Soc, VI, p. 51, PI. 10, fig. 8, a, b, c, 1849. Petaloconchus sculpturatm Gabb, Trans. Amer. Phil. Soc, XV, p. 240, 1873. Not of Lea. Petaloconchus sculpturatus Guppy, Quart. Jour. Geol. Soc, XXXII, p. 519, 1876. Vermetus {Petaloconchus) sculpturatus Dall (in part). Trans. Wagner Inst. Sci., Ill, pt. 2, pp. 305, 306, 1892. Petaloconchus domingensis Brown and Pilsbry, Proc Acad. Nat. Sci. Phila., p. 359, 1911. Petaloconchus domingensis Maury, Bull. Amer. Paleont, V, No. 29, p. 128, PI. 22, fig. 11, 1917. A single specimen was found, showing part of the coiled portion of the shell. There seems to be no doubt as to the identity. The fragment with three whorls measures in height 8 mm., diameter 6 mm., diameter of whorl about 3 mm. Locality. — 204. Horizon. — Quebradillas limestone. 140 SCIENTIFIC SURVEY OF PORTO RICO Petaloconchus? collazoensis, new species Plate XXI, Figure 16 Tube of small diameter (2.5 mm.); surface smooth except for faint growth lines; tip of spire missing in specimens, but one specimen shows a loosely, irregularly coiled cylindrical spire of five whorls; diameter of spire 10 mm.; whorls are in contact, but coiled with a 5 mm, radius, leaving large umbilical opening, the whole resembling the coiled steam pipes of a cylindrical boiler. The specimens show that the coiling habit ceases abruptly, the tubes then growing in all directions, almost tying knots. Localities. — 263, Collazo Falls. Horizon. — San Sebastian shale. Genus Pyramidella Lamarck Pyramidella portoricoensis, new species Plate XXI, Figure 17 Shell small, conic, smooth, rather low spired; seven flat whorls with sharp median spiral offset, forming, with the sharply beveled shoulder of the overlapping whorl, a broad, square sutural channel. The median offset, or angulation, continues around the body whorl; suture of earlier whorls is narrow; large circular perforation extends the entire length of the axis, and is very conspicuous as seen from the base; columella tri-plicate; posterior plication large and distinct; the other two small and obscure, of which the anterior one is situated at the extreme base of the columella; aperture oval; outer lip simple; height 4.75 mm., diameter 2 mm. This shell resembles P. (LongchcBus) forulata Guppy, but its umbilicus places it with the typical group of Pyramidella. The suture is of the type of P. semicanaliculatus Maury. Locality. — 204. Horizon. — Quebradillas limestone. Genus Modulus Gray Modulus modulus Linne variety basileus Guppy Monodonta basilea Guppy, Geol. Mag., new ser. decade 2, I, p. 434, PI. 16, fig. 2, 1874. Modulus hasilea Ball, Trans. Wagner Inst. Sci., Ill, pt. 2, p. 294, 1S92. Modulus modulus var. hasilea Dall, Trans. Wagner Inst. Sci., Ill, pt. 6, p. 1585, 1903. HUBBARD, TERTIARY MOLLUSC A FROM LARES DISTRICT 141 This shell is apparently the same as that described by Giippy from the Bowdeii. It agrees with Guppy's description, especially in its low spire, and tubercnlated, carinate whorls. On the lower portion of the final whorl, however, it apparently differs somewhat from the Bowden shell in having spirals of variable size, the primaries alter- anting with one or more fine secondary threads. Jndging from Guppy's illustration, the aperture of the Porto Kican shell is broader. The outer lip is denticulate, but the interior is not preserved, so as to show whether the Urate character exists or not. Height of shell is 14 mm., diameter 13 mm. Apparently the modern representative of this shell is M. lenticu- laris (Chemn.) Auct., which according to Tryon is synonymous with M. modulus Linne. Locality. — East side of Vieques Island (Collected by A. K. Lobeck). Horizon. — Quebradillas limestone. Genus Bittium (Leach M. S.) Gray Bittium species inclet. Plate XXI, Figure 18 Whorls six (or more), with three raised spirals crossed by sharp ribs of equal height, and about twelve to a whorl; prominent nodes produced at intersections of ribs and spirals; suture deeply Impressed; height (6 whorls) 11 mm., diameter 5 mm. A single specimen, too imperfect to identify or describe fully. Locality. — 204. Horizon. — Quebradillas limestone. Genus Cerithium Adanson Cerithium portoricoensis, new species Plate XXII, Figures 5, 6 Shell small, slender; protoconch unknown; whorls twelve or thirteen, rounded, with obscure varices of irregular occurrence on the last six whorls, and usually a prominent varix on the body whorl; first three whorls with three primary spirals crossed by about twelve narrow rounded ribs, more prominent than the spirals, and slightly nodose where they cross the latter, the whole producing a cancellate structure with rectangular pits; a fourth 142 SCIENTIFIC SURVEY OF PORTO RICO primary spiral appears on the fourth whorl; beginning with the sixth whorl, secondary, less prominent spiral threads appear, two immediately above the suture, and one between each of the four primaries. The two uppermost secondaries do not appear until the seventh whorl ; on last four whorls the ribs have increased to about twenty in number, are irregular, low, and obscure, be- ing marked chiefly by the lines of nodes where they cross the primary spirals. The ribs usually become obsolete on the anterior two-thirds of the body whorl; aperture short, slightly more than half the length of the body whorl; posterior canal small, U-shaped; anterior canal not quite covered by inner lip; outer lip slightly thickened; three average specimens have the following measurements: Height 20 mm., Maximum Diameter 6 mm. Height 17 mm., Maximum Diameter 5 mm. Height 16 mm., Maximum Diameter 5 mm. This shell has a superficial resemblance to C. coccodes Dall, and to Clava chipolana Dall. Locality. — 204. Horizon. — Quebradillas limestone (abundant). Cerithium quebradillensis, new species Plate XXI, Figures 7, S A single decollate specimen with six whorls; earliest whorls roundly angular, with shoulder and peripheral keel; last three whorls with about 12 high, rounded ribs, sub-nodose at the keel; all whorls with primary spirals and secondary spiral threads; last four whorls with very fine tertiary spiral threads; earliest whorls show two closely grouped primary spirals at the- keel, the upper one slightly stronger and forming the periphery of the whorl; a third primary occurs at (just posterior to) the suture; these primaries persist throughout, and a fourth primary spiral occurs on the base, being covered in the preceding whorls. It lies close to the suture; the secondary spirals at first form a group of three on the shoulder, another group of three below the keel, and a single one between the primary spirals of the keel. The three on the shoulder are markedly stronger than the other secondaries, a feature which persists throughout; on the third whorl (of the decollate specimen) the tertiary spiral threads appear, the first just above the sutural primary, the next ones intercalated between the secondaries in the group below the keel. The tertiaries do not appear on the shoulder until the fourth whorl; the shoulder, adjacent to the suture, is, in all whorls, marked by a rather wide band, at first smooth, but later marked by from one to six tertiary threads; all whorls are marked by fine, raised growth lines, or longitudinal threads, which produce a minute beading on crossing the spirals, and which in the earlier whorls, produce with the spirals a fine cancellate ornamenta- tion; the primary spirals of the later whorls have a secondary superimposed' thread, which is beaded by the growth lines; the base is slightly concave,. HUBBARD, TERTIARY MOLLUSGA FROM LARES DISTRICT 143 and is sculptured with spirals of four sizes, more or less alternating. The single primary, above described, is sculptured with numerous minute spiral threads; pillar smooth, with no callus, anterior notch small, short, recurved; slight suggestion of a posterior notch; aperture ovate. Height 32 mm., diameter 15 mm. The most striking feature of this shell is the sudden change in the character of ornamentation of the whorls, with the appearance of the ribs. This final ribbed sculpture resembles that of C. calli- soma Dall, from the Caloosahatchie Pliocene, but the Porto Kican shell differs from the latter in its concave base and general turritel- olid aspect. It is apparently not closely related to any species of the Antillean region. Locality. — 204. Horizon. — Quebradillas limestone. Cerithium (Campanile) collazum, new species Plate XXIII, Figures 1, 2 Shell large, conic, rapidly enlarging; whorls flattened to slightly rounded, with a median concavity in the later whorls of adult individuals; suture deeply impressed; each whorl with a flat, narrow shelf or shoulder pro- jecting beyond the preceeding whorl; surface of whorls with medium fine, nodose spirals like those of C. halense Dall, but with different arrangement. Just behind the suture are one to two very finely nodose spiral threads, one of which is frequently obscured by the overlapping whorl. Behind these is the largest spiral, ornamented with small round nodes about 2 mm. apart. The remainder of the whorl is ornamented with finer spirals, nodose, and alternating in size. The median concavity of the later whorls comes im- mediately posterior to the large noded spiral above described. The inter- spaces are slightly narrower than the spirals; base flattened, slightly convex, smooth; aperture roundly quadrate; columella stout, with single prominent fold, anterior notch short, large, recurved. A large decollate specimen with six whorls measures in height 48 mm., maximum diameter 28 mm. An interesting feature of this shell is the presence in the adult stage of an impressed spiral line near the periphery of the base, which marks the line of juncture with each succeeding whorl. This is an adult character, and is lacking in the earlier whorls. It is apparently an illustration of the inheritance of an acquired charac- ter. Another interesting feature is the change in form of the whorl from rounded in youth to flattened, and finally to a concave form in the final stages of old individuals. 144 SCIENTIFIC SURVEY OF PORTO RICO It would seem that this shell is related to some of the large, coni- cal forms from the Flint Eiver Oligocene, such as C. halense Ball, as its ornamentation is quite similar to the latter. It differs notably in the shape of its whorls, and in this latter respect, is more like the hu-ge species of Campanile from the Paris Basin Eocene. Maury has described the internal mold of a larger, but similar shell from the San Sebastian shale Campanile (Portoricia) laricum Maury, which may be a related species or a variety. Maury's species differs chiefly in its flatter and more concave whorls, as well as in its considerably greater size. Localities.— 262, Collazo Falls, 142?, 236, 749(a). Range. — San Sebastian shale to lower Lares limestone. Genus Strombus Linne Strombus proximus? Sowerby strombus i)roximas Sowerby, Quart. Jour. Geol. Soc, VI, p. 4S, PI. 9, fig. 8, 1849. Strombus pugilis Gabb (in part), Trans. Amer. Phil. Soc, XV, p. 233, 1873. Not pugilis Linne. Exclude other synonyms. Strombus proximus Guppy, Quart. Jour. Geol. Soc, XXXII, p. 521, 1876. Strombus pugilis Dall (in part), Trans. Wagner Inst. Sci., Ill, pt. 1, p. 177, 1890. Not pugilis Linne. Strombus proximus Maury, Bull. Aiuer. Paleont., V, No. 29, p. 119, PI. 20, figs, 4, 5, 1917. Only incomplete specimens were found, Avhich are identified doubt- fully with this species. Localities. — 204, Mona Island (Lobeck). Tlorizon. — Quebradilhis limestone, Ponce limestone. Strombus bifrons? Sowerby A single specimen was found among the mnt(>rial collected from the Quebradillas limestone near Bayamon by Dr. Lobeck. The in- ternal mold shows that the shell is the same shape as S. Ufrons, and measures in height (with tip of spire missing) 32 mm., diameter 19 mm. It is apparently an immature specimen, f(n- the outer \\\^ does not flare to any extent, and the anterior notch is not noticeably recurved. The ornamentation consists of spirals like those of S. bifrons, but with tlic nodes or spines rather obsolete on the final HUBBARD. TERTIARY MOLLUSCA FROM LARES DISTRICT 14& whorl, in which respect it resembles S. gatunensis Toula. S. bifrons is characteristic of the Aphera and Sconsia formations in Santo Domingo. Locality. — Near Bayamon (Lobeck), Horizon. — Qnebradillas limestone. Genus Orthaulax Gabb Orthaulax gabbi? Dall Plate XXIII, Fignre 3 A single specimen was obtained from the sea cliff at the mouth of the Guajataca Eiver in a stratum of very resistant limestone. The speci- men had been exposed to the polishing action of the waves, and the original coloring, a light mottled brown, is preserved on the body of the shell near the aperture. The surface of the body whorl and spire are marked by the numerous faint longitudinal growth lines characteristic of the genus. Viewed from above, the outline is sub- circular, with a slight triangularity as in 0. pugnax Heilprin and 0. gabhi Dall. The apical angle is about the same as that of 0. gabhi, but considerably more acute than that of 0. aguadillensis Maury, a shell apparently from the same horizon. A marked sub- angularity of the body whorl is very similar to that of 0. gabbi. Above this angulation the whorl is strongly convex; below it, the whorl is concave, with a slight convexity midway between the angu- larity and the anterior end of the shell. The angularity is strongest at the aperture, on the inner lip, but is obsolete over the final two thirds of the body whorl, A slight shoulder is developed above the angulation on the final portion of the body. Height of shell 65 mm., diameter 38 mm. This shell was examined by Dr. Dall, who states that while the specimen is too poor for positive identification, it is probably 0. gabhi, a shell characteristic of the Chipola horizon. A comparison with 0. aguadillensis Maury, a fragmental specimen, shows that the shell here described is quite a dift'erent species. Locality. — 204. Horizon. — Quebradillas limestone. 146 SCIENTIFIC SURVEY OF PORTO RICO Orthaulax portoricoensis, new species Plate XXV, Figures 1, 2, 3, 4, 5 Under this heading are included all the large Strombus-like in- ternal molds, which are among the most abundant forms to be found in the Quebradillas limestone. One of these internal molds was figured by Semmes, in his report on the San Juan District, as Stromhus, species indet. The internal molds, taken by themselves, are generically indeterminate, but resemble the molds of a large Strombus. Fortunately, a few of these internal molds were found with portions of the external mold adhering to the specimens, which proves beyond a doubt that these large shells belong to the genus Orthaulax. They attain a size much greater than any species of Orthaulax yet known. The specimens illustrated by figures 1 and 2 were examined by Dr. Dall, who states that: 1 . Figure 1 is undoubtedly an Orthaulax, possibly the same species as Figure 2, but having a more acute or pointed spire. 2. Figure 2 is also an Orthaulax, new species, almost cer- tainly the same species as an undescribed form from Santa Cruz. 3. Xeither of these specimens is the same species as the shell described in this report as 0. gahhi? Dall. A large number of specimens, in addition to the two examined by Dr. Dall, show considerable variation in size and elevation, or acute- ness of the spire. The fact that all these variations can be found in shells collected from a single fossiliferous pocket in the limestone, would indicate that they are for the most part variations of a single species, although more than one species of Orthaulax occur in the Quebradillas formation. The apical angle as measured from the external molds, varies from 78° to 90^^ (largest specimen) ; and from the internal molds, varies from 60° to 85°. The following are measurements taken from a few of the internal molds: HUBBARD, TERTIARY MOLLUSCA FROM LARES DISTRICT 147 Height Diameter Apical Angle 1 155 mm 105 mm 83° 2 115 80 85 S 75 82 4 60 75 5 . . . (Fig. 1) 40 60 6. 7 ...(Fig. 2) 80 75 8 70 9 70 0. 60 53 73 50 75 43 72 40 71 35 68 From these measurements, it will be observed that the smaller shells have a higher, more acute spire. This may be a varietal dif- ference. Figures 3 and 4 show the internal mold of one of these smaller shells in which the final whorl has begun to envelop the spire, thus indicating that all of these small shells may be immature individuals. Figure 1 shows the internal mold of the spire, and the external mold of the enveloping dome of an adult shell. The curious openwork structure replacing the shell Avail is the cast of the burrow- ing of a sponge or some other organism. This boring animal seems to have taken a special liking to the thick-shelled Orthaulax and other ponderous gastropods of the Quebradillas formation. The burrows became infiltrated with lime carbonate, which solidified with the surrounding limestone. The shell structures were then removed by solution, leaving the casts of these burrows in the form of a complex network resembling calcareovis algae. Figure 5 shows the dome of an adult shell of the low spired type, and illustrating the degree to which the shell had been riddled with globular chambers, frequently connected by long w^orm-like passages. Localities. — 204, 41, 52(a), 801, and numerous other localities not recorded. An excellent specimen was collected by Dr. A. K. Lobeck on the east coast of Vieques Island. Horizon. — Quebradillas limestone, Los Puertos limestone. Char- acteristic of the Quebradillas of the entire north coast of Porto Rico, as shown by the collections of Berkey and Semmes. 148 SCIENTIFIC SURVEY OF PORTO RICO Genus Cyprsea Linne Cyprsea spurcoides? Gabb Cyprwa spurcoides Gabb, Trans. Amer. Phil. Soc, V, p. 235, 1873. Cyprcea spurcoides Maury, Bull. Amer. Paleont., V, No. 29, p. 115, PI. 19, figs. 7, 8, 9, 1917. Apparently the same as the form described from Santo Domingo, except that the crenulations of the two lips are more nearly equal in number. Height 24 mm., maximum diameter 18 mm. Locality. — 204. Honzon. — Quebradillas limestone. Cypraea sancti-sebastiani Maury Localities.— 2Q2, 232(b), 263?, 233, 235, 237, Juana Diaz. Horizon. — San Sebastian shale, Juana Diaz shale. Genus Cassis Lamarck Cassis, species indet. Plate XXIT, Figure 9 Internal molds of a shell which may possibly be a new species. The distinguishing feature is the presence of a strong varix on the body whorl on the side opposite the aperture. Outer lip simple, ap- parently not thickened. The specimens are uniform in size and appear to l)e adult. The external sculpture, judging from impres- sions on the internal molds, consists of from eighteen to twenty broad spirals on the last whorl, with correspondingly fewer spirals on the earlier whorls. Height 30 mm., diameter 23 mm. This shell is apparently of the type of Phalium aldrichi Dall, but is not quite as broad as the latter. Localities. — 232(b), 236. Horizon. — San Sebastian shale. Genus Dolium Lamarck Subgenus Malea Valenciennes Malea camura Guppy i¥a/eaca?H/n a Guppy, Quart, .lour. Geol. Soc, XXII, p. 287, PI. 17, fig. 9, 1866. Malea ringcnsG&hh (in part), Trans. Amer. Phil. Soc, XV, p. 223, 1873. HUBBARD, TERTIARY MOLLUSGA FROM LARES DISTRICT 149 Malea camnra Guppy, Quart. Jour. Geol. Soc, XXXII, p. 525, 1876. Malea camnra Dall, Trans. Wagner Inst. Sci., Ill, pt. C, p. 1584, 1903. Malea camur a Msiury, Bull. Amer. Paleont., V, No. 29, p. 112, PI. 19, fig. 3, 1917. The specimens average about 37 mm. by 27 mm. Only internal molds were found with a few fragments of external mold. Locality. — 204; and Berkey 98. Horizon. — Quebradillas limestone (abundant). Genus Columbella Lamarck Subgenus Strombina Moerch Strombina portoricana, new species Plate XXIIT, Figure 4 Protoconch of two whorls, rather large, smooth, naticoid; post-nuclear whorls six to seven, moderately convex; suture distinct, but not channelled, and becoming obscure on last whorl; surface of whorls smooth except for faint growth striae, and by hump-like varices, or swellings, occurring as early as the third post-nuclear whorl; body whorl with spiral lines, alternating in strength, on the pillar and lower part of -whorl, but very faint over the remainder of the whorl, and more closely spaced near the suture and on the pillar. A specimen with eight whorls and base of pillar missing measures in height 19.5 mm., diameter 9 mm. The largest specimen measures in height 33 mm., diameter 17 mm. ITo complete specimens were found, and none showing the lip or aperture. This shell is apparently related to the group represented by 8. lessepsiana Brown & Pilsbry, S. prisma Pilsbry & Johnson, and S. cyphonotus Pilsbry & Johnson. Locality. — 204. Horizon. — Quebradillas limestone. Genus Phos Montfort Phos costatus Gabb Plate XXII, Figure 10 Id Phos costatus Gabb, Trans. Amer. Phil. Soc, XV, p. 212, 1873. Phos costatus Maury, Bull. Amer. Paleont., V, No. 29, p. 88, PI. 14, figs. 13, 14, 1917. The specimens correspond closely to those figured by Maury from Santo Domingo. Protoconch not preserved in any of the specimens. 150 SCIENTIFIC SURVEY OF PORTO RICO An adnlt specimen measures iii height 29 mm., diameter 14 mm. Xiimber of whorls eight. Locality. — 204. Horizon. — Qiiebradilhis limestone (abundant). Phos elegans Guppy variety portoricoensis, new variety Plate XXII, Figure 11 Shell small (height about 20 mm.), high spired; whorls rounded, with about fourteen rather sharply rounded ribs, crossed by primary, and fre- quently alternating smaller secondary spirals; small tubercules at inter- sections of spirals and ribs; occasionally two or three longitudinal threads between the ribs which do not cross the spirals; ribs irregularly spaced, especially on the body whorl, where a varix is formed at or near the aperture; ribs on the spire frequently vary in size, but do not form varices. All the specimens are fragmental, but sufficient to show the entire shell except the protoconch. This variety is probably closest to the Santo Domingan occurrence of the species. Locality. — 192. Horizon. — Quebradillas limestone. Genius Murex Linne Murex (Phyllonotus) cornurectus Guppy M. (Chicoreiis) megacerus Gabb, Trans. Amer. Phil. Soc, XV, p. 202, 1873. Not M. raegacerus Sowerby. M. cornurectus Guppy, Quart. Jour. Geol. Soc, XXXII, p. 521, PI. 28, fig. 4, 1876. M. {Phyllonotus) cornurectus Maury, Bull. Amer. Paleont., V, No. 29, p. 103, PI. 16, figs. 9, 10, 1917. An internal mold, together with a portion of the external mold, was found of a large Murex which seems to be this species. Height (of last three whorls) 80 mm., diameter (of internal mold) 45 mm. Locality. — 204. Horizon. — Quebradillas limestone. HUBBARD, TERTIARY MOLLUSCA FROM LARES DISTRICT 151 Genus Alectrion Montfort Alectrion gurabensis Maury variety portoricoensis, new variety variety varicum, new variety Plate XXII, Figures 12, 13, 14, 15 Plate XXIV, Figure 19 The Arecibo limestone contains an abundance of a small shell which is like A. gurabensis Manry, a Santo Domingan form, but which has some very evident diiferences from the latter. These shells fall into two distinct varieties, one with coarse ribs, the other with finer, more numerous ribs and a pronounced varix. The former are here called variety ■portoricoensis, the latter variety varicum. The former is by far the most abundant of the two. Both forms are smaller and have fewer whorls than the Santo Domingo type, but they are not young shells as shown by the uniformity in size among a large number of specimens. Xo specimen of either variety was found which measures more than 8 mm. in height. The following table shows the relation of the two varieties to the tvpe form from Santo Domingo, as described by Maury (Bull. Amer. Paleont., V, Xo. 29, p. 91, PL 15, fig. 21, 1911) : Locality. — 204, Horizon. — Quebradillas 1 imestone. Genus Fusus Lamarck Fusus henekeni Sowerby Plate XVIII, Figure 5 Fusus henekeri Sowerby, Quart. Jour. Geol. Soc, VI, p. 49, 1849. Fusus henekeni Gabb, Trans. Amer. Phil. Soc, XV, p. 204, 1873. Fusus henekeni Guppy, Geol. Mag., p. 439, 1874; Quart. Jour. Geol. Soc, XXXII, p. 524, PI. 28, fig. 6 (type), fig. 2 (var. haitensis) , 1876. Fusus henekeri Ball, Trans. Wagner Inst. Sci., Ill, pt. 6, p. 1584, 1903. Fusus henekeni Grabau, Smithsonian Misc. Coll., XLIV, No. 1417, p. 19, 1904. Fusus henekeni Maury, Bull. Amer. Paleont., V, No. 29, p. 78, PI. 12, fig. 1. 1917. A single specimen was found, with protoconch and body whorl missing. Only primary and secondary spirals show on the specimen. 152 SCIENTIFIC SURYEY OF PORTO RICO Table 3 Characters A. gurabensi.-> ISIaury Santo Domingo A . gurabensis var. portoricensis Porto Rico •4. gurabensis var. variciim Porto Rico Size Height 11 mm. Diameter 6 mm. Height 7 mm. Diameter 4.5 mm. Height 8 mm. Diameter 4.5 mm. Nuclear whorls 3 smooth 2-2.5 2? Post-nuclear whorls 6 5 5 Character of post - nuclear whorls Convex, angulated and shouldered ; ribs narrow, round ; wider interspaces. Convex, sub-angular and shouldered. EarUer wliorls sub- angular — the last two are rounded. Suture Wavy Wavy Slightly wavy Ribs on body whorl 11 12 Slightly oblique 13-14 Oblique Spirals on spire. Penultimate whorl 3 strong, widely spr ced 3 weaker above 3 weaker below Penultimate whorl same as in the San- to Domingo shell 3rd post-nuclear 1 weak posterior 3 strong median 2 weak anterior enultiniate 3 weak posterior 3 strong median 4 weak anterior Spirals on body 3 or 4 weak posterior 3 strong median 12 w(>ak anterior- 2-3 weak posterior 3-4 strong median 10-13 weaic anterior 3 weak posterior 3 strong median 12 weak anterior Spirals on pillar 8 6-7, arrangement — 2 strong alternate with 2 weak ? Aperture Round Sub-Round Round Outer lip 8 lirae Thin 15 lirae, altoriiatc long and short Thickened b>- strong varix Inner lip Posterior denticle Smooth, moderately thick callus. Posterior denticle absent or obscure. Smooth, moderately thick callus. Posterior denticle Anterior canal.. Well defined Broad and deep •) HUBBARD, TERTIARY MOLLUSGA FROM LARES DISTRICT 153 This is probably a small variety, otherAvise very close to the type form, but more specimens are required for a complete description. The spire of eight whorls measures in height 20 mm., maximum diameter 8.5 mm. Locality. — 204. Horizon. — Quebradillas limestone. Genus Turbinella Lamarck Turbinella chipolana Dall variety precursor, new variety Plate XXIII, Figures 6, 7 Shell small, fusiform, consisting of three and one-half nuclear whorls and three post-nuclear whorls; protoconch like T. chipolana Dall and T. regina Heilprin, in having a swollen tip, succeeded by smaller nuclear whorls. The tip consists of one and one-half volutions, is transversely ovate, and is peculiar in having a shallow median constriction. The two succeeding nuclear whorls considerably smaller in diameter, the first being gently convex, the last nearly flat. The latter is succeeded abruptly by sculptured whorls with seven rounded, not nodose ribs, and prominent spirals, of which there are four on the first two, and about twenty-three on the final whorl and pillar. The spirals are separated by wider interspaces. The whorls are marked by a prominent, narrow and wavy sub-sutural band or shoulder, sculptured with at first one and later with two spirals. The interspace immediately below the shoulder is strongly impressed. The spirals are of equal strength, except on the pillar, where faint intercalated threads appear. The entire post-nuclear surface is marked by fine raised longitudinal growth lines, strongest in the interspaces between the spirals. Height of shell 17.5 mm., maximum diameter 7 mm.; height of protoconch 3^^ mm., diameter , of tip of protoconch 2% mm. This shell is probably the ancestor of T. chipolana Dall. It is apparently adult, its small size being indicative of its primitive character. The same is true of variety arecihoense, a mutation from the San Sebastian shell occurring in the Quebradillas lime- stone. All the material available was gone over thoroughly with the purpose of trying to find some molds or fragments which would show that these shells reach a larger size, but the results were nega- tive. It seems, therefore, safe to assume that the primitive Turhi- nellas were small forms, which did not reach their maximum size 154 SCIENTIFIC SURVEY OF PORTO RICO until the Pliocene. Analogous cases may be seen in some of the genetic groups of Fusiis (Graban, A. W. Phyl. Fusns, Smithson. Miscl. Coll., XLIV, Xo. 1417, 1904). Locality. — 84. Horizon. — San Sebastian shale. Turbinella chipolana Dall variety areciboense, new variety Plate XXIII, Pignre 8 A single specimen in which the lower half of the body whorl is missing. Protoconch like that of variety precursor, but without median construction of the tip, and with the third whorl almost entirely concealed by the over- lapping of the first post-nuclear whorl, giving the appearance of a two- whorl protoconch. The tip is slightly larger (diameter 3 mm.) than that of precursor, and slightly more swollen when compared with the succeeding whorls. The whorl following the tip is nearly flat, whereas in precursor it is noticeably convex. The first post-nuclear whorl has ribs, spirals, and sub- sutural band, or shoulder, as in precursor and represents the precursor stage. On the next and following whorls, however, the ribs become nodose, and on later whorls there are more spirals than in precursor, with occasionally a fine intercalated spiral. The longitudinal growth lines are similar to those of precursor, but show a tendency to alternate in strength. The sub- sutural band, or shoulder, is prominent in the first two post-nuclear whorls and marked by a single spiral; on the third whorl it is inconspicuous, and marked by two spirals, as in the later whorls of precursor, while on the fourth and fifth (or final) whorls it has disappeared altogether. This fact is interesting in showing the tendency to lose a primitive character which has apparently been completely lost in the Chipolan shell. Height of frag- ment 20 mm., maximum diameter 11 mm. This shell is a mutation from \ariety precursor, the precursor, or simple ribbed stage being accelerated and limited to the first post- nuclear M-horl. The later whorls have nodose ribs and repro^oiif what may be termed the areciboense stage. This shell is best inter- preted as a local mutation iioi in the direcf line leading to T. chipo- lana and T. regina. Locality. — 204. Horizon. — Quebradillas limestone. 1 HUBBARD. TERTIARY M0LLU8CA FROM LARES DISTRICT 155 Genus Xancus Bolten Xancus validus Sowerby Turbinellus valid us Sow erhy. Quart. Jour. Geol. Soc, VI, p. 50, 1849. TurMnella valida Gabb, Trans. Amer. Phil. Soc, *XV, p. 218, 1873. Turbinellus validus Guppy, Geol. Mag., p. 438, 1874; Quart. Jour. Geol. Soc, XXXII, p. 523, 1876. Turbinella scolymus Tryon (in part) Man. Conch., IV, p. 70, 1882. Not the recent shell. Turbinella validus Dall, Trans. Wagner Inst. Sci., Ill, pt. 1, p. 99, 1890. Turbinella scolymus Dall (in part). Trans. Wagner Inst. Sci., Ill, pt. 1, p. 97, 1890. Xancus validus Maury, Bull. Amer. Paleont, V, No. 29, p. 83, PI, 13, fig. 5, 1917. Apparently rare. Only a few fragments were found, showing the sculpture. Locality. — 204. Horizon. — Quebradillas limestone. Xancus, new species I Plate XXIIT, Figure 9 Known only from fragments. Spire unknown; shell apparently like X. validus Sowerby and X. icilsoni Conrad, but differing chiefly in the greater number of tubercles, or spines, which are along a contracted zone below the rounded shoulder. There are four high, strong columellar folds, the anterior one being smaller than the others. The lower portion of the body whorl is sculptured with strong spirals as in X. validus. The exterior of the remainder of the shell is unknown, except for the presence of the tubercles. Height of body whorl 67 mm., maximum diameter about 50 mm. Locality.— 24:4, 254-5-6. Horizon. — Lares limestone. Genus Mitra Lamarck Mitra henekeni Sowerby Plate XXIV, Figure 1 Mitra henekeri Sowerby, Quart. Jour. Geol. Soc, VI, p. 46, PI. 9, fig. 5, 1849. Mitra henekeni Gabb, Trans. Amer. Phil. Soc, XV., p. 219, 1873. Mitra henekeni GuTppy (in part). Quart. Jour. Geol. Soc, XXXII, p. 528, 1876. Mitra henekeni Maury, Bull. Amer. Paleont., V, No. 29, p. 74, PI. 12, figs. 5, 5a, 1917. 156 SCIENTIFIC SURVEY OF PORTO RICO A single fragmental specimen of a voiing shell. Protoconcli miss- ing. Seven whorls. Height 20 mm., diameter 7 mm. Locality. — 204. Horizon. — Quebradillas limestone. Mitra symmetrica Gabb Mitra symmetrica Gabb, Trans. Amer. Phil. Soc, XV, p. 220, 1873. Mitra symmetricus Maury, Bull. Amer. Paleont., V, No. 29, p. 75, 1917. A single fragmental specimen, consisting of the last two whorls with anterior portion missing. Columellar folds not shown; whorls flattened, slightly convex; suture distinct, but not impressed; aperture elongate-ovate; surface with flattened spirals and much narrower grooves between, which are marked by growth lines. Height 25 mm., diameter 12 mm. It has been suggested that M. symmetrica may be the young of M. titan Gabb. Locality. — 280. Horizon. — Lares formation. Genus Olivella Swainson Olivella muticoides Gabb variety portoricoensis, new variety Plate XXIV, Figures 2, 3 The shells vary from high to low spired, and also in the number of columellar plications. They include the types represented by 0. muticoides and 0. muticoides variety canaliculata of Santo Domingo, and there are all gradations between. The sutural canal is extremely broad in all specimens; columellas callus very thick; three to four columellar plications, with several faint ones posterior to these; largest specimen measures in height 12 mm., diameter 4.5 mm. Localities. — 204, 671 ; Aguila Point, K 75.2 Ponce-Penuelas road (Mitchell). Horizon. — Quebradillas limestone (extremely abundant) ; and Ponce limestone. HUBBARD, TERTIARY M0LLU8CA FROM LARES DISTRICT 157 Olivella portoricoensis, new species Plate XXIV, Figiire 4 Shell small, smooth, moderately high spired; whorls about six; protoconch missing on all specimens; suture distinct, impressed below upper margin of the succeeding whorl, leaving a gap between top of whorl and surface of the preceding whorl; a raised spiral thread appears above the suture, barely showing above the upper margin of the overlapping whorl; columella with two spiral grooves; callus moderately thick; anterior notch shallow, incon- spicuous. Height of shell 12 mm., diameter 5 mm. The distinguishing feature of this shell is the spiral thread at the suture. This thread is not continued on the body whorl. Localities. — 204, 216. Horizon. — Quebradillas limestone. Genus Cancellaria Lamarck Cancellaria laevescens Guppy Plate XXIV, Figures 5, 6 Cancellaria Iwvescens Guppy, Quart. Jour. Geol. Soc, XXII, p. 289, PI. 17, fig. 12, 1866. Cancellaria Iwvescens Gabb, Trans. Amer. Phil. Soc, XV, p. 239, 1873. Cancellaria Iwvescens Maury, Bull. Amer. Paleont, V, No. 29, p. 64, PI. 10, fig. 6, 1917. Cancellaria Iwvescens Dall, Trans. Wagner Inst. Sci., Ill, pt. 1, p. 43, 1890. The specimens are smaller than the Jamaican shells, and show all variations between complete loss of cancellation on the body whorl and merely a diminution of strength of cancellation near the aper- ture. In the latter cases, the spirals continue, but the ribs are ir- reg-ular in occurrence and indistinct or absent for considerable in- tervals. In some shells the ribs on the body whorl become merely irregular crowded growth lines ; callus very thick ; columella with at least three prominent plaits; aperture not widening anteriorly as does that of the Jamaican shell. An average shell measures in height 23 mm., diameter 14 mm. The largest complete specimen measures 26 mm. by 15 mm., while a fragment indicates that the shells attain a size slightly larger than this. Locality. — 204. Horizon. — Quebradillas limestone (abundant). 158 SCIENTIFIC SURVEY OF PORTO RICO Genus Turris Bolten Turris albida Perry vaiiety haitensis Sowerby Plate XXIV, Figure 7 Plenrotoma haitensis Sowerby, Quart. Jour. Geol. Soc, VI, p. 50, 1849. Turris albida var. haitensis Maury, Bull. Amer. Paleont, V, No. 29, p. 50, PI. 8, fig. 4, 1917. A few specimens were collected Avhich agree closely with this variety. The largest specimen (spire with body whorl missing) measures in height 22 mm., diameter 8 mm. Locality. — 204. Horizon. — Quebradillas limestone. Turris albida Perry variety cf. virgo Lamarck Plenrotoma virgo Lamarck, Anim. s. Vert., VII, p. 94, 1822. Plenrotoma viryo Moore, Quart. Jour. Geol. Soc, IX, p. 130, 1853. Turris (Surcula) virgo Gabb, Trans. Amer. Phil. Soc, XV, pp. 206, L'OV, 1873. Turris alhida var. virgo Maury, Bull. Amer. Paleont. V, No. 29, p. 50, PI. 8, fig. 6, 1917. A single fragment showing portions of the last two whorls, and indicating; a shell of about 60 mm. heia'ht. An unusual feature is the absence of secondary spirals. It seems to be an adult form whicli has retained the form of the young variety virgo. Another special feature is the coarseness of the spirals, the prominent median keel being especially broad and thick, resembling the keels of Ecphora qiiadricostata. The suture is indistinct. Interspaces between s]>irals marked only by the cliaracteristic oblique growth lines. Locality. — 204. Horizon. — Quebradillas limestone. Genus Drillia Grav Drillia censors Sowerby variety portoricoensis, new variety Plate XXIV, Figures 8, J) Shell small, moderately slender; whorls about nine; sutural band as in the type, but nearly half the axial width of the whorl; sculpture of from HUBBARD. TERTIARY MOLLUSC A FROM LARES DISTRICT 159 four to five prominent spirals, and about 18 sharp ribs which do not cross the sub-sutural band, the latter being smooth, and concave on either side of the spiral ridge which divides the band. The concavity below this spiral ridge is over twice the width of that above the ridge, and much more deeply concave. Height 15 mm., diameter 5 mm. This shell is close to D. consors, but has some distinctive differ- ences. One of the specimens shows the protoconch fairly well. It is rather large, smooth, naticoid, and apparently consists of two whorls. The first post-nnclear whorl is sculptured with ribs only, the spirals being too fine to observe (if present at all) and the sub- sutural band not appearing until the second sculptured whorl. Locality.— '204: ; Berkey 98. Horizon. — Quebradillas limestone. Drillia grabaui, new species Plate XXIV, Figure 10 Shell small, slender; decollate specimen (5 whorls) measures in height 14 mm., diameter 4 mm.; protoconch of two smooth, rather large, globular whorls; post-nuclear whorls with sharp, widely spaced ribs and fine, flattened spiral threads, not crossing the ribs and slightly arcuate between the ribs; sub-sutural band like that of D. consors, but narrower and wavy, or fluted, and marked with faint spiral threads and arcuate growth lines indicating a fairly deep, semi-circular posterior sinus. On the penultimate whorl, there are six strong spirals, with three additional ones just below the band, which are fine, crowded, and about the same strength as those on the band. On the body whorl there are twenty-two spirals, and the ribs extend well toward the anterior portion of the shell before dying out. The ribs, spirals, and band appear together immediately following the protoconch stage. This shell has a slight resemblance to D. maonrisparum Maury, of the Santo Domingan Oligocene, but differs in its slightly larger size, more slender form, different protoconch, and absence of any varix. It probably belongs to the D. consors group. Locality. — 204. Horizon. — Quebradillas limestone. DriUia portoricoensis, new species Plate XXIV, Fig-ures 11, 12 Shell with ten whorls, including protoconch of two smooth, globular volutions; post-nuclear whorls sculptured with low, straight longitudinal ribs and faint spirals, visible only between the ribs, except on the anterior half 160 SCIENTIFIC SURYEY OF PORTO RICO of the body whorl, where they are much stronger, and slightly nodose where they cross the ribs; sub-sutural band of the D. consors type, with dividing spiral ridge prominent in the earlier whorls, but becoming low and incon- spicuous in the adult; band slightly wrinkled where it laps over the ribs of the preceding whorl; lower half of band slightly concave, and smooth except for faint, microscopic spiral lines, and growth lines indicating a broad, sub-angular posterior sinus; adult shells with a strong, thick varix at the outer lip. Height of shell 16 mm., diameter (excluding varix) 5.5 mm. This shell is easily recognizable from its conspicuous varix, and its almost obsolete sculpture. Locality. — 204. Horizon. — Quebradillas limestone. Drillia semmesi, new species Plate XXIV, Figure 13 Shell small, broader and lower spired than is usual in members of this genus; whorls seven, including a large protoconch of two smooth, rounded whorls, the first small, the second considerably larger; first post-nuclear whorl sculptured with about thirteen sharp ribs extending from suture to suture, and crossed by three raised spiral ridges, producing a cancellate sculpture; all post-nuclear whorls sharply shouldered, and the ribs, on crossing the shoulder, are strongly arched backward, a feature most noticeable in the later whorls; shoulders of the last four whorls are marked by obscure spirals; later whorls retain the three primary spirals, and on the fourth whorl, smaller intercalated secondary spirals appear. The uppermost primary spiral marks the shoulder angulation. The body whorl has about twelve prominent (or "primary") spirals, with some of the interspaces marked by one or two faint (or "secondary") spirals. All post-nuclear whorls show longitudinal growth lines, arcuate on the shoulder, indicating a wide U-shaped posterior notch. Aperture and anterior portion of shell unknown. Length of most complete specimen, with lower portion of body whorl missing, 8 mm., diameter 4 mm. This shell is named in honor of Dr. D. R. Semmes, in recognition of his work on the San Juan District of Porto Rico. Locality. — 204. Horizon. — Quebradillas limestone. Genus Cythara Schumacher Cythara cf. elongata Gabb Mangilia elongata Gabb, Trans. Amer. Phil. Soc, XV, p. 211, 1873; Jonr. Acad. Nat. Sci. Phila., VIII, p. 351, PI. 46, fig. 34, 1881. HUBBARD, TERTIARY MOLLUSCA FROM LARES DISTRICT 161 Cythara elongata Dall, Trans. Wagner Inst. Sci., Ill, pt. 6, p. 1583, 1903. Cythara elongata Maury, Bull. Amer. Paleont., V, No. 29, p. 59, PI. 9, fig. 12, 1917. A single poorly preserved specimen, which seems to be this species or one very closely related. Height 7.5 mm., diameter 3 mm. Locality. — 204. ZZ^ori^o n.—Quebradillas limestone. Genus Terebra Adanson Terebra quebradillensis, new species Plate XXIV, Figure 14 Shell small, slender; protoconch of two smooth globular whorls, slightly larger than the succeeding post-nuclear whorl; twelve post-nuclear whorls, rounded as in T. herlinerce Maury; sub-sutural band nearly one-third the axial width of the whorl, marked off by deep, narrow furrow, and faintly striated by one or two spiral threads as in T. wolfgangi Toula; about eighteen arcuate ribs on each whorl, which cross the furrow and sub-sutural band without interruption or change of direction; remainder of whorl below sub-sutural band marked by about nine spiral threads (sixteen on last whorl) which vary in size. A prominent spiral is situated next to the furrow. The rest are finer, and crowded together in groups of two or three. In the earlier whorls they are more uniform in size and distribution; and fewer in number; columella short and non-plicate. The' largest specimen measures in height 18 mm., diameter 4 mm. This shell resembles T. herlinerce Maury in its convex whorls and the character of the spiral threads. It differs in its smaller size and wider sub-sutural band. It resembles T. ivolfgangi Toula in its bisected or trisected sub-sutural band. Localities. — 204, and near Ponce (Mitchell). Horizon. — Quebradillas limestone, Ponce limestone. Genus Conus Linne Conus catenatus Sowerby Plate XXIV, Figure 15 Conus catenatus Sowerby, Quart. Jour. Geol. Soc, VI, p. 45, PI. 9, fig. 2, 1S49. Conus interstinctus Guppy, Quart. Jour. Geol. Soc, XXII, p. 288, PI. 16, fig. 3, 1866. Conus catenatus Gabb, Trans. Amer. Phil. Soc, XV, p. 230, 1873. 162 SCIENTIFIC SURVEY OF PORTO RICO Conus catenatus Guppy, Quart. Jour. Geol. Soc, XXXII, p. 527, 1876. Gonus catenatus Maury, Bull. Amer. Paleont., V, No. 29, p. 38, PI. 5, fig. 4; PI. 6, figs. 1, 2, 1917. The specimens vary from concave to only slightly concave spire, from high to moderately low spire, and there is considerable range in size. Upper whorls carinated ; last five whorls rounded on shoulder and broadly channelled above, with faint arcuate growth lines crossing the channels; posterior third of body whorl smooth; anterior two- thirds marked by spiral threads which alternate with much finer, obscure threads. There are many variations, some of the specimens resembling C. cercadensis Maury, other resembling C. kitieredgel Maury, but all evidently variations of C. catenatus. A typical shell measures in height 31 nmi., diameter 17 mm. Locality. — 204. Also found by Berkey in the vicinity of Que- bradillas. Horizon. — Quebradillas limestone. Conus cf. marginatus Sowerby Plate XXIV, Figure 16 Conus marginatus Sowerby, Quart. Jour. Geol. Soc, VI, p. 44, 1849. Conus marginatus Gabb, Trans. Amer. Phil. Soc, XV, p. 230, 1873. Conus marginatus Guppy, Quart. Jour. Geol. Soc, XXXII, p. 528, PI. 29, fig. 5, 1876. Contis (Chelyconus) maroinatus Cossmann, Jour, de Conch., LXI, pp. 44-46, PI. 3, figs. 14, 15, 1913. Conus marginatus Maury, Bull. Amer. Paleont., V, No. 29, p. 46, PI. 7, fig. 11, 1917. A small conus which has features in common with C. marginatus and may be the young of the latter. Whorls eight, including two nuclear; spire moderately high, scarcely concave; whorls carinate except last whorl, which is rounded at the shoulder; whorls of the spire flat above, while the final whorl is slightly channelled above; posterior half of body smooth; anterior half with twelve smooth, flat, spiral bands separated by deep grooves marked by raised longi- tudinal threads, which arc limited to the grooves and do not cross the spiral bands. Height 7.5 mm., diameter 4 mm. Locality. — 204. Horizon. — Quebradillas limestone. HUBBARD. TERTIARY MOLLUSCA FROM LARES DISTRICT 163 Genus Bullaria Rafinesque Bullaria paupercula Sowerby Bulla pauper tula Sowerby, Quart. Jour. Geol. See, VI, p. 52, 1849. Bulla paupercula Gabb, Trans. Amer. Phil. Soc, XV, p. 246, 1873. Bxaia paupercula Gwv^y , Geol. Mag., p. 437, 1874; Quart. Jour. Geol. Soc, XXXII, p. 518, 1876. Bulla paupervulaDdM, Trans. Wagner Inst. Sci., Ill, pt. 1, p. 18, 1890. Bulla striata JidiW, Trans. Wagner Inst. Sci., Ill, pt. 6, p. 1583, 1903. Bullaria pauper culaMoMiy, Bull. Amer. Paleont., V, No. 29, p. 18, PI. 3, fig. 8, 1917. The specimens are internal molds, but their general outline seems sufficient to identify them with this species. Fragments of external mold show faint spiral grooves on the anterior part of the shell. Spire depressed. Height 15 mm., diameter 10 mm. Localities.-— 204:, 6Q4:, 235 ?. Horizon. — Quebradillas limestone. Ponce limestone. Bullaria portoricoensis, new species Plate XXIV, Figure 17 Shell cylindrical, length nearly twice the diameter; spire sunken, imper- forate; aperture longer than shell, extending above the truncate apex; surface with fine, slightly wavy impressed spiral lines, covering all of the shell except the summit, and crossed by slightly curved growth lines. An average specimen measures in height 20 mm., diameter 11.5 mm. The most striking characteristic of this shell is its cylindrical form. The young shell is more globose in form, resembling one of the elongate forms of B. granosa. Locality. — 204. Horizon. — Quebradillas limestone. Bullaria granosa Sowerby Plate XXIV, Figure 18 Bulla granosa Sowerhy, Quart. Jour. Geol. Soc, VI, p. 51, PI. 10, fig. 10, 1849. Bulla granosa Gabb, Trans. Amer. Phil. Soc, XV, p. 246, 1873. Bulla granosa GuY)i)y, Geol. Mag., p. 437, 1874; Quart. Jour. Geol. Soc, XXXII, p. 518, 1876. Bullaria granosa Msiury, Bull. Amer. Paleont., V, No. 29, p. 20, PI. 3, fig. 10, 1917. 164 SCIENTIFIC SURVEY OF PORTO RICO The specimens are slightly more globular than the Santo Domingo shells. There is a great range in size, but the specimens fall into two groups, one of a large size averaging 20 mm. in height and 16.5 mm. in diameter, and the other a small size averaging 14 mm. in height and 11 mm. in diameter. The small shells have a large range in size and proportions, some of the measurements being: Height Diameter 17 mm. 15 mm. 11 mm. 8 mm. 16.5 mm. 12.5 mm. 11.5 mm. 10 mm. The surface sculpture of the shells agrees closely with the shells from the Santo Domingo Oligocene. Localities.— ^04:, 209?, 216, 52(a), 191, 225, 678. San Juan district (Semmes). Culebrinas Point? (Mitchell). Horizons. — Lares limestone to Quebradillas limestone. Ponce lime- stone? Extremely abundant in the Quebradillas limestone. Plate X Fig. 1. Atrina rahelli n. s. ; nat. size; external mold. Fig. 2. Pecten (Pecten) laresense n. s. ; x 1.6; left valve. Fig. 3, Pecten (Pecten) laresense n. s. ; x 2 ; right valve. Fig. 4. Pecten (Euvola) reliquus Brown and Pilsbry, var. porto- ricoensis n. var. ; x 2.3. Fig. 5. Pecten (Nodipecten) nodosus Linne; x 1.8. Fig. 6. Phacoides (Lucinisca) calhounensis Dall; x2.6; immature shell ; gutta-percha cast. Fig. 7. Metis trinitaria Dall ; x 0.75 ; internal mold, left valve. SC'IENTIFIC SURVEV (iF I'dKTK UUO AND THi; VlIKilN ISLANDS Volume 111, I'late X Fig. 1. Fig. 2. Fig. 3. Fig. 4. Fig. 5. Fig. 6. Fig. 7. Fig. 8. Plate XI Pecten (Chlamys) collazoensis n. s. ; x 3.5. Pecten (Chlamys) portoricoensis n. s. ; x 2.25 ; right valve. Pecten (Chlamys) portoricoensis n. s. ; x 2 ; left valve. Pecten portoricoensis var. reticulatis; x 2.25, Pecten portoricoensis var. grandis; x 3.3 ; fragment show- ing prominent ribs, Pecten portoricoensis var. grandis; x 3,3 ; fragment of another shell showing low ribs. Tellina sp. aif, T. (Angulus) atossa Dall; xl.8; gutta- percha cast. Solen (Plectosolen) collazoensis n, s. ; x 14. SciKNTiFic Survey of Pouto Kuo and the Vikuin Islands Volume III, 1'late XI Plate XII Fig. 1. Peden (Chlamys) grdbaui n. s. ; x 2.25. Fig. 2. Pecten grabaui var. aguadensls; x 2.2, Fig. 3. Pecten grabaui var, hatoensis; x 2, Fig, 4. Pecten grabaui var. guayabensis ; x 2 ; ears defective. Fig. 5, Pecten (Chlamys) hodgii n, s. ; x 2,8. Fig, 6. Pecten (Plagiocteniuni) sebastiensis n, s. ; x 1.4. SriE.NTiFic Survey of Porto Rico and the Virgin Islands Volume III, Plate XH Plate XIII Fig. 1. Pecten (Aequipecten) lobecki n. s. ; x 2.5. Figs. 2, 3. Pecten (Plagiodenium) cercadica Maury; x 1.9. Fig. 4. Ostrea sellceformis Conrad, variety portoricoensis n. var. ; nat. size; lower valve, adult shell. Figs. 5, 6. Ostrea sellceformis var. portoricoensis; nat. size ; upper valves, exterior and interior; young. Fig. 7. Chione woodwardi Guppy; nat. size; gutta percha cast. SciK.NTiFic Survey of 1'orto Rico and the Viu«in Islands Volume III, Plate XIII Plate XIV Fig. 1. Pecten (Plagioctenium) borinquenense n. s, ; x 2.5 ; right valve. Fig. 2. Pecten borinquenense; x 2.5 ; loft valve. Fig. 3. Amusium mauryi n. s. ; x 3.4. Fig. 4. Barbatia cf. bonaczyi Gabb; x2.9; gutta-peroha cast. Fig. 5. Area dariensis Brown and Pilsbry ; x 2.2. Fig. 6. Area (Scapharca) cf. donacia Dall; x3.7; right valve, gutta-percha cast. Fig. 7. Area cf. donacia; x 4; left valve, gutta-percha cast. Scientific Strvey (if 1'outo Uico and the Vihci.n Islands Volume HI, Plate XIV Plate XV Fig. 1. Ostrea antiguensis Brown; xO.9; lower valve. Fig. 2. Ostrea haitensis Sowerby var. ? ; x 1.3 ; interior, young shell. Fig. 3. Ostrea haitensis var. ? ; x 1.3 ; exterior, young. Figs. 4, 5. Glycimeris collazoensis n. s. ; x 1.5. Fig. 6. Glycimeris portoricoensis n. s. ; x 1.5 ; gutta-percha cast. SCIENTIKIC SUR\ EV OF I'ORTO RiCO AND THE VlHGIN ISLANDS VOLUMK III, I'l-ATK XV Plate XVI Fig. 1. Ostrea cahohasensis Pilsbry and Brown, variety porto- ricana n. var. ; x 0.6 ; valves together; specimen collected by A. K. Lobeck. Fig. 2. Pitaria (Hyphantosoma) carbasea Guppy; x2.2; gutta- percha cast. Figs. 3, 4. Tellina (Scissida) grabaui n. s. ; x 2.5 ; gutta-percha casts. Scientific Survey of Pouto Rico and thk Vikgin Islands Volume III, Plate XVI Plate XVII Fig. 1. Ostrea cahohasensis Pilsbry and Bro\vii, variety porto- ricana n, var. ; xO.6; lower valve, specimen collected by Lobeck. Figs. 2, 3. Area sp. indet. ; x 2.2. Fig. 4. Codakia (Jagonia) magnoliana Dall, variety borinquenense n. var. ; x 2.5 ; gutta-percha cast. Fig. 5. Chama portoricana n. s. ; x 0.8 ; internal mold with por- tions of external mold of the lamellae at the margin. Fig. 6. Leda peltella Dall x 2.4; gutta-percha cast. SCIENTIFU' SVRVEY OK roKTO TJ K'O ANO THK VlliCIN ISLANDS VOLUME III. PLATE XVII Plate XVIII Fig. 1, Lucina collazoensis n. s. ; x 0.7 ; left Talve, int. mold. Fig. 2. The same specimen, — dorsal view; x 1.1. Fig. 3. Lucina cf. chrysostoma (Meusch.) Philippi; nat. size; internal mold. Fig. 4. Phacoides (Miltha) sp. indet. ; x 1.4 ; gutta-percha cast. Fig. 5. Phacoides (Miltha) sp. indet; x2.6; gutta-percha cast. Fig. 6. Phacoides (Pseudomiltha) laresensis n. s. ; x 1.4 ; internal mold, with portions of the shell preserved near the beak, showing the sculpture. Scientific Survey of Pouto Rico and the ViiuiiN I.si.amis VOMIMK HI, I'l.ATK. XVIII Fig. Fig. 1. 2. Figs. 3, ■ Figs. 5, Fig. 7. Fig. Fig. 8. 9. Fig. 10. Fig. 11. Fig. 12. Plate XIX Cardiutn muricoides n. s, ; x2.6; gutta-percha cast. Cardium (^Trachycardlum) cinderellw Maury, variety alternatum n. var. ; x 2.2 ; int. mold, 4, Cardium cinderellce var. alternatum; x2.6; gutta- percha casts showing variations in the sculpture. 6. Cardium (Trigonocardia) samhaicum Maury, variety portoricoensis n. var. ; x 2 ; gutta-percha casts. Cardium n. s. ? (aff. C. samhaicum Maury) ; x 2.5 ; gutta- percha cast. Maury ; x 2.5 ; gutta-percha cast. Cardium {Trigonocardia) haitense var. cercadicum Cardium haitense variety areciboense n. var. ; x 2.5 ; gutta-percha cast. dementia dariena Conrad; x 1.1 int. mold, specimen from Robles, P. R. Clementia dariena Conrad; x 0.95 ; int. mold, specimen from Collazo, P. P., somewhat distorted by pressure. Clementia dariena Conrad; x 0.95 ; specimen from the Gatun formation, Panama, collected bv Prof. J. F. Kemp. SCIENTU'IC SUU\EV OF I'OUTO UiCO AND THE VlIUilN ISLANDS Vui.UMK III, I'LATE XIX Plate XX Fig, 1. Cytherea (Cytherea) herkeyi n. s. ; x 0.75 ; int. mold, speci- men collected by Prof. Berkey. Pig. 2. Same specimen, dorsal view ; x 0.7. Pig. 3. Same specimen; x 0.75 ; gutta-percha cast from fragment of the external mold. Pig. 4. Tellina strophoidea n. s. ; x 2.5 ; right valve, gutta-percha cast. Pig. 5. Tellina strophoidea; x 1.3; left valve, gutta-percha cast. Pig. 6. Tellina portoricoensis n. s. ; x 1.5 ; right valve, gutta-percha cast. Scientific Survev of Pouto Rico and the Virgin Islands Volume III, I'l ate XX Plate XXI Fig. 1. Calliostoma portoricoensis n. s. ; x 2.5 ; gutta-percha cast. Fig. 2. Turbo fetkii n. s. ; x2.6; gutta-percha cast. Fig. 3. Liotia (Arene) coronata Dall, variety portoricoensis u. var. ; X 4 ; gutta-percha cast. Fig. 4. Neritina (chipolana var. ?) collazoensis n. s. ; x 2. Fig. 5. Hipponyx portoricoensis n. s. ; x 2 ; gutta-percha cast. Fig. 6. Hipponyx portoricoensis; x 2 ; gutta-percha cast, showing variation with strongly recurved apex. Fig. 7. Crucibulum auricula Gmelin variety portoricoensis n. var. ; x 1.3 ; gutta-percha cast, specimen collected by Prof. Berkey. Fig. 8. Same specimen; x3.3; gutta-percha cast of the internal . cup. Fig. 9. Crucibulum auricula? var.; x2.3; gutta-percha cast. Fig. 10. Crucibulum (Dispotcea) collazum n. s. ; x 2 ; interior, showing cup. Fig. 11. Natica (Ampullina?) collazoensis n. s. ; xO.75; decollate shell with rounded whorls. Fig. 12. Natica collazoensis; x 0.7 ; decollate shell with angulated whorls (int. mold). Figs. 13, 14. Turritella tornata Guppy variety portoricoensis n. var. ; x 1.1 ; int. molds. Fig. 15. Turritella portoricoensis n. s. ; x 2.7 ; gutta-percha cast. Fig. 16. Petaloconchu^? collazoensis n. s. ; x 1.5. Fig. 17. Pyramidella portoricoensis n. s. ; x 3.6 ; gutta-percha cast. Fig. 18. Bittium sp. indet. ; x 2 ; gutta-percha cast. S< iKNl'iFic SUK\ KV <)i' I'm; 1(1 Kn iiik Vi i,i:i .\ Islands VtiiiMi: II 1. I'l.ATK XXI Fig. 1. Fig. 2. Fig. 3. Fig. 4. Fig. 5. Plate XXII Epitonium {Girsotrema) collazoensh n. s. ; x 2.2. Same specimen; x 1.5. Turritella mitchelli n. s. ; x 1.4. Turritella herkeyi n. s. ; x 0.85 ; gutta-percha cast. Ceritliium 'portoricoensis n. s. ; x 3.7 ; giitta-percha east, showing protoconch. Fig. 6. Ceritliium portoricoensis; x2.3; gutta-percha cast, show- ing varix on body whorL Cerithium quebradillensis n. s. ; x 2.6; plastolin cast. Same specimen; x2.2; showing aperture and pillar. Cassis sp, indet. ; x 0.8 ; int. mold. Phos costatus Gabb; x 0.8 ; gutta-percha cast. Phos elegans Guppy variety portoricoensis n. var. ; x 1.4 ; gutta-percha cast. Fig. 12. Alectrion gurabensis Maury variety portoricoensis n. var. ; X 2.2 ; gutta-percha cast. Fig. 13. Alectrion gurabensis var. poi'toricoensis ; x 2.2 ; gutta- percha cast, sho^dng aperture. Figs. 14, 15. Alectrion gurabensis variety varicum n. var.; x 2.2 ; gutta-percha casts. Fig. 7. Fig. 8. Fig. 9. Fig. 10. Fig. 11. Scientific Survey of I'outo Rico and the ViiuiiN Islands VOl.U.MK III, I'l-AIK XXll Plate XXIII Figs, 1, 2. Cerithium (Campanile) collazum n. s. ; x 1.1. Fig. 3. Orthaulax gahhif DaW; X 0.9. Fig. 4. Strombina portoricana n. s. ; x 2.Y ; gutta-percha cast. Fig. 5. Fusus henekeni Sowerby ; x2.3; gutta-percha cast. Fig. 6. Turhinella chipolana Dall variety precursor n. var. ; x 2.5. Fig. 7. Same specimen; x 3.7; aperture view. Fig. 8. Turhinella chipolana variety arecihoense n. var.; x2.6; gutta-percha cast. Fig. 9. Xancus n. s. ? ; nat. size; int. mold of portion of body whorl. SciKNTiFir Survey ov 1'outo Uico ani> the Viucin Islands VOLUMK III, I'l.ATK XXIII Plate XXIV Fig. 1. Mitra henekeni Sowerby ; x 2 ; gutta-percha cast. Fig, 2. Olivella muticoides Gabb variety portoricoensis n. var. ; X 1.75 ; gutta-percha cast, showing a low spired shell. Fig. 3. Olivella muticoides var. portoricoensis; x 1.75 ; gutta- percha cast, showing a high spired shell. Fig. 4. Olivella portoricoensis n. s. ; x 3.1; gutta-percha cast. Fig. 5. Cancellaria Icevescens Guppy; xO.8; gutta-percha cast showing shell with smooth body whorl. Fig. 6. Cancellaria Icevescens; x 0.8 ; gutta-percha cast, showing reappearance of cancellate ornamentation on the final portion of the body whorl. Fig. 7. Turris alhida Perry variety haitensis Sowerby; xl.4; gutta-percha cast. Fig. 8. DrilUa censors Sowerby variety portoricoensis n. var. ; X 3.5 ; gutta-percha cast. Fig. 9. Drillia consors var. portoricoensis; x 2.3 ; gutta-percha cast. Fig. 10. Drillia grabaui n. s. ; x2.3; gutta-percha cast. Fig. 11. Drillia portoricoensis n. s. ; x 2.3; gutta-percha cast show- ing varix. Fig. 12. Drillia portoricoensis; x 2.3 ; gutta-percha cast. Fig. 13. Drillia semmesi n. s. ; x 2.25 ; gutta-percha cast. Fig. 14. Terehra quebradillensis n. s. ; xl8; gutta-percha cast. Fig. 15. Conus catenatus Sowerby; xl.5; gutta-percha cast of specimen collected by Prof. Berkej. Fig. 16. Conus cf. viarginatus Sowerby; x3.6; gutta-percha cast. Fig. 17. Bullaria portoricoensis n. s. ; x2.2; int. mold. Fig. 18. Bullaria granosa Sowerby; x2.25; gutta-percha cast. Fig. 1!). Alectiion gurahensis Maury variety portoricoensis n. var. ; X 4 ; gutta-percha cast, showing the aperture and pillar SCIENTIFir SURVKV OF POISTO K I CO AM' THi: VlKCI.N ISLANDS Vol. L. Ml. Ill, I'l.ATK XXIV Plate XXV Fig. 1. Orthaulax portorlcoensis n. s. ; x 0.85 ; internal and ex- ternal mold of the upper half of the shell showing most of the shell wall replaced by casts of a burrows made by a boring sponge or other organism. Fig. 2. Orthaulax portorlcoensis; x 0.85 ; internal mold of a large, low spired shell. Fig. 3. Orthaulax portorlcoensis ; x 0.85 ; internal mold of a small, high spired shell, probably an immature individual. Fig. 4. Reverse side of the same specimen shown in Fig. 3. Fig. 5. Orthaulax portorlcoensis ; x 0.85 ; case of the upper portion of a mature shell, made up entirely of the burrow cast= illustrated in Fig. 1. SCIEXTIKU- SURVKVOI- I'olM" K I CO AND THE Vl 1!.: I N ISI.ANI.S Vol.IIMK 111. IM.ATK \XV NEW YORK ACADEMY OF SCIENCES SCIENTIFIC SURVEY OF Porto Rico and the Virgin Islands VOLUME III— Part 3 Fossil Corals of Porto Rico, with Descriptions also of a Few Recent Species — H. N. Coryell and Violet Ohlsen NEW YORK Published by the Academy 1929 FOSSIL CORALS OF PORTO RICO With Descriptions also of a Few Recent Species By H. N. Coryell and Violet Ohlsen CONTENTS .(Those subtitles marked by an (*) were prepared jointly by H. N. Coryell and Violet Ohlsen and the remaining by the senior author.) Page Introduction 169 Purpose and resume 169 Collections studied 170 Previous work on West Indian coral fauna 170 Acknowledgments 171 Correlations based on the coral fauna 171 Tabulation of collecting localities 175 Collecting localities of the Reeds collection 176 Collecting locaHties of the Hubbard collection 181 ♦Glossary 184 Systematic description of genera and species 185 Madreporaria Imperforata 185 *Seriatoporidae Milne Edwards and Haime 185 Pocillopora Lamarck 185 Pocillopora portoricensis Corj^ell, n. sp 186 *Stylophora Schweigger 186 *Stylophora afflnis Duncan 186 *Stylophora goethalsi Vaughan 187 *Stylophora granulata Duncan 187 *Stylophora macdonaldi Vaughan 188 *Stylophora panamensis Vaughan 188 *Stylophora ponderosa Vaughan 188 *Stylophora portobellensis Vaughan 189 *Astrocoeniidae Koby 189 *Astrocoenia Milne Edwards and Haime 189 *Astrocoenia decaturensis Vaughan 189 *Astrocoenia guantanamensis Vaughan 190 *Astrocoenia meinzeri Vaughan 190 *Astrocoenia ornata Milne Edwards and Haime 191 *Astrocoenia portoricensis Vaughan 191 *Orbicellidae Vaughan 191 Antiguastrea Vaughan 112 Antiguastrea cellulosa (Duncan) ; 193 Antiguastrea cellulosa curvata (Duncan) 193 *Orbicella Dana 194 *Orbicella annularis (Elhs and Solander) 195 *Orbicella cavernosa (Linnaeus) 199 Plate XXVI XXVI XXVI XXVI XXVI XXVI XXVI XXVI XXVI XXVII XXVII XXVII XXVII XXVIII XXVII XXVIII XXIX 1C)S sviESTiric SI h'\ i:y of ro/no mco Page Plate *Orbicella costata (Duiu-an) 196 XXIX *Orhicella limbata (Duncan) 197 XXIX Orbicella tampaensis Vaughan 197 XXX *Faviidae Gregory 198 CaUitnophyllia Blainville 198 CalajnophylUa dendroidea CoryeW, n. sp 198 XXX Calamophyllia portoricensis Coryell, n. sp 199 XXX, XXXI Favites Link 200 Fanites expansa Coryell, n. sp 200 XXXII Fairites irregularis Coryell, n. sp 200 XXXII Goniastrea Milne Edwards and Haime 201 Goniastrea crassa Coryell n. sp 201 XXXIII Goniastrea pectinata (Ehrenberg) 202 XXXIII Hydnophora Fischer de Waldheim 202 Hydnophora hubbardi Coryell, n. sp 203 XXXIII Lamellastraea Duncan 203 Lamellastraea crassa Coryell, n. sp 203 XXXIII Latomeandra d'Orbigny 204 Latomeandra lata Coryell, n. sp 204 XXXIII Leptoria Milne Edwards and Haime 205 Leptoria areolata hispida (\'errill) 205 XXXIV Leptoria phrygia (Ellis and Solander) 205 XXXIV Maeandra Oken 206 Maeandrn nntiguensis Vaughan 206 XXXIV Maeandra labyrinthiformis (Linnaeus) 207 XXXIV *Manicina Ehrenberg 208 "Manicina unlloiighhiensis Vaughan 208 XXXV Metastraea Milne Edwards and Haime 208 XXXV Mefnstraen planulala Coryell, n. sp 209 XXXV *Agaricidae Verrill 209 *Agaricia Lamarck 209 Agaricia agiii.si:\. /ossn. co/.m/.n or roirro h'ico i69 Page Plate Acropora palmata (Lamarck) 219 XL Acropora panamensis Vaughan 220 XL Astreopora Blainville 223 Astreopora nntiguensis Vaughan 221 XL *Poritidae Dana 221 *Goniopora Blainville 221 *Gonwpora canalis Vaughan 222 XL *Goniopora cascadensis Vaughan 222 XL *Goniopora clevei Vaughan 222 XL *Goniopora decaturensis Vaughan 223 XL *Goniopora decaturen,ns silicensis Vaughan 223 XL *Goniopora h Hit Vaughan 224 XLI *Goniopora imperatoris Vaughan 224 XLI *Goniopora jacobiana Vaughan 225 XLI *Goniopora panamensis Vaughan 225 XLI *Goniopora portoricensis Vaughan 225 XLI *Goniopora regularis (Duncan) 226 XLII *Porites Link 226 *Porites anguillensis Vaughan 227 XLII *Porites astreoides Lamarck 227 XLII *Porites baracoaensis Vaughan 228 XLII *Porites douvillei Vaughan 228 XLII Porites (Synaraea) macdonaldi Vaughan 229 XLIII *Porites panamensis Vaughan 229 XLIII, XLIV Porites porites (Pallas) 230 XLIV * Porites toulai Vaughan 232 BibUography 232 IXTRODUCTIOX Purpose and Eesume The essential purpose of this paper is to include a specific and generic description of the corals collected by Doctors Chester A. Reeds and Bela Hubbard. Sixty-seven species and 24 genera are described. Thir- teen of the species are new. Thirty-eight of the 67 species were from the San Sebastian, 13 from the Lares, 19 from the Cibao, 3 from the Los Puertos, 2 from the Quebradillas, 39 from the Ponce and 1-t from the Pleistocene and Eecent. Every old species was identified by the aid of the original descriptions and figures, and their names have been corrected under the Laws of Xomenclature. The correlation of the Porto Rican coral fauna with the stratigraphic distribution of other similar faunas in the Caribbean Region is given in Table No. 1, pages 172-3. In Table II, page 175, are shown the probable 1-Q 8CIE\TIF/C SURVEY OF PORTO RICO equivalents of the formations of Porto Rico in other islands of the West Indies and in the subtropical and tropical portions of North America. Collections Studied In 1915 Dr. Chester A. Eeeds and Mr. P. B. Hill visited Porto Eico for the purpose of making one of the most elaborate scientific collections of fossils that had ever been assembled from that island. Their work was to forni the basis of a publication on the fossil fauna. Some of the groups of forms collected by them have already been studied by dif- ferent students^ and the results have been published either as separate numbers of the Scientific Survey of Porto Rico or have been incorporated within other reports of this survey. The coral specimens were shipped to Washington, D. C, to Dr. T. Wayland Vaughan. He was recognized as an authority on Tertiary corals and was selected as the proper person to undertake the work, but Dr. Yaughan's other duties left no time for the identification and description of the specimens, many of which were similar to those he had already described. So after a few years they were returned to the headquarters of the New York Academy of Sciences and later forwarded to Columbia University, where they have been stored since 1927. Dr. Bela Hubbard's geological studies on the Lares area of Porto Eico were published in 1923 after he had presented a report on the results of his field observation before the New York Academy of Sciences in 1917. The corals of his collections were forwarded from Porto Rico directly to Dr. T. Wayland Yaughan, Washington, D. C. They were set aside there with Dr. Eeeds' specimens and later forwarded to New York and stored in Columbia University. Nothing more was done with these two collections until the spring of 1928. At that time the work which has resulted in the present paper was initiated and the specimens gathered by both parties were studied. Previous Work on West Indian Coral Fauna The wriiers who have made incidental mention of the fossil corals of the West Indies are indeed very numerous. Those who have obtained a few specimens from the Islands and inserted a description of a new species or variety in a stratigraphic discussion of the Caribbean Region are less numerous, but yet make up a considerable list. Detailed refer- ence to these students is given in the specific and generic descriptions. Tbo zoological .'nid jjalcontological students who liavo made a s.ys- tematic classification and study of the coral fauna of tlio lands around C0RYELL-0HL8EN, FOSSIL CORALS OF PORTO RICO 171 the Caribbean Sea are few. One of the earliest of these was Duncan, an English paleontologist, who obtained several collections of corals from the West Indies. He began his publications as early as 1884 in the Zoological Journal of the Linnean Society of London and followed these with a series of publications in the Quaterly Journal of the Geological Society of London, which appeared in volumes XIX to XXVIII. A. E. Verrill was an enthusiastic worker in unravelling the life history of reef-making corals and in reorganizing the classification of the Hexa- coralla. He published many genera and species from the Porto Rican region in the Transactions of the Connecticut Academy of Arts and Sci- ence for 1901. The work of Dr. T. Wayland Vaughan in Bulletin 103 of the U. S. National Museum for 1919 probably ranks as the best organized sys- tematic paper on fossil corals of the Caribbean Province among the several papers that he has published. It is the one found most useful in the work of identifying the Reeds and Hubbard collections. Acknowledgments The writers wish to take this opportunity to express their gratitude especially to Doctors Chester A. Reeds and Bela Hubbard, who collected the Porto Rican specimens, to Miss Marjorie Elton for her splendid assistance, to Miss Amy Hepburn for making available the numerous references and to Professor Howard A. Meyerhoff for many useful sug- gestions. As the junior author was able to collaborate only during the study of a portion of the Reeds collection, those genera, species and pages of dis- cussion that were prepared jointly are designated by an asterisk (*) preceding the subtitles, both in the index and the text. CORRELATIONS BASED ON THE CORAL FAUNA The Tertiary and later formations in Porto Rico are arranged in a series of irregular "off-lapping" beds upon a median eastward-westward axis of older rock. The youngest beds are along the north and south shores with the older members outcropping in irregularly parallel bands farther inland, leaving the older sediments and igneous masses exposed in the interior. The detailed classification of the various divisions of the Tertiary are still under discussion as to their proper position in the geological time scale. Some of the determinations are based upon faunal evidences 112 SCJJ::\T11'JC SI h'VIJ) OF I'ORTO RICO primarily, while others lay claim to their position upon the interpretation of tlie structural features. The work (Ui the coral collections here presented should not l)e ex- pected to add much that is new to the stratigraphy. It should aid, however, in tlie correlation of the horizons determined in Porto Eico with those from which other coral faunas have been collected. One of the available works to form a basis for a correlation is that of Dr. T. Wayland Vaughan in Bulletin 103 of the U. S. National Museum. The following table gives the range of the Tertiary and later corals of the Reeds and Hubbard collections in the Porto Pico formations and their occurrence in other formations as determined by Vaughan. TABLE I Species from Porto Rico Porto Rican Horizons Acropora craxsa, n. sp Acropora palmata Acropora panamensis Agariria auaricites crassa Ayariria irreyularis, n. sp Ai/aricia sinuata, n. sp AniiyuaMrea cellulosa Antiynastrea cellulosa curvala . . . . Astreopora antiyuensis Aslroroenia Heraturfnaia Asiroroimia ynantanamensis Aslrocoenia meinzeri Astrocoenia ornata Astrororiiia porlnrtrensis Caiamiijihi/llia ilrniiroidca, n. sp . . Calnmoiilii/tlia purtoricensi.i, n. sp Cyathomorplia antiyuensis Cyathomorpha tenuis Diploastrra rrassolamellata Favitrs expari.ia, u. sp Favites irreyularis, n. sp Ooniastrea crassa, n. sp Goniastrea pectinata Goniopora canalis Goninpnra rasradensis Goniopora rlnii Goniopora decaturensis Goniopora decaturensis silicensis. . Vaughan, Bull. 103,U. S.N. M. 01 c C 0) u o .2 OS S«8 COh'yi:}J.-()HLs/:\. fossil cou als or roirro inco 173 TAHLE I (Continued) Species from Porto Rico Porto Rican Horizons Goniopora hilli Goniopora imperatoris Goniopora jacobiana Goniopora panamensis Goniopora portoricensis Goniopora regularis Hydnophora hubbardi Lamellastraea crassa, n. sp. . . . Latomeandra lata, n. sp Leptoria areolata hispida Leptoria phrygia Maeandra antiguensis Maeandra labyrinthiformis. . . . Manicina willoughbiensis Metastraea planulata, n. sp. . . Orbicella annularis Orbicella cavernosa Orbicella costata Orbicella limbata Orbicella tampaensis Pironastraea anguillensis Pironastraea antiguensis Pocillopora portoricensis, n. sp Porites anguillensis Porites astreoides Porites baracoaensis Porites douvillei Porites (Synaraea) macdonaldi Porites panamensis Porites porites Porites toviai Stylophora affinis Stylophora goethalsi Stylophora granulata Stylophora macdonaldi Stylophora panamensis Stylophora ponderosa Stylophora portobellensis Siderastrea conferta Vaughan, Bull. 103, U. S. N. M. O a V o « S»8 Of the 67 species identified here, 43 occur in the Oligocene and Mio- cene beds as tabulated by Vaughan. The Antigua is represented by 23 of the 42 species, 8 of which extend into the Anguilla or Emperador beds. The Antiguan corals were col- 174 SCIENTIFIC SURVEY OF PORTO RICO lected from the base of the formation. Of the 33 species, 19 occur in the San Sebastian Shale of Porto Rico as designated by Doctor Chester A. Reeds * and Professor Howard A. Meyerhoff.** The San Sebastian thus appears equivalent to the base of the Antigua formation. Twenty-one of the 42 species occur in the Lares Limestone. Of these, 7 occur in the Antigua only, 4 in the Antigua and xA.nguilla, or Antigua and Emperador and 7 in the Anguilla or Emperador only. The Lares Limestone on this basis appears transitional, but on comparing the occurrence of the Lares species with the San Sebastian, it is found that 17 of the 21 from the Lares Limestone were collected also from the San Sebastian. The Lares Limestone then appears closely related to the later formation. Only two of the Lares species were found in the Cibao above. Six of the 42 species occur in the Cibao as limited by Meyerhoff.* Of these^ 1 occurs in the Antigua only, in the Antigua and Anguilla or Antigua and Emperador, and 2 in the Anguilla or Emperador only. The Cibao appears to be related to the formations of Anguilla age rather than to older beds, but an impression based on so few species cannot be as conclusive as one could wish. From the collections of Reeds and Hubbard, 39 species were identified from the Ponce formation as determined by Doctor Reeds in the field. Of these 39 species, 21 have been identified from the San Sebastian Shale, 17 from the Lares Limestone, 4 from the Cibao Limestone, 2 from Los Puertos and 2 from the Quebradillas Formation. The dis- tribution indicates that the collection was made from the beds on the soutli side of Puerto Rico without any particular field recognition as to the zonal divisions or lithologic phases more readily determined in the fiolil (III the north side of the island. The Ponce Formation, as here deterniiiird irom the coral collection, includes beds equivalent to the San Scliasiiaii. Lares, Cibao, Los Puertos and Quebradillas of the north- ern section. TIh^ lower part of the Ponce Formation, as designated in llir III' III hy Reeds, would include the Guanica, Juana Diaz and Collazo bods as given hy Vaughan.* This would leave only the upper limestone deposits to lie included in the limited Ponce. With detailed field work added tn the faiinal studies still other limitations should be possible. * Reeds, C. A., Field Tabulation r.ist, impnblisliod. •* Mcyerlioff, II. A., personal communication. * Meyerliciir, II. A., personal communication. * Vjiuyhaii. '1'. \V,iyl:ui o O. a fi bC P s < O o3 J2 W Cibao 3 QJ O Lares Guanica 'O (S O Antigii San Sebastian Juana Diaz TABULATION OF THE COLLECTING LOCALITIES The localities from which the corals were collected are grouped into two lists. The first one gives the stations visited by Dr. Chester A. Reeds and Mr. P. B. Hill during the months of June and July, 1915. ♦Vaughan, T. W., BuU. Geol. Soc. of Amer., XXXV, 1924, opposite p. 720. ** Vaughan, T. W., Bull. 103, U. S. Nat. Mus., 1919. j-p, i<.vii:\'riif x( mi:) of j'oinu Riro The number 440 is the accession number allotted to the collection in the American Museum of Natural History. The other number or num- bers designate the field locality. The description of each station is preceded bv the age of the formation from which the collection was made as determined by the field interpretation and the study of the fossil specimens. The second list groups in consecutive order, the stations visited by Dr. Bela Hubbard during his study of the Lares district. Many of the specimens in his collection are excellently preserved. He was fortunate in securing representatives of a number of genera not commonly present in a collection of Porto Eican fossils. COLLKCTIXG LOCALITIKS OF THE KkHDS COLLECTION 440-3 Lu.s racrlus — Arecibu Lime-'iionc. Near south edge of Agua- dilla, about 150 feet above the sea on an old road lead- ing eastward up the hill. 440-11 Los Puertos — Areciho Limesione. Collection troin rock wall ill railroad cut just south of railroad station, Aguadilla. 440-14 Los Piierlos — Areciho Lintcsfonc. Collection from south end of cut just south of I'ailroad station. Aguadilla. 44i»-n Jjos I'licrlos — Areciho Liineslonc. Collection from the field 1/4 mile oast of the south edge of Aguadilla. al)out .'UM) feet al)Ove the sea. 440-18 Recent. Collection from the beach simtli of Aguadilla. 41(»-"-^o Sun Schiisliiin. ColU'ction from slialy limestone south side of I'oad near Km. post -V-), on road from San Sebastian to Lares. 4 10-4 mile north of Rabell's ranch house along the road leading northeast of San Sebastian about 10 Km. 440-116 San Sebastian. Collection from just below the last falls on the Collazo Eiver. 440-117 Lares. Collection from the weathered marl beds along the roadside near Km. post 2, Hm. post 3, in an excavation for the roadway of road No. 2, leading from Aguadilla to Eincon. 440-119 Lares. Collection from weathered marly limestone in em- bankment on the roadside near Km. post 9, Hm. post 9, on road No. 2, leading from Aguadilla to Eincon. 440-123 Lares. Collection from weathered limestone on north, bank of road near bridge at Km. post 7, Hm. post 5, on the government highway leading from Aguadilla to Eincon. 440-126, Quehradillas. Collection from the cut at west end of rail- -129,-130, road bridge over Cuajataca Eiver near Qucbradillas. -131,-133, -134,-135. 440-136, Quebradillas. Collection from along the railroad tracks be- -137. tween the wagon road crossing and the railroad bridge over the river near Quebradillas. 440-142 Quebradillas. Collection from the cut along the roadside near Km. post 40, on road from Quebradillas to Camuy. 440-170 Juatia Diaz. Collection from the "fifth ledge" of fossiliferous sillily limestone in Jacaguas Eiver 1»ed, 1-2 Km. north- west of Juana Diaz. CORYELL-OHLSEN, FOiiSIL CORALS OF PORTO RICO 179 4-10-242 Juana Diaz? Collection from the road embankment 25 feet above the river on the west bank of Coamo Eiver below Coanio Eeservoir. 440-286, Ponce. Collection from embankment of rock in situ on the -291,-292, north side of the road near Km. post 1, Hm. post 8-9, -293,-294, on the highway leading from Ponce to Penuelas. -296,-297. 440-298, Ponce. Collection from a cliff bank on the north side of the -299,-300. road near Km. post 2, Hm. post 8-9, on the highway from Ponce to Penuelas about 5 Km. west of Ponce. 440-307 Police. Collection from the south wall of road near a prospec- tive' bridge over a stream near Km. post 5, Hm. post 2, on the highway from Ponce to Penuelas. 440-320, Ponce. Collection from a reef in a northwardly-facing bluff -321,-322, along the roadside near Km. post 25, Hm. post 2, on -323. the road from Penuelas to Guayamilla. 440-325, Ponce. Collection from a coral and algal reef in a high bluff -326,-327, along roadside near Km. post 25, Hm. post 1-3, on the -328,-329, road from Ponce to Penuelas and Guayamilla. -330,-331. 440-339, Po7ice. -Collection from a small bat cave in a great wall of -340,-341, coral and foraminiferal reef limestone on the east shore -342,-344, of Guanica Harbor. -346. 440-354, Ponce. Collection from the cliff just south of the lighthouse -355,-359, pier, Guanica Harbor. -360,-361, -362,-364, -365,-366, -367. 440-368, Potice. Collection from the face of cliff, 300 feet south of -369,-370, the small bat cave, east shore Guanica Harbor. -373,-375, -376,-378, -380. 440-381, Ponce. Collection from the foraminiferal beds designated -383,-384, above (440-339, etc.) and with 30-40 feet of limestone 180 SCIEyriFW SURVEY OF PORTO RICO -385,-386, overlying the reef at 340-389, east shore, Guanira TTar- -387,-389, ' bor. -390. 44()--l()l, Cretaceous. Collection from the island 1/4 mile southeast of -402,-403, Parguera, Porto Pico. The Island forms the east shore -405,-407, of the Parguera Harbor. -408,-409, -410,-412, -413,-415, -418. 440- ill, Lares. Collection from the basal portion of the limestone .44.9^.443^ overlying the tuff near Km. post 64, Hm. post 9, on the -444. road from Arecibo to Ponce. 440-445, Lares. Collection from the bluff on the west side of the road -446,-449, near Km. post 66, Hm. post 6-T, leading from Arecibo -450. to Ponce. 440-451, Lares. Collection from near Km. post 66, Hm. post 7-8, on -452,-453, the road from Arecibo to Ponce. •454,-455, -456,458. 440-460, Cibao. Collection from the talus slope and wash neai- Km. -4(il. post 69, II in. post 2.5, on the road from Art'cihu to Ponce. 440-462 Cibao. Collection from a high bluff on the west side of the road, near Km. post 69, Hm. post 3, leading from Are- cibo to Ponce. 440-465, Cibao. Collection from a high bluff along the roadside near -466,-467, Km. 69, Hm. i)osts 6-7, on the higliway from Arecibo to -468. Ponce. l|n. isCi Los Piicrlos. Collection from blocks along roadway taken from boulders in the talus slope near Km. post 2. on the road from Manati to Ciales. 440-490 CihiKK Collection from limestone 17; situ in tlie east blnlT along tlie roadsi(k^ neai' Km. post I. II m. post 6, on the highway leading from Manati to Ciales. I IO-507 Lares. Collection from a Mulf 60 feet high and 200 feet long on the east side of the roadway near Km. post !». on tile road from Manati to Ciales. CORYELL-()TrL>iEX. FOSSll. rO/M/.N (>/■' I'ORTO RICO 181 4-4:0-542 Qiiebradillas. Collection i'rom a railruad cut in a blull on the west bank of Eiver Loiza, 14 n^ile northwest of Central Canovanas, near Loiza. Collecting Localities of the Hubbard Collection 800 801 802 803 804 805 806 807 808 811 812 ), Zone C, Km. 32.8, Lares Road. L Zone C, Lares Eoad. 814 815 ^^^ I Zone C, Collazo-Lares Road. 817 j 818 ] 819 I Zone C, Lares Road. 822 J 824 ] 826 y Zone D, Lares. 827 J 828 ] 829 I I Zone C, Collazo-Lares Road. 830 833 834 ] 835 i Zone C or D, West of San Sebastian. 836 J 839 ] 2one C, Lares Road. 840 5 841 I Zone D, Lares. 182 SCIEXTIFIC SURVEY OF PORTO RICO 846 847 849 850 852 853 I Zone C, Collazo-Lares Eoad. 854 I 855 I 856 I 857 I 859 I 860 J 863 I Zone D, Lares. 864 ] 865 I 866 I 867 868 86!) 870 874 877 878 879 880 881 882 i Zone C, Collazo Lares Road. 886 I 887 888 890 891 893 894 896 897 898 I 899 I 900 I 901 I 909 J ('()ini:TJ,-()iiLsi:\. fosxil corals of J'orto rivo 183 912 913 911 916 917 918 922 924 925 933 934 935 936 937 942 943 945 948 955 956 957 958 959 960 964 968 974 975 980 982 984 986 987 989 1 [ Zone D, Lares. Zone C, East of Lares. \- Lares District. Zone C or D. East of Lares. > Zone C, Lares Iioad. L;ires District. Zone G, South of Caniuy. i Lares District. I Zo]ie C, Collazo near Lares lioad. I Zone C\ Collazo-Lares EoacL J 18-t .s(7/;\77/7t' .sr/i'i7;i' of I'oirro inco 9itO I Zone If, Kincoii Point. 991 993 995 996 998 999 y Lares District 1000 1001 1004 ^1006 I ' 1089 J H 2 \ Lares District H 3 J S 5 ] S 7 \ P.IL 82 a-d ) .. ,, > /.one I P.IJ. 98 c-g \ ' *OLOSSAPY :\r(.na Island (Collected by X. L. Rritton). or D, Lares Peef. Calyx-ifc . . . Cociiciichyiuu .The cuii-shiipcd upinT end of a corallite. . rdiims iiiU'rcdi-allilc tissue. CoUine \ cah-areous ridiie separatinj; series of ealicular centers as in Manichifi. Columella ("oiallilc ( "oralluiii A solid or porous rod of calcareous tissue that occupies the center of a corallite. .\n individual of a eoralluui. An eidire sl ; calices shallow, circular, 0.8 to 1.0 mm. in diameter, surrounded by a raised margin and from 0.8 to 1.0 mm. apart; six prominent septa reach to the large, dense and styliform columella; coenenchyma papil- lose, the granules often in rows. Locality.— U0-6-i:,-o00, -321,-339, -384:,-4:U, Eeeds Collection, Ameri- can Museum of Natural History. Occurrence. — San Sebastian Shale Ponce Formation and Lares Lime- stone. * Stylophora goethalsi Vauglian, 1919 Plate XXVI, Figures 3, 3a Stylophora goethalsi Vaughan, U. S. Nat. Mus. Bull. 103, 1919, p. 338, PL LXXV, figs. 2-3. Description. — Corallum composed of compressed branches with mar- ginal protuberances; calices shallow, 0.5 to 0.75 mm. in diameter and from 0.5 to 1.5 mm. apart; six well developed septa extend to the promi- nent columella, short secondary septa occur; coenenchymal surface, granulose. Cotype.—l^o. 334767, U. S. National Museum. Locality. — 440-460, Reeds Collection, American Museum of Xatural History. Occurrence. — Cibao of the Arecibo group. * Stylophora granulata Duncan, 1864 Plate XXVI, Figure 4 Stylophora granulata Duncan, Geol. Soc. London Quart. .Torn-.. XXT. 1864, p. 10, ri. II, fig. 3; Duncan, Geol. Soc. London Quart. .Tdur.. XXIV, 1867, p. 25; Duncan, Geol. Soc. London Quart. Jour., XXIX, 1873, p. 551; Vaughan, U. S. Nat. Mus. Bull. 103, 1919, p. 340. Description.— Covsillum composed of nearly cylindrical branches; calices deep, 0.75 to 1.35 mm. in diameter, irregularly spaced and sur- rounded by a raised margin formed by the septa and costae; six prin- cipal septa reach to the columella, secondary septa are present; the coenenchyma is conspicuously granulose and in places costate. Locality.— 4:4:0-327, Eeeds Collection, American Museum of X^atural History. Occurrence. — Ponce Formation. 188 SCIEyTfFIC SURVEY OF PORTO RICO * Stylophora macdonaldi Vauglian, 1919 Plate XXVI. Figures 5, 5a Htylophont inacdonaldi Vaughan. U. S. Nat. Mus. Bull. 103, 1919, p. 339, PI. LXXV, tigs. 5-7a. Description. — Corallum consisting of slender branches, compressed wherever bifurcating; calices shallow, 1.0 mm. in diameter and from 0.5 to 1.5 mm. ajDart; six well developed septa extend to the prominent columella with their outer edges raised at the margin of the calyx; coenenchymous tissue granulated. * Cotypes.—^o. 324769, 324770, U. S. National Museum. Locality. — 440-117,-126,-321,-331, Eeeds Collection, American Mu- seum of Xatural History, and 901, Hubbard Collection, Columbia Uni- versity, Xew York City. Occurrence. — Lares Limestone, Quebradillas Formation and Ponce Formation. * Stylophora panamensis Vauglian, 1919 Plate XXVI, Figure 6 Htylophorn pfnuininisift Vaughan, U. S. Nat. Mus. Bull. 103, 1919, p. 335, PI. I.XXV, figs. 1. la. Description. — Corallum consists of irregularly shaped plates; calices shallow, 0.5 to 0.75 mm. in diameter, from 0.5 to 1.0 mm. apart and margins slightly elevated; six primary septa join the compressed col- umella; coenenchyma inconspicuously granulose. Type.—lSio. 324768, U. S. National Museum. Locality. — 440-20,-39, Eeeds Collection, American Museum of Natural History. Occurrence. — San Sebastian Shale. * Stylophora ponderosa Vaughan, 1900 Plate .\.\\I, Figure 7 Hti/lni)hliaii, 1919 Plate XWIl, Figure 2 Astrorornia meinzeri Vauglian, U. S. Nat. Mus. I'.ull. 103, 1919, p. 349, PI. LXXIX, figs. 3, 3a. Vc.scripUon. — Corallum coni|t()stMl of lirauchcs elliptical in cross-sec- tion; calices from 2.5 to 3.0 imu. in .lianictcr and from 1.25 to 1.5 mm. deep; the walls are from 0.5 to 1.5 nun. iWn-k, with the upper surface ridged or grooved longitudinally: tiuM-f arc sixteen septa, with eight reaching Ihe styliform coluindla whci'c Ihc ends of the septa thicken and fuse around the columellar mass; dissepiments are present. Type.— 'No. 324791, U. S. National Museum. Locality. — 440-346, Reeds Collection, Anicriean :\rnscum of Natural History. Ornirrcncc. — Ponce Formal ion. VORYEJ.L-OHLSi:\. FOSSIL COl^'ALS Ul' I'OliTO h'lCO I'Jl * Astrocoenia ornata .AFiliie Edwards and Haime, 1857 Plate XXVII, Figure 3 Astrocoenia ornata Milne Edwards and Haime. Hist. Nat. Corall., II, 1857, p. 357. Description. — Corallum forms a rounded or hemispherical mass; calices poh'gonal, measuring from 3.0 to 2.5 mm. in diameter; walls thick and very granulose; the principal septa are thickest near the wall and Join with the large columella at the center of the calyx. Localitij. — 1:40-51, Eeeds Collection, American Museum of Natural •History. Occurrence. — San Sebastian Shale. * Astrocoenia portoricensis Vaughan, 1919 Plate XXVII, Figures 4, 4a Astrocoenia portoricensis Vaughan, U. S. Nat. Mus. Bull. 103, 1919, p. 350, PI. LXXVI, figs. 4, 4a ; PL LXXVIII figs. 1, la. Astrocoenia ornata Yaughau, Geol. Soc. Londou Quart. Jour., LVII, 1901, p. 497: (not) Milne Edwards and Haime, Hist. Nat. Corall., II, 1857, p. 257. Description. — Corallum composed of palmate branches; calices shal- low, polygonal, from 1.0 to 1.5 mm. in diameter; walls from 0.3 to 0.5 mm. thick ; there are sixteen septa, eight of which reach the compressed styliform columella; endothecal dissepiments present; interseptal spaces about as wide as the thickness of a septum; septal costae present but inconspicuous. Type. — X^o. 324785, U. S. National Museum. i^oca^i^i/.— 440-20,-38,-39,-40,-46,-59,-321,-325,-331,-354,-355,-383,-454, Reeds Collection, American Museum of X^atural History. Occurrence.— ^2in Sebastian Shale, Lares Limestone and Ponce For- mation. * FAMILY. — ORBICELLIDAE VAUCxHAN Genus Antiguastrea Vaughan, 1919 Antiguastrea Vaughan, U. S. Nat. Mus. Bull. 103, 1919, p. 401. Heterastraea Reis (not Hcterastraea Tomes) preoccupied. Bayer. Geognost. Landesuntersuch. geognost. .Tahrg, II. 1889. pp. 150-152. Genotype. — Antiguastrea ccllii losa ( Duncan ) . 292 8vii:\'i iFic siRvin' of roirro hico Astraca ccUulosa Duncan. Genl. Soo. Tvondon Quart. Journ., XIX. 1863. pp. 417, 418, PI. XIII. tij,'. 10. Antiguastrea ccllulo.sa Vauj^han. U. S. Nat. Mu.s. Bull. lO.'i. 1919. p. 402. PI. XCVIII, figs. 3-4a ; PI. XCIX, figs. l-3a ; PL C figs. l-4a ; PI. CI, figs. 2-2a. ( For a more complete synonymy see the description of tlie siiecies. ) Description. — The coral lum forms dome-shaped, tuberose or expla- nate massive heads of variable sizes, ranging from a few inches to five or more indies in diameter; corallites are shallow, rounded or sub- polygonal and vary from 2.0 to 10.75 mm. in diameter; intercalicular tissue is porous; calicular walls are thick; the costae are subequal and as numerous as the larger septa, and usually not present over the entire exothecal tissue; the columella is lamellar and usually conspicuous; the septa are arranged in four cycles, rarely in five ; dissepiments are present. Bangs. — Oligocene. Remarks. — This genus is distinguished from OrhiceUa by its lamellar columella and less well developed costae. Antiguastrea cellulosa (Duncan), 1863 Plate X.W Til. Figure 1 Astraea cellulosa Duncan, Geol. Soe. London Quart. Jour.. XIX, 1863. pp. 417, 418, PI. XIII, fig. 10. Isfistrncd turhiiKtta Duncan. Gool. Soc. London Quart. .Tour.. XIX. 1863, p. 423. PI. XI \', ligs. la-c; Duchassaing and Miclielotti, Sup. Mem. Corall. Antilles, 1866. p. 89 (of reprint) ; Duncan, Geol. Soe. London Quart. Jour., XXIV. ISC.T, ]i. 2."» ; I >uc]iassaing. Rev. Zoopli. et Spong. Antilles, 1870. p. Ml. IJfUustrucu cellulosa Ducha.ssaing and Miclielotti, Sup. Mem. Corall. Antilles 1866, p. 86 (of reprint) ; Duncan. (Jeol. Soe. London Quart. Jour., XXIV, lNf,7, p. 21: Ducliassaiiig. Kcv. Zoopli. et Spong. Antilles. 1870, p. 30. Orbicclla ccllulosn \aii;,'lian. (Jcol. Soe. London Quiirt. Jour.. LII. 1!K)2. p. 497; Vaughan, Cirnegie Inst. Wiisliington Yearbook. No. 13. 1915. ji. 360. Antif/uastrea vclluhisn N'auglian. I'. S. \at. Mus. P.ull. lo:*.. 1919. p. 402. PI. X<'\ I IT. figs. 3-4a : PI. XCIX, figs. l-3a ; PI. C, figs. l-4a ; PI. CI. figs. 2-2a. Description. — Corallum massive heads of variable shapes as tuberose, hemisi)lu'rieal and subplanate; the corallites are rounded or subpoly- gonal. usually aiitiguous, leaving small areas of interstitial tissue; the ealieiiJar depressions are sliallow and varv in diameter from 2 to 9 mm., averaging about 4.5 mm.; se])ta are numerous, arrangecl in four cycles in the average-sized calices; the columella is small, usually distinct and composed of lamellate tissue; the walls of the calices are well developed coryi:ll-(Hii.si:\. fossil coif \ls or roirro rico 193 and thick; eostae are as luuueruus as tla- se|jUi and usually not well de- veloped. Asexual reproduction takes place by submarginal or intercalcinal gemmation. Type. — Coll. Geol. Soc. London; "Conglomerate" of Antigua. Locality. — 440-14,-34,-40,-41,-92,-242,-339,-454, Keeds Collection, American Museum of Natural History, and 826, 863, 900, 984, and P. R. 82 (a) and (b). Hu])bard Collection, Coluinl)ia T^niversity, New York City. Occurrence. — Los Puertos Limestone, Lares Limestone, San Sebas- tian Shale, Juana Diaz ? and Ponce Formation. Antiguastrea cellulosa curvata (Duncan) Plate XXVII, Figure 5 Astraca ccUulvsa curnita Duncan, Geol. Soc. London Quart. Journ., XIX, 1863, p. 418. Antiguastrea ceJlidosd curnita Vaughan. U. S. Nat. Mus. Bull. 10.3, 1919, p. 408, PI. XCVIII, figs. 4. 4a. Description. — Corallum irregular and massive; the corallites are cir- cular or compressed into oval forms as seen in cross-section and average 4.5 mm. in diameter; costae delicate and unequal; calicular walls thick; septa in four cycles, the primary are large and conspicuously denticu- late; the columella consists of porous structure; dissepiments occur within the calices ; exotheeal tissue is abundant in the interspaces. Asexual reproduction is by intercalicular or marginal gemmation. Type.— Coll. Geol. Soc. London; "chert-formation" of Antigua. Locality. — 440-39,-46, Reeds Collection, American Museum of Natural History. Occurrence. — San Sebastian Shale. * Orbicella Dana, 1846 Orbicclla Dana. U. S. Expl. Exped. Zooph., 1846. p. 205 ; Vaughan. Geol. Reichs Mus. Leiden Samnd.. ser. 2, II, 19(11. p. 21 ; VerriU, Trans. ,Conn. Acad. Arts, and Sci., XI, 1!>02, p. 93 ; Vaughan, Carnegie Inst. Washington Pub. 213, 1918, p. 85. Phymcornia Milne Edwards and Ilainie. ("oiiipte Rendu de TAcad. des Sci.. XXVII. 1848. p. 4(!9. Genotype. — Mudrciiora (iiuiiiJaria Ellis and Snlander. (For a more complete synonymy see the description of the species.) Description.— The following is the original description of the genus : "Cells nearly circular, more or less prominent, not sub-dividing by 1(^4 SVIEXTIFIC .SLRVm OF PORTO RICO growth, or rarely so; stars (calices) with distinct limits formed by the coalescence laterally of the lamellae, and therefore cells appear tubular and separated by interstices." Range. — Jurassic to Recent. * Orbicella annularis (Ellis and Solander), 1786 Plate XXVIII, Figure 2 Madreporn anniiUiris EUis and Solander, Nat. Hist. Zoopli., 1TS6. p. 169, PI. LIII, tigs. 1, 2. Modrepora faveolata Ellis aud Solander, Nat. Hist. Zooph., 1786, p. 166, PI. LIII, figs. 5, 6; Gmelin, Linn. Syst. Nat., ed. 13, 1790, p. 3709. Mudrcponi acropora (imelin, Linn. Syst. Nat., ed. 13, 1790, p. 3707; Esper, I'flauzenth., Fortsetz., I, 1797, p. 21, PI. XXXYIII. Astrea annularis Lamarck, Hist. Nat. Anim. sans Vert., II, 1816, p. 259; Lam- ouroux, Exp. Meth. Genres des Polyp., 1821, p. 58, pi. LIII, figs. 1, 2. Explanaria annularis Ebrenberg, Corall. Roth. Meer., 1834, p. 84 (of reprint). Astraea {Orhicclla) annularis Dana, U. S. Expl. Exped. Zoophytes, p. 214, PI. X, fig. 0. Hcliastraea annularis Milne Edwards and Haime, Hist. Nat. Corall.. II, 1857, p. 473; Duchassaing and Michelotti, Mem. Corall. Antilles. 1801, p. 70 (of reprint) ; Duchassaing and Michelotti, Snp. Mem. (\)rall. Antilles, 1806, p. 84 (of reprint). Heliastraca acropora Duchassaing and Michelotti, Mem. Corall. Antilles. 1861, p. 70 (of reprint) ; Duchassaing and Michelotti, Sup. Mem. Corall. Antilles, 1866, p. 84 (of reprint). Hcliastraea lamarcki Duchassaing and Michelotti, Mem. Corall. Antilles. 1801, p. 76 (of reprint) ; Sup. Mem. Corall. Antilles, 1800, p. 84 (of reprint). Cyphastraca costata Duncan (part), Geol. Soc. London Quart. Journ., XIX, 1863, pp. 441, 443; Duchassaing and Michelotti, Sup, Mem. Corall. An- tilles, 1866, p. 85 (of reprint) ; Duncan, Geol. Soc. London Quart. .Tour.. XXIV. 1868, p. 24; Gregory, Geol. Soc. London Quart. Jour., LI, 1895. p. 274. Astraea harbadensis Duncan, Geol. Soc. Londtni Quart. Jour., XIX. 1863, PI). 421, 444, PI. XV, figs. 6a, Oh. Orhicclla annularis Verrill. Mus. Comp. Zool. Lull.. I. 1864, No. 3. p. 4S : Ver- rill. Loston Soc. Nal. Hist. Proc. X. lS(i5, p. 323; Verrill. Trans. Conn. A.'.id. .\ils :iiid Sci.. XI. 1901. ii. 91. I'l. W. Im. 1; Vaughan. lUol. Soc. Washington Proc, X^', 1902, p. 50; Duerden. Nat. Acad. Sci. Mem.. VIII. ]9n;;. p. .504, Pis. VIII-X, figs. 64-73; Vaughan, Washington Acad. Sci. j'liir., \', 1915, p. o'.Ki; \:nighan. Carnegie Inst. Washington Year Book No. 14, 1916, p. 227. Hcliastraea harhadensis Ducha.ssaing and Michelotti. Sup. Mem. Corall. An- tilles, 1866, p. 85 (of reprint » : Dinuiin. Geol. Soc. London Quart, .lour.. XXIV, 1868, p. 24. Echinopora franski Gregory, Geol. Soc. Lomlnn (juaii. .loiir.. I.l. 1S!I5. ],. •2'!4. coh'yi:J-L-<)iii.sj:\. fo^sul c.); Vaughan, U. S. Nat. Mus. Bull. 103. p. 416. Genot.vpe. — Atitrca rctiforiiiis Lamarck. Milne Edwards and Ilaime. Compte Rendu de I'Acad. des 8ci., XXVII, 1848, p. 495. Description. — Corallum forms massive heads with a domed or undulat- ing upper surface ; the calices are irregularly polygonal, separated by simple compact walls and reproduce by fission : tlie septa are well de- veloped with pali occurring before the inner end of all except the last cycle ; the columella is spongy. Range. — Cretaceous to Eecent. Goniastrea crassa Coryell, new species Plate XXXIIT. Figure 1 Dcscrl[)ti(in. — The corallum forms heads 13 cm. or more in diameter: the calices are deep, irregularly polygonal, separated by thickened simi)le crested walls and vary from 4 to 10 mm. in their longer diameter and 3 to 5 nun. in the shorter diameters, measured from crest to crest of the walls; the septa are well developed, consisting of alternating long and short plates, the longer expanding at their iiuier ends to form paliform structures; the ends df only the last cycle of septa are free; fwm thirty-four to forty septa are ])resent in the larger calices and from twenty-four to twenty-six in the average size, which are 5 mm. in their greater diameter; the columella is spongy, not well developed in the smaller individual corallites. Type and Localily. — Xo. !)8(), Hubbard Collection, Columbia Uni- versity, Xew York City. Occurrence. — Areeibo grou]). Remarks. — This species is distinguished from Goniastrea cfinalis Vaughan by the larger calices, fewer septa and thicker common wall. It differs from Goniastrea variaUUs Duncan by tlie thicker common w all. 202 SCJEXTIFIC SLRVJjy OF PORTO RICO Goniastrea pectinata (Ehrenberg), 1834 Plate XXXIII, Figure 2 Astraca pectinata Ehreiiberg. Koralleutli. Roth. Meer.. 1834, p. 96. Astraca faviatcUa Dana. U. S. Expl. Exped., Zooph.. 1846, p. 241, PI. XIII, figs. 2-2(1. Astraea simiosa Dana, U. S. Expl. Exped., 1846, p. 243, PI. XIII. figs. 5-5c. Goniastraea pectinata Klunzinger, Korall. Roth. Meer., 1879, Pt. 3, p. 34, PI. IV, fig. 6; Yaughan, Carnegie Inst. ^Yashington, Dept. of Marine Biology, Pub. 213, 1918. Goniastraea favus Klunzinger, Korall. Roth. Meer., 1879, Pt. 3, p. 35, PL IV, fig. 4, PI. X, fig. 7. Description. — Corallum forms massive heads; the corallites are poly- gonal, varying from 5 to 12 mm. in maximum diameter; the larger are oval in cross-section and present stages in reproduction by fission; the walls are thick, costated and with acute summits formed on the costa in the central part of the domed surface; the outer septal ends of adjacent calices meet ui)on the wall forming costa ; often a large septum is opposed by a small one ; the larger septa end in paliform lobes at the margin of the trabecular columella. Locality. — i4<)-:i(;T. lieeds Collection. American Museum of Xatural History. Occurrence.- — Eecent. Genus. — Hydnophora Fischer de Waldheim, 1807 H jid nophora ¥isL-]wr de Waldhi'im. Mus. Deniidoff. III. 1807, p. 29.".. 1 pi.; Fischer de Waldheim. Recherches sur les Ilydnophores. 1810. pp. 7-13. 1 pi.; Fischer de Waldln'ini. Oryctographie du gouvernement de Mos- cou. 18;i()-.".7, p. 155; Milne Edwards and Haime. Compte Rendu de I'Acad. des Sci., 1848, p. 493; Milne Edwards and Ilainie, Ann. des Set. Nat., ser. 3. XI. 1849. p. 299: Milne Edwards and Ilaimo. Hist. Nat. Corall.. II, 1S."7. i». 41S: VauLrhan. Carnegie Inst. Washington. Dept. of Marine I'.iol. Pub. 2i::. 191s. p. liil. Hiirella Delage and Ileroiiard. Traite d(> Zool. concrete. II. 19()1. Pt. 2. p. (i-js. Genotype. H jidiiorithnrd (h itiid'irii Fischer de Waldheini := Mtiilrctxtni c.re.sa Pallas. Elench. Zooph., 1766. p. 290. l)ef luni. : the c(dlines are thin, irregularly curved and sliarply crested: the <-((liiMicll;i is pniiily developed. Type anti LoraUiy. — No. !M2. Hubbard Collection, Columbia Uni- versity, New ^'()l■k City. Occurrence. San Scliastian Sliale. C0RYEr.L-(}fiLsr:\. rossfi.' con \ls or roirro inco 205 Genus. — Leptoria Milne Edwards and Haime, 1848 Lcptoriu. — Milne Edwards and Ilaiuu'. ("oinpte lU-iidu, XXVII, 184. p. 422, I'l. CIV, tigs. 2, 2a; PI. CV. Description. — Corallum forms a pedunculate expansion with llattened or curved surface; calcinal valleys deep, steep-sided, sinuuus and often long, measuring o to IG mm. in width and from 8 to 10 mm. in depth; calcinal centers distinct and from to 2\ mm. apart: culliiial summits marked by a narrow furrow, l.n to 1.") mm. wide: 19 to 22 septa in 1 cm.; columella pi-esent or absent: dissepiments well developed: the lower sur- face of the coiallniii is radiallv costated, the costae averaging I imn. apart. Ti/fie. — Xo. ;52r)()OGa, I'. S. National Musenui, LinaJihl. — M()-T-^,-T;), iJeeds Collection, .\nicrican Museum of Xatui'al History. ()( iitrrence. — San Sebastian Shale. (JciHis. — Metastraea Mihic h^dwards ami llaiiiu', 1857 Mcttistnicit Millie Edwards and ilaiine. Hist. .Nnl. des Corall.. II. 1N'>T. p. r>25. (ienotype. — rriomtstrrii . .' iiriniiitiacd Milne Edwards and Ilainie. Ann. des Sci. .Xat.. ser. :;. XII. IMM.. p. ll'.T (Recent). M, hishiicd \ I'allas, 17(;6|. Miidiciiiini (jnlaxca Ellis and Solander, Nat. Hist. Zooph., 1786. p. 16S. A.stnii nitlidns astroitcs Oken, Lehrb. Naturg., Th. 3, Abth. 1. 1S15. p. 65. A.^trca (Siderastrea) galaxca Blainville, Diet. Sci. Nat., LX, 1830, \). 3.".5. Descripiion. — Tiie following is tlic original description as given by Blainville: — CORYELL-OIILSIjy. FOtt I i. Sup. .Mriii. ('(•rail. AntiUcs. 1866, p. 83 ( nf rcpiiiil ) ; Ducbassaing. Rev. Zoopb. Antilles, 1S70. p. 30. Ill li(islr(i< (t inili-nirn^ix Ducbassaing and Miclu'lut ti. Sup. Mem. Corall. Antil- les, l.S6(;, p. 86 (of reprint) ; Duncan, <4eol. Soc. London Quart. Jour., XXIV, 1867. p. 24; Ducbassaing, Rev. Zoopb. .\ntilles, 1870, p. 30. C/J(illi"iiiiiriih(i iiiitUjnvnsis Vaugban, U. S. Nat. Mu.^. Bull. 103, 191!). p. 463, I'l. ex XIX, fig. 2; PI. GXXX. figs. 1-3; PI. CXXXI. figs. 1-4: PI. CXXXII, figs, l-2b; PI, CXXXIII, tig. 1, coryI':l!.-ali : the columella is trabci-ulate. CORYELL-oin.si:\. FOSSIL coh'M.s or I'orto RICO 2ir Bange. — Tertiary to Eecent. Remarks. — Diploastrea is distinguished from Cyalhomorplta in its coarse perforation and dentation and in the absence of pali. Agathiphyllia is similar to Diploastrea in having no pali according to Eeuss. It is necessary to study the type specimen before the relation of these two genera can be determined and until that is done the genus of Diploastrea will be recognized. Brachyphyllia cannot be recognized without further study of the type, B. dormitzeri. Diploastrea crassolamellata (Duncan), 1863 Plate XXXIX, Figure 2 Astraea crassolamellata Duncan, Geol. Soc. London Quart. Jour., XIX, 1863, pp. 412-417, PI. XIII, figs. 1-7. Heliastraea crassolamellata Duchassaing and Michelotti, Sup. Corall. Antilles, 1866, p. 86 (of reprint) ; Duncan, Geol. Soc. London Quart. Jour., XXIV, 1868, p. 24; Duchassaing, Rev. Zooph. Antilles, 1870, p. 30. OrUcella crassolamellata Vaughan, Geol. Soc. London Quart. Jour., LVII, 1901, p. 497. BrachyphylUa sp. Vaughan, Geol. Soc. London Quart. Jour., LVII, 1901, p. 497. Diploastrea crassolamellata Vaughan, U. S. Nat. Mus. Bull. 103, 1919, p. 470, PI. CXXXV, figs. l-5b ; PI. CXXXVI, figs. 1-lb ; PI. CXXXVII, figs. 1-5. Description.— Corallmn massive, expending upward and outward from a small base, the upper surface flat or dome-shaped, deeply grooved between the calices; the lower surface is incompletely epithecated; the caUces are large, shallow, raised above the intercalicular tissue and vary from 6.25 to 27.5 mm. in diameter; the septa are in several cycles (four to nine) and very porous; synapticulae and trabeculae are abundant within and without the calyx ; the columella is well developed, large and trabeculate, often measuring 5 mm. in diameter ; the calicular walls are porous ; costae are present and observed wherever the epitheca is absent. Xype,— Coll. Geol. Soc. London; "Marl formation," Antigua. Locality.— U0-57,-58,-U0,-S78,-4:U, Eeeds Collection, American Mu- seum of Natural History, and 80?, 81? Hubbard Collection, Columbia University, New York City. Occurrence.— Ssin Sebastian Shale, Ponce Formation and Lares Lime- stone. • 2X8 sviExriF/c .S77n'/;v of porto rico Madreporaria Perforata family. acroporidae verrill Genus Acropora Okeii, 1815 Acroporidae Verrill. Trans. Conn. Acad. Arts and Sci., XI, 1901. p. 16.3; (not Acroporidae Canu, 1913, which was made to include several genera of Cheilostome Bryozoa). Genus — Acropora Oken, 1815. Acropora Olien, Lehrb. Naturg., Th. 3, Abth. 1. 1815, p. 66; Verrill, Trans. Conn. Acad. Arts and Sci., XI. 1901. p. 161; 1902. p. 208; Yaughan. Carnegie Inst. Washington, Dept. of Marine Biol. Pub. 213, p. 159; Vaughan, U. S. Nat. Mus. Bull. 103, 1919, p. 479; Zittel, Text Book of Paleon., I, 1913, p. 107. Madrcpora Lamarck (not Linne, ed. X, 1758, p. 792), Hist. Anim. sans Vert., II, 1816, p. 277; ed. 2, 1836, p. 445; Dana, Zooph., 1846, p. 435; Milne Edwards and Haime, Hist. Corall., Ill, 1860, p. 1.32 ; Rathbun. Catalog, r. S. Nat. Mus., X, 1887. pp. 10-19 ; Klunzinger. Korall. Koth. Meer., II, 1879, p. 2 ; Brook, Catalog. Mad. Brit. Mus., I, 1893, p. 22. /sopora..— Studer (as subgenus), Monatsber. Akad. Wiss. Berlin, 1878, p. 535; Vaugban, Fossil Corals Curaco, 1901, p. 68; Vaughan, Bull. U. S. Fish Comm. for 1900, II, 1901, p. 312. Heteropora Ehreuberg, Korall. Koth. Meer., Abhandl. K. Acad. Wiss. Berlin for 1832, 1834, p. 323; (not Blainville, whose genus was referred to a Bryozoan). Acropora Reuss, Die fossilen Anthozoen und Bryozoan der Scbicbtenguippe von Crosaro, Denkschrift. der K. Akad. der Wissenschaft. Wein, XXIX, 1869, p. 277, was used for a Bryozoan species. In U. S. Nat. Mus. Bull. 106, 1920, pp. 318-320. ("iiiiu and Bassler added two new species of Cheilo-stome Bryozoa to the genus .of Reuss. In Zittel, Traite de ralcontob)gie, 1883, Acropora (a Bryozoan genus) was included under Vltivularia as a synonym. Further study is necessary to determine what should be done with the Bryozoan species now assigned to the genus Acropora, a name that can be properly used only for a coral species. Genotype. — Millcpora muricata Linnaeus. Millcpora miiricata Linnaeus, Syst. ed. 10, 1758, p. 792. Madrcpora tnuricuta I'allas, Elench. Zooph., 1706, p. 327; Linnaeus, Syst. ed. 12, 1767, p. 1279. Madrcpora ccrrironii.s Lamaink. Hist. Anim. sans Vi-rt.. 1816. pp. 278, 281. Acropora inuricata Oken, Lehrb. Naturg., Th. 3, Abth. 1, 1815, p. 66. Description. — Corallum forms branching, flabellate or palmate colo- nies; the calices are small, circular pits at the end of tubular pro- jections raised above the surface of the porous coenenchyma; they pro- trude most conspicuously upon the upper surfaces of the palmate forms or nearer the ends of the branches in the branching colonies; the septa are often rudimentary and in two cycles; the two major ov directive CORYELL-OllLtiEy. FOSSIL COh'ALS Of /'Oh'TO lUCO 2l\) septa in some calices form a conspicuous partition ; the columella is rndimentary or absent; the costal ridges are numerous oi' rcplaciMl by papillose tissue. Range. — Tertiary to Recent. Acropora crassa Coryell, new species Plate XXXIX, Figure 3 Description. — Corallum forms bilaminate, palmate fronds or masses with the lamina on the lower side of the broad fronds much thinner and carrying a less number of corallites than the upper lamina ; fronds vary in thickness from 1/2 to 31/2 inches; the corallites are small, 0.5 to 0.8 mm. in diameter, circular in cross-section and separated from one another by a distance equal to or less than their diameter; the coeuenchyma is coarsely porous; the calcinal walls are thick and often in contact in adjacent individuals; the directive septum is well developed and in many calices it unites with the opposite one. Type. — X"o. 23006, American Museum of Natural History. i^ocaZi^//.— 440-346,-368,-369, Reeds Collection, American Museum of Xatural History, and 918, 968, 987, 989, 1089, Hubbard Collection, Co- lumbia University, Xew York City. Occurrence. — Ponce Formation. Remarl-s. — This species is distinguished from Acropora palmata by the thick, calcinal walls, closer average spacing of the corallites and the thicker and less undulating palmate corallum. Acropora palmata (Lamarck), 1816 Plate XL, Figure 1 Madrcpora palmata Lamarck, Hist. Nat. Anim. sans Vert., II, 1816, p. 270; Gregory, Ann. Mag. Nat. Hist., ser. 7, VI, 1900, p. 29. 3Iadrepora muricata palmata Brook, Brit. Mus. Cat. Madrep. corals, gen. Madrepora, 1893, p. 25. Isopora muricata palmata Vaughan, U. S. Fish Comm. Bull, for 190O, II, 1901, p. 313, Pis. XXVI, XXVII. Acropora muricata palmata Verrill, Trans. Conn. Acad. Arts and Sci., XI, 1901, p. 166. icropora palmata Vaughan, Washington Acad. Sci. Jour., V, 1915, pp. 597, 598- Vaughan, Nat. Acad. Sci. Proc, II, 1916, pp. 95, 100; Vaughan, Carnegie Inst. Washington Yearbook No. 14, 1916, pp. 227-230; Vaughan, U. S. Nat. Mus. Bull. 103, 1919, p. 483. Description.—CovaWnm forms undulating and palmate branching colonies; the corallites are circular and protrude above the finely porous coenenchyma, extending out farthest on the margins of the branches ; the 220 SCIENTIFIC SURVEY OF PORTO RICO calcinal pits vary from 0.5 to 0.8 mm. in diameter; costal ridges are numerous; six septa are well developed, a single directive or two oppo- sites are more prominent than the rest; the coeuench)'ma of well pre- served or recent specimens is very spinous ; calcinal walls are thin. Localitij. — 937, H 2, Hubbard Collection, Columbia Universit}^ Xew York City. Occurrence and Range. — Pleistocene and Eecent. Acropora panamensis Vauglian, 1919 Plate XL, Figure 2 Acropora panamensis A'aughan, U. S. Nat. ]\Ius. Bull. 103, 1919, p. 480, PI. CXLI, figs. 1-2. Description. — Corallum composed of thick branches varying from 12 to 20 mm. in diameter and becoming compressed below a point of bi- furcation, where the greatest diameter often measures 30 mm. ; the coral- lites are protuberant, projecting upward and outward; the average in- dividual extends from 1.5 to 2.0 mm. above the coenenchyma; the di- ameters of the calices vary from 1.5 to 3.5 mm.; they are scattered over the branches irregularly, in some places more closely spaced than in others, but commonly separated from one another by a distance equal to the diameter of the calice; costal ridges are numerous; synapticulae are present; the septa occur in two cycles; the directives are well de- veloped but do not form dimidiate calices; coenenchyma is porous and near a corallite protuberance it is covered with the extended costal ridges. Type. — No. 325042a, U. S. National Museum ; Emperador Limestone. Locality. — 440-51-,-()2,-64,-330,-340,-454, Eeeds Collection, American Museum of Natural History, and 880, 896, 897, Hubbard Collection, Columbia University, New York City. Occurrence. — San Sebastian Shale, Ponce Formation and Lares Lime- stone. Genus. — Astreopora Blainville, 1830 Astreopora Blainville, Diet. Sci. Nat., LX. 1830. p. .348; Milne Edwards and Ilainic. Coniptc Itcndu do I'Acad. 8 ; Milne Kdwards iiiid llaiiiu-. Arclii. du Mus. d'llist. Nat.. \'. Is.'.l, p. 141; Milne Edwards and Ilainie. Hist. Nat. des Corall.. 111. 1S(K). p. 167; Bernard, Brit. .Mus. Cat. Madreporaria. II. IS'ti;. p. 77: Vauglian. Carnegie Inst. Wasliingtdii I'ul.. •2\:\. litis, i-. 14.".; Vauirliaii. I'. S. Nat. Mus. Bull. 103, 1919, p. 48;',. Genotype. — Astreopora nii/riophthalma (Lamarck) . .istrea tinniophthnlma Lamarck, Hist. Nat. Anim. sans Vert., ISIG. p. 260. CORYELL-OHLSEN, FOSSIL CORALS OF PORTO lilCO 221 Description. — Corallum consists of massive or branching colonics with porous coenenchymous tissue filling the spaces between each individual; the surface of the intercalicular area is smooth, canaliculate or bearing small spine-like structures; the calices are irregularly rounded, slightly protuberant; the septa are irregularly developed, consisting of a dif- ferent number of cycles in different individuals of the same colony; a columella is present but often poorly developed. Range. — Tertiary to Eecent. Astreopora antiguensis Vaiighan, 1919 Plate XL, Figue 3 Astreopora antiguensis Vaughan, U. S. Nat. Mus. Bull. 103, 1919, p. 484, PI. CXXXIX, figs. 3, 3a ; PI. CXL, fig. 1. Description. — Corallum consists of subterete or palmate branches ; the calices are protuberant, irregular in outline, 2 to 4 mm. in diameter and varying from 1.5 to 2.5 mm. apart; the individual corallites are costate ; the coenenchymous tissue appears somewhat smooth and porous except where the costae are present; the septa appear arranged in two or three incomplete cycles; the columella may or may not be present. Type. — Museum of Comparative Zoology, Harvard University. j^o^ality.— 4:4:0-321, Reeds Collection, American Museum of Natural History. Occurrence. — Ponce Formation. *FAMILY. — PORITIDAE DANA * Genus. — Goniopora Blainville, 1830 Goniopora Blainville, Diet. Sci. Nat, LX, 1830, p 359; Quoy and Gaimard, ■ Voyage de TAstrolabe, ZooL, IV, 1833, p. 218. Genotype.— Gowjopo/o pedunculata Quoy and Gaimard. Description.— CoTsdlum forms compressed or cylindrical branches, lamelliform foliations or subhemispherical masses; calices polygonal, 1.5 to 4.0 mm. in diameter; wall costated, spiny or granular; the septa, usually 24 in number, are arranged in the gonioporid plan, six primaries extending directly to the columella with a triplet group of a secondary and two tertiaries between each, and sometimes a directive plane formed by two opposite primaries can be distinguished; the columella is com- posed of a network tangle of calcareous tissue; synapticulae and pali are present. Range. — Tertiary and Eecent. 222 SCIESTlFir SilRVFY OF PORTO RICO * Goniopora canalis Vaughaii, 1919 Plate XL, Figure 4 Goniopora canalis Vaughan, U. S. Nat. Mus. Bull. 10.3, 1919, p. 494, PI. CXLVI, figs. 1-3. Description. — Corallum forms compressed branches ; the calices are polygonal, varying in diameter from 2 to 4 mm. and on unweathered surfaces have a depth of 1.0 to 1.25 mm.; the walls are costate, reticu- late, ridge-like, enclosing a single calice or a longitudinal series of two to four and varying in thickness from 0.75 to 1.25 mm.; the septa are gonioporoid, tapering inward with two cycles reaching the inconspicuous columella tangle; no pali observed. Tye.—No. 325052, U. S. National Museum. Locality. — 440-39,-451, Eeeds Collection, American Museum of Natural History, and 8?0, 878, 891, 996, Hubbard Collection, Columbia University, New York City. Occurrence. — San Sebastian Shale and Lares Limestone. * Goniopora cascadensis Vaughan, 1919 Plate XL, Figure 5 Goniopora cascadenms Vaughan, U. S. Nat. Mus. Bull. 103, 1919, p. 497, PL CXT.VI, figs. 6-9. Description. — Corallum forms small subcylindrical branches; the calices are polygonal and vary in diameter from 1.5 to 2.5 mm. : the walls are thin, costate and reticulate or apparently absent in some places ; the septa are in three cycles, two of which reach the well developed, often styloform columellar tangle; the pali are slightly developed as a cycle of thickenings around the columella. Type.—^o. 325072, U. S. National Museum. Zom/i///.— 440-38,-339,-373, Keeds Collection, American Museum of Natural History. Occurrence. — San Sebastian Shale and Ponce Formation. * Goniopora clevei Vaugliaii, 1919 Plate XL, Figure G (li,nii>itcra IILSi:\. FOSSIL COh'ALS OF /'Oh'TO RICO 225 * Goniopora jacobiana Vaiii^lian, 1919 Plate XLI, Figure 3 Goniopdia jacoUana Vaughan, U. S. Nat. Mus. Bull. 103, 1919, p. 492, PL CXLIV, figs. l-3a. Description. — Coralluiu forms a hemispherical or columniforiu mass of superimposed lamellae; the calices are polygonal and var}^ from 2.5 to 3.5 mm. in diameter; the walls are thin, costated or liaving a retic- ulate structure where the thickness reaches 1 mm. ; septa are gonioporid and thin : the columella is inconspicuous and apparently absent in some calices ; no pali observed. Tijpe.—y.o. 325077, U. S. National Museum. Locality. — 440-342, Reeds Collection, American Museum of Natural History and 942, 943, Hubbard Collection, Columbia University, New York City. Occurrence. — Ponce Formation. * Goniopora panamensis Vaughan, 1919 Plate XLI, Figure 4 Ooniopora panamensis Vaughan, U. S. Nat. Mus. Bull. 103, 1919, p. 4S9, PI. CXLII, figs. 2-2b. Description. — Corallum is large, consisting of superimposed, flattened or domed plates; the calices are polygonal, varying from 2.5 to 3.5 mm. in diameter and often reaching a depth of 2.0 mm.; in many places the calices occur in series surrounded by a definite, common, ridge-like wall ; the individual walls are thick, reticulated, and at the junction angles, acervuline; the septa are gonioporid in arrangement and number, two cycles of which reach the small columellar tangle ; pali are inconspicuous. .Type.~^o. 325053, U. S. National Museum. Locality. — i40-39,-47,-50,-366, Eeeds Collection, American Museum of Natural History, and 860, Hubbard Collection, Columbia University, New York City. Occurrence. — San Sebastian Shale and Ponce Formation. * Goniopora portoricensis Vaughan, 1919 Plate XLI, Figure 5 Gonioponi portoricensis Vaughan. U. S. Nat. Mus. Bull. 103, 1919. p. 49.5, PI. CXLVI, figs. 4, 5. Description. — Corallum forms cylindrical or compressed branches; the calices are polygonal, shallow and vary in diameter from L5 to 2.0 mm. : 226 SVIEXTIFJC sr RVEY OF PORTO RICO the walls are thin and appear as if formed by the fusion of the ends of the septa; tlie septa are thin and in three cycles; the columella is incon- spicuous; the pali are not always sufficiently developed to be distin- guished from the septal structures. Type. — Xo. 325061, U. S. Xational Museum. Locality. — i40-38,-5 7,-62,-454, Eeeds Collection, American Museum of Xatural History. Occurrence. — San Sebastian Shale and Lares Limestone. * Goniopora regularis (Duncan), 1863 Plate XLII, Figures 1, la Alveopnra dacdalaea var. regularis Duncan, Geo. Soe. London Quart: Jour., XIX. 1S63. p. 426. PI. XIV, figs. 4a, 4e. Alveopora dacduluca Dimeau, Geol. Soe. London Quart. .Juur., XXIV, 1867, p. 25. Alveopora regularis Vaughan, Geol. Reich. Mus. Leiden Samml., ser. 2. II, p. 71. Uoniopora regularia Vaughan, U. S. Nat. Mus. Bull. 1U3, lUlU, p. 491. Description. — Corallum is of an undulate, turbinate or lobulate (■(iluiiiniform shape; calices are polygonal, varying from 1.5 to 2.5 mm. in tliameter ; tlie walls are distinct in places, consisting of reticulate structure; the septa are tliin, granular and gonioporid in arrangement and number; the ])late-like pali are conspicuous. Locality. — 440-320, Reeds Collection, American Museum of Natural History, and 865, Hubbard Collection, Columbia University, Xew York City. Occurrence. — San Sebastian Shale and Ponce Formation. * Gemis. — Pontes Link, 1807 I'ltriti.s I.iiiU. lU'schroih. Xatur. Samiul. Kostock. IsoT. ]i. ICl': Vaughan, S.iiuiiil. (Jcol. Kciclis. Mus. Leiden, ser. 2. II, V.HH. ]). T."*. : Vaughan, U. S. Conini. of Fish and Fisheries Bull, for 1900, II, 1901. p, .S14. Tl. XXVIIl: Vauglian. I'roc. Biol. Soe. Wash., XV, 1902, p. ">(>: Bernard, I'oritos of tlic lndo-1'adfic Kegion, 190"), p. 3n.3. (35 plates) : Bernard, I'orites of tlie Atlantic and West Indies. 1906, ]>. 144, (17 plates) ; \'auglian. Carnegie Inst. Washington I'nh. 2i:>, 191S, p. i;is. Gcimtype. Mailri i>(ira poritcs Pallas. prscripfioii. — Coralhim forms foliaceous or ramose tuffs, incrusta- tions. lieiiiis|ilieri( al or lolird masses with a l)asal epitheca; corallites with tralieculate walls; calices small, ])olygc)niil ; septa trabeculate. CORYELL-OHLSEX, FOSSIL CORALS OE J'ORTO RICO 227 spinose, twelve in number; eolnniella a network tangle at the center of the corallite and often surmounted hy a styliform trabecula; pali five or six, often indistinct, forming a circle about the columellar tangle; synapticules and dissepiments occur; tabulae are porous. Range. — Cretaceous to Eecent. * Porites anguillensis Yauglian, 1919 Plate XLII, Figure 2 Pontes anguiUcusis Vauglian, U. S. Nat. Mus. Bull. 103, 1919, p. 504, PI. CXLIX, figs. 1-lb; PI. CL, fig. 5. Description. — Corallum composed of thin, undulating, superimposed laminae; calices shallow, subcircular, 1.7 to 2.3 mm. in diameter; walls of dense, costate and perforate coenenchymous tissue, 0.8 to 1.0 mm. in thickness; the twelve septa thick, in poritid arrangement; the six pali are before the inner ends of the septal groups; synapticulae are well de- veloped, forming three rows in the wall and a ring about the pali; trabeculae in columellar tangle are coarse ; an axial tubercle is present. Type. — University of Upsala. Lomitf?/. — 110 -40,-45',-16,-50,-l 19,-311,-446,-458. Eeeds Collection, American Museum of Natural History, and P. E. 98 (b), Hubbard Col- lection, Columbia University, New York City . Occurrence. — San Sebastian Shale, Lares Limestone and Ponce For- mation. * Porites astreoides Lamarck, 1816 Plate XLII, Figures 3, 3a Porites astreoides Lamarck, Hist. Nat. Anini. sans Vert., II, 1816, p. 2(59; Rathbun, U. H. Nat. Mus. Proc, X, 1887, p. 354; Vaughan, U. S. Fish Comm. Bull, for 1900, II, 1901, p. 317, PI. XXXII; PL XXXIII; PI. XXXIV. figs. 1, 2; Verrill, Trans. Conn. Acad. Arts and ScL, XI, 1902, p. 160, PI. X-XXI, fig. 4 ; Vaughan, Carnegie Inst. Washington Year- book No. 10, 1912, pp. 148-156, PL IV, figs. 3a, 3d, 3e ; PL V, fig. 5b; PL VI, figs. Ic, 2e ; Vaughan, Washington Acad. Sci. Jour., V, 1915, p. 597; Vaughan, Nat. Acad. Sci. Proc, II, 1916, p. 98; Vaughan, Carnegie Inst. Washington Yearbook No. 12, 1916, pp. 226-228, 231; Duerden, Nat. Acad. Sci. Mem., VIII, 1903, p. 550, Pis. III-V. Porites verrilll Verrill, Trans. Conn. Acad. Arts and Sci., XI, 1902, p. 101, PL XXXI, fig. 5. Description. — Corallum massive, subhemispherical, with a regularly curved or knobby surface; calices polygonal, deep, 1 to 2 mm. in diame- 228 {iClESTIFIC SURVEY OF PORTO RICO ter; there are twelve principal porous septa with dentate upper margins and arranged in regular poritid form ; walls are thin ; the columella is composed of a network tangle ; the pali are poorly developed. Locality. — i40-18,-3GT,-162, Eeeds Collection, American Museum of Natural History. Occurrence. — Recent, Ponce Formation ? and Cibao Limestone ?. " Porites baracoaensis Vaughan, 1919 Plate XLII, Figure 4 Porites baracoaensis Vauglian, U. S. Nat. Mus. Bull. lO.'',, 1919, p. 499. PI. CXLVII, figs. 1, la. Description. — Corallum composed of slender branches; calices shal- low, polygonal, 1.25 to 2.25 mm. in diameter; walls thin, costate and with squamae present within most of the calices; septa of poritid ar- rangement; there are usually six pali, one before the inner end of each group of septa and united at the base of the calyx to the columellar tangle. Type. — Xo. 325009, U. S. National Museum. Localily. — 440-362, Reeds Collection, American Museum of Natural History, and 1)55, Hubbard Collection, Columbia University, New York City. ' Occurrence. — Ponce Formation. * Porites douvillei Vaughan, 1919 Plate XI.IT. Figures 5-7 roriti's . 47; Vaughan. /'. forma rhtvaria. U. S. Fish Cr»mm. P.iill. for 1!XM>, II. 1!M)1. p. 316, PI. XXIX; PI. XXXI. fig. 2; Vaughan, Carnegie Inst. Wasliington Yearbook No. 10. 1912, pp. 148, 152, 156, PI. IV. fig. 4c: PI. VI. figs. 3, 4; Vaughan. Washington Acad. Sci. .Idur.. V, 191.5. ]>. .597: Vaughan. Nat. Acad. Sci. Proc. 11. 1916. l»p. 95, 98; Vaughan, Carnegie Inst. Washington Yearbook No. 14, 1916,. 11. 228; Cregory. Quart. .lour. Ceol. Soc. Lcuidon. LI. 1895, p. 182. I'orilis JlciHitxa Dana, Zoojili. Explor. i;\|ied. Wilkes. 1S48. )>. .554. PI. LI II. lig. 6; Milne Edwards niid ll.'iiinc. .\nu. Sci. \;il. /mil., .>r;ill.. .\nl. Mem, UdV. .Vccad. Sci. CORYELL-OHLSEX, FOSSIL CORALS (>/•' I'ORTO RICO 231 Tor., ser. 2, XXIII. 180G, p. 191; Dncliassaiiiff. Rev. Zoopli. .\nt., 1S70, p. 32. Porites flabeUiformis Lesueur, Mom. Mus. Hi.st. Nat., \'I, 1S2(>, p. 28!); I)e.s- longchamps, Encycl. Meth. Zoopli., 1824, p. G52 ; Milne Edward.s and Haime, Ann. Sci. Nat. Zool., ser. 3, XVI, 1S51, p. 31; Milne Edwards and Haime, Hist. Nat. Corall., Ill, 1860, jt. 178; Duchassainj,' and Mielielotti, Suppl. Mem. Roy. Aeoad. Sci. Tor., ser. 2, XXllI, 18GG, p. 190; Duchassaing, Rev. Zooph. Ant., 1870, p. 32. Porites solanderi Duchassaing and Mielielotti, Mem. Roy. Accad. Sci. Tor., ser. 2, XIX, 18G1, p. 358, Suppl. Mem. Roy. Accad. Hci. Tor., .ser. 2, XXIII, 186G, p. 189; Duchassaing, Rev. Zooph. Ant., 1870, p. 32; (^uelch, Zool. Challenger Exped., Pt. XLVI, 1886, p. 13. Porites plumieri Duchassaing and Michelotti, Suppl. Mem. Roy. Accad. Sci. Tor., ser. 2, XXIII, 1866, p. 190, PI. X, tig. 14; Duchas.saing, Rev. Zooph. Ant., 1870, p. 32; Quelch, Zool. Challenger p]xped., I't. XLVI, 1886, p. 13. Porites macrocephala Duchassaing and Michelotti, Suppl. Mem. Roy. Accad. Sci. Tor., ser. 2, XXIII, 1866, p. 189, PI. X, fig. 15; Duchassaing, Rev. Zooph. Ant., 1870, p. 32. Porites recta Lesueur, Mem. Mus. Hist. Nat., VI, 1820, p. 288, I'l. XVII, tig. 16; Deslongchamps, Encycl. Meth. Zooph., 1824, p. 651; Dana, Zooph. Explor. Exped. Wilkes, 1848, p. 556. Porites valula Duchassaing and Michelotti, Suppl. Mem. Roy. Accad. Sci. Tor., ser. 2, XXIII, 1866, p. 94 (of reprint). Porites nodifera Klunzinger, Die Koralleuthiere des Rothen Meeres, Pt. 2, p. 41. Porites porites Vaughan, Biol. Soc. Washington Proc, XV, 1902, p. 56 ; var. Vaughan, Carnegie Inst. Washington I'earhook No. 7, 1909. p. 135. Description. — The corallum forms ramose tufts with basal epitheca; the calices are shallow pits or even flush with the surface, varying from 1.5 to 2.0 mm. in diameter; the twelve septa are trabeculate, and often show a bilateral S3'mmetry; they are grouped as follow, a single directive, four lateral pairs and a ventral triplet; a palus is present before each septal group, forming a ring of six (sometimes five) about the columellar tangle; the upper end of the columella may end in a single tubercle or appear papillose; the wall is porous and varies con- siderably from a thin division among the younger polyps on the end of the branches to as much as 0.5 mm. on the older portion of the coral- lum; synapticulae and dissepiments are present. Locality. — This species occurs in the Bermudas, Bahamas, Florida, Vera Cruz and other Caribbean regions, also in the Indian Ocean reefs. The specimen in the Hubbard Collection of Columbia University is from Porto Eico, locality 935. 232 SCIENTIFIC SURVEY OF PORTO RICO Occurrence and Range. — Miocene to Eecent. The specimen that forms the basis of this description is Eecent. * Porites toulai Vanglian, 1919 Pontes toulai Vaughan, U. S. Nat. Mus. Bull. 103, 1919, p. 501, PI. CL, figs. 1-4. Descriptian. — Corallum composed of slender, subcylindrical or slightly compressed branches; calices shallow, 1.5 to 2.5 mm. in diameter, either surrounded by an individual wall or a wall enclosing several calices in longitudinal series with indistinct divisions separating each calyx; the septal arrangement is irregularly poritid; synapticular tissue forms a palar ring and also a circle near the wall ; an irregularly developed ring of five or six pali surrounds and is connected to the columellar tangle by radial extensions; in some cases a central tubercle is present on top of the columella. Type. — No. 32105a, U. S. National Museum. Locality. — 440-24,-25,-27,-33,-38,-39,-44,-45,-46,-48,-49,-50,-5]^-52,- 56,-58,-62,-63,-64,-92,-123,-299,-325,-330,-331,-339,-362, -383, -442, -451,- 452,-454, Eeeds Collection, American Museum of Natural History, and 924, Hubbard Collection, Columbia University, New York City. Occurrence. — San Sebastian Shale, Lares Limestone and Ponce For- mation. Other fossil corals from Porto Eico, not included in this paper but de- scribed by T. W. A^auglian in U. S. Nat. Mus. Bull. 103, are as follows: Antrcopora portorircnsix Nan^luiu, p. 485. Diploastrea crassolamellata mag- nifica (Duncau), i». 470. BimJOGRAPHY AcHiAuui, A. D. 1)'. 1875. Corall. eocen. del Friuli, p. 70. Bayer, .T. 18SJ>. (Jeognost. Landesuntersucli. Geognost. Jahresh. IT. pp. 150-152. Beknaiu), II. M. 1890, British Mu.>^. Cat. Madrcporaria. 11, p. 77. 1905. Porites of the Indo-I'acilie Region, p. 303. 1900. Porites of the Atlantic and West Indies, p. 144. Blainvim.e, H. D. ue ]82(;. Diet. Sei. Nat., XMII, p. 50. l,s:i(). Diet. Sei. Nat.. LX. pp. 312, 319, 335, 348, 351, 359, 361. BOLTEN, .1. 1798. Museum Boltenianuni. \k 70. Brook, G. 1S03. Calalng. Madrepdia. I'.iilish Mus., 1. pp. 22, 25. CORYELL-OHLSEN , FOSSIL CORALS (>/' I'OUTO RICO 233 Dana, J. D. 184G. Structure and Classificiition of Zoophytes, V. S. Expl. Exped., \)\). 205-2();», 214, 241, 24:5, 2(i:{, :«:!. •.v.^-^, 4:!:.. 1S48. Zoopliytes, IJ. S. Expl. Exped. Wilkes, pp. r)23, 5n4. Delage, Y. and Herouabd, E. lOOl. Traite de Zool, concrete, II, Tt. 2. p. (L'S. Deslongchamps, E. 1824. Encycl. Metb. Zooph., p. G52. DUCHASSAING DE FoNBRESSIN, F. 1850. Anim. Had. Ant., p. IT. 1870. Rev. Zooph. et Spong. Antilles, pp. 26, 28, 30, 32. DUCHASSAING DE FONERESSIN, P. AND MiCHELLOTTI, G. 1861. Mem. Koy. Accad. Sci. Tor., Ser. 2. XIX, p. 358. 1866. Sup. Mem. Roy. Accad. Sci., Tor., Ser. 2, XXIII. DUEBDEN, J. E. 1903. Nat. Acad. Sci. Mem., VIII. pp. 55<), 564. Duncan, P. M. 1863. Geol. Soc. London Quart. Jour., XIX. pp. 412-410, 421-423, 425, 426, 432, 433, 436, 441, 443, 444. 1864. Geol. Soc. London Quart. Jour., XX, pp. 39, 66. 1864. Geol. Soc. London Quart. Jour., XXI. p. 10. 1868. Geol. Soc. London Quart. Jour.. XXIV, pp. 19, 20, 23-25. 1873. Geol. Soc. Loudon Quart. Jour., XXIX, p. 551. 1884. Linn. Soc. London Jour. Zool., XVIII, pp. 45, 105. 1895. Geol. Soc. London Quart. Jour., LI, p. 274. ESPER, J. F. 1797. Ptianzenth., Fortsetz., I. pp. is, 21, 74, 135. Ehrenbekg, C. G, 1834. Korallenthiere des Rothen Meeres, Abhandl. K. Akad. Wiss. Berlin for 1832, 1834 (reprint), pp. 84, 96, 99, 101, 323, 341. Ellis J. and Solandeb, D. O. 1786. Nat. Hist. Zooph., pp. 162, 166, 168. 169, 172. Fromentel, E. de 1861. Introd. Polyp, a I'Etude des Foss., pp. 179, 235. Gardiner, C. I. 1904. Fauna and Geol. Maldive and Laccadive Arch., II, p. 774. Gmelin, J. G. 1790. Linn. Syst. Nat., ed. 13, pp. 3767, 3769, 3774. GOLDFUSS, A. 1826. Petref. Germ., p. 45. Gregory, J. W. 1895. Geol. Soc. Tendon Quart. Jour., LI, pp. 270, 274, 277, 282. 1900. Pah-po. Indica, Ser. 9, II, Pt. 2, p. 59. 1900. Ann. Mag. Nat. Hist., ser. 7, VI, p. 29. GUETTABD, J. E. 1770. Mem. sur les Sci. et les Arts, II, p. 406. 234 SCIENTIFIC SURVEY OF PORTO RICO Hall, J. 18.11. Aiuer. Jour. Sti. and Arts, XI, pp. 398-401. Hoffmehster, J. E. See Vaughau, T. W. Klunzixcer, C. B. 1879. Die Koralleiithiere des Kotlien Meeres, I't. o, pp. 2, 34, 35. Lamarck, J. B. 1801. Syst. Anim. sans Vex't., pp. 871, 375. 1816. Hist. Nat. Anim. sans Vert., II. pp. 246, 258-200, 265, 269, 270, 273, 277-279, 281, 4<>4. 18:^ Hist. Nat. Anim. sans Vert., II, pp. 380, 415, 441, 445. Lamourolx, J. F. 1821. Exp. Meth. (Jenres des Polyps., pp. 58, 61. 1824. Eneycl. Meth., pp. 57, 131. Lesuer, C. a. 1820. Mem. Mus. Hist. Nat. \l, p. 289. LiXDSTROM, (}. 1877. Hiindl. k. Svensk. Vet.-Akad., XIV, No. 6, p. 24. LixK, H. F. 1807. Besehreib. Nat. Samml. Rostock, Pt. 3, !>. 162. Linn 6, C. 1758. Syst. Nat., ed. 10. |)|.. 792, 794. 1767. Syst. Nat., ed. 12, pp. 1276, 1279. Lonsdale, W. 1847. Amer. .Jour. Sti. and Arts, Ser. 2, IV, i>p. 357, 362. MLatthai, C. C. 1914. Linn. Soc. lA)ndon Trans-Zool., Ser. 2, XIIT, p. 72. Mayer, A, G. 1918. Dept. Marino Biology. Carnegie Inst, of Washington. IX. pp. .3-48. MiCHELIN, H. 1840-47. Ic'onograplue Zoophytologiquc, p](. ."i8. 94. MiLNK i;i)\\ Ai{i)s, IL and Haimic, .T. 1848. Compte Rendu de I'Acad. des Sei.. XXVil. i)p. 258, 469, 492, 493, 495. 1849. Compte Rendu de TAoad. des Sei., XXIX. p. 261. 1849. Ann. des Sei. Nat.. Ser. .3, XI. pp. 261. 27(1. 299. 1849. Ann. des Sei. Xat., Ser, 3, XII, p. 137. 18.51. Ann. des Sei. .Nat.. Ser. 3, XVI. im'. 26, 31 1851. Anhi. dn Mus. d'Hist. Nat.. V. p. 141. 1851. Pol. fo.ss. des terr. palaeoz.. i>p. 80, 83, 85 18.57. Hist. Nat. des Corall., II, pp. 2.57, 342, ;i47, 3,57, .384, 402, 418, 463, 47:;, .105, .525, 54.-.. 1860. Hist. Nat. des Corall.. Ill, jip. 72. so. i:','_'. l.-,.-,. KJT. 171. 176. 178. :!01. Oken, L. 181.5. Lehrl». dcr .N.itiug.. Tli. ::. .\hlli. 1. |)p. t;.-.-(;8, 7(h 75. CORYELL-OHLSEy, F0S8IL CORALS OF I'ORTO RICO 235 Orbigny, a. iV. 1S49. Notes snr 1. Fossil Corals, Curaco, p. 68. 1902. Biol. Soc. Washington Proc, XV. p. 5r.. 1902. Geol. Soc. London Quart. Jour., LII, p. 497. 1907. U. S. Nat. Mus. Bull. 59, p. 136. 236 SVIEM'IFIC SURVEY OF PORTO RICO Vaughan, T. W. ltX)T. I'roc. U. S. Nnt. Mus.. XXXII. ]>. 25'J. I'.HY.). Carnegie Inst. Washington Yearboolc No. 7, p. 135. 11)12. Carnegie Inst. Washington Yearl)ook No, 10, pp. 148-156. 1!>15. Carnegie Inst. Washington Yearbook. Nto. 13, p. 3G0. l!Uo. Wasliington Acad. Sci. Jour.. V, pp. oDG, 597. IDKJ. Carnegie Inst. Washington Yearbook. Xo. 14, pp. L'27-230. 1910. Nat. Acad. Sci. Proc, II, pp. 95, 98, 100. 1918. Carnegie Inst. Washington, Dept. Marine Biology, IX, Pub. 213, pp. 75, 85, 117, 119, 121. i:«. 142, 145, 159. 1919. U. S. Nat. Mus. Bull. 103, pp. 335, 336, 338-340, 342, 348-350. 380. 387, 3<.H), 401, 402, 408, 416, 417, 421, 422, 427, 432-435, 451, 454, 463, 467, 469, 470, 479, 480, 483, 484, 488, 489, 490-497, 499, 501, 503, 506. 1922. Geo!, and Pale. Djiua's Corals and Coral Islands, ed. 3, p. 519. 1900. Trans. Conn. Acad. Arts and Sci.. X. pp. 552, 553. 1!K>1-1902. Trans. Conn. Acad. Arts and Sci.. XI. pp. 66. 70, 83, 92-JM, 96. 1(10, 145, KiO. 161, 163, 164. 166. 2{)6. 208. Waldhkim, Fisiieu he 1807. Mus. Demidoff, III, p. 295. 1810, Recherches sur les Hydrophores. pp. 7-13. 18.30-37, Oryctographie du gouveruement dc Moscou, ]>. 1.55, WlUTKIf:!.!), R. P. 1901. Bull. Amer. Mus. Nat. Hist., N, Y., XIV, p. 223. ZrrrEL, K, A, 18S3. Tr.iitc dc I'aleontologie. I. i)p. 247. 257, 2.59. 260, 261, 191.*!. Text Book of Paleon., 1. i)p. 9S. 99. 107. Plate XXVI PAGE Pocillopora portoricerisis Coryell, new species, 32 Fig. 1. Lateral view, about x % Stylophora affinis Duncan, 33 Fig. 2. Showing the arrangement of the septa, about x 5/6 Fig. 2a. Lateral view of the entire specimen, about 5/6 Stylophora goethalsi Vaughan, 33 Fig. 3. Lateral view of the entire specimen ; the lower 2(. is covered with matrix, about x 4/5 Stylophora granulata Duncan, 3-4 Fig. 4. Lateral view of the entire specimen, about x 9/10 Stylophora macdonaldi Vaughan, 34 Fig. 5. A fragment, about x 9/10 Fig. 5a. Diagram showing septal arrangement of Stylophora. Stylophora panamensis Vaughan, 35 Fig. 6. A view showing the nodular surface, about x % Stylophom ponderosa Vaughan, 36 Fig. 7. A specimen showing the massive growth, about x 4/5 Stylophora portohellensis Vaughan, 36 Fig. 8. The terminal of a compressed branch, about x 4/5 Fig. 8a. Diagram showing septal arrangement. Astrococnia decaturensis Vaughan, 37 Fig. 9. A dendroidal s])ecimen, about x 4/5 Scientific Suuvey uf I'okto Kicd a\d tiiio Viecin Islands Vdl,. HI, I'l.ATK XXVI S So, Ba. Plate XXVII PAGE Astrocoenia guantanamensis Vaughan, 38 Fig. 1. A weathered massive specimen, about X 4/5 Astrocoenia meinzeri Vaughan, 38 Fig. 2. A very broad branching specimen, about x 4/5 Astrocoenia ornata Milne Edwards and Haime, 39 Fig. 3. A fragment of a large massive specimen, about x 3^ Astrocoenia portoricensis Vaughan, 39 Fig. 4. A portion of a branching colony, about x 4/5 Fig. 4a. Diagram showing the septal arrangement of Antiguasirea cellulosa ciirrata (Duncan) 42 Fig. 5. A fragment of a corallum, about x 4/5 SciE.vTii-ic ScrtvKY OF Porto Rico axd the Virgin Islands Vol.. Ill, l'i,.\ii; XXVII A- •■■.• ir->. "^^'^-: i'-^A. ,5■^'- ^■^ . •*- »■,'■■, lO' Plate XXVIII PAGE Antiguastrea cellulosa (Duncan), 41 Fig. 1. A portion of a subplanate massive specimen, about X 4/5 Orhicella annularis (Ellis and Solander), 43 Fig. 2. A fragment of a massive corallum, about x % Scientific Survey of Porto Rico axd the Virgin Islands Vol. Ill, Plate XXVIII t- 'S\ :-'#":f r;-'-' ; If'- (»« > *> ^1 X. i'" ^4T '4f ^ rf ..^*» V' Plate XXIX PAGE Orbicella cavernosa (Linnaeus), -15 Fig. 1. A portion of a hemispherical mass, about x 4/5 Orbicella costata (Duncan), -16 Fig. 2. A hemispherical colony, about x 7/1 Orbicella limbata (Duncan), 47 Fig, 3. A portion of an undulating corallum, about x 4/5 Scientific Si;rvev ok I'orto Rico and the Virgin Islands Vol. Ill, Plate XXIX 5'-?^- -.. -T tH-» ^ '' A ».&-'-. JPB ^^^^ «^,t^ ^-^^ ^.''^ '*v^^1K^- '■>:% Plate XXX PAGE Orbicella tampaensis Yaughan, 48 Fig. 1. A head-shaped corallum, about x 4/5 Calamophyllia dendroidea Coryell, new species, 50 Fig. 2. A view of the type specimen, about x % Fig. 3. A few branches showing the habit of reproduction and the fine costae, about x % Calamophyllia portoricensis Coryell, new species, 51 Fig. 4. A view of the type specimen, about x % Scientific Survey of Porto Rico an'd the Virgix Islands Vol. Ill, Plate XXX ^';,:.' ■^*-'. - A-\ J'\ ^ - ~*-,V i- ^ :f^' \^ i t ^ y - / Plate XXXI PAGE C alamo phyllia portoricensis Coryell, new species, 51 Fig. 1, Another specimen somewhat weathered, showing the coarse costa and the manner of branching, about x4/5 Fig. 2. End view of the specimen in fig. 1, about x 4/5 Fig, 3. Lateral view of the type specimen, about 5/6 Fig. 4. End view of the type specimen, showing rounded and compressed stems, about x 5/6 Scn:\TiFic SunvEV of roisTO Rico and thk Virgix Islands Vol. Ill, TlateXXXI »« .•.•" '•».-' .1% ' .i, ■■1 -<«f. " "^ ••' 1^ . ': .4r ; If , ^'"^'V^ ^^M. ^ Plate XXXII PAGE Favites expansa Coryell, new species, S2 Fig. 1. Surface of type specimen, about x % Fig. la. Diagram of the septal arrangement Favites irregularis Coryell, new species, 53 Fig. 2. Surface of type specimen, showing the variability in size of the corallites, about x 4/5 Scientific Survf.y of Porto Rico and the Virgin Islands Vol. Ill, Plate XXXII ^.; ;l. 'I Plate XXXIII PAGE Goniastrea crassa Coryell, new species, 54 Fig. 1. A view of the surface of the type, about x 4/5 Goniastrea pectinata (Ehrenberg), 55 Fig. 2. A portion of the surface of a massive head, about X 7/10 Hydnophora hubhardi Coryell, new species, 56 Fig. 3. A view of the surface of the type, about x 4/5 Lamellastraea crassa Coryell, new species, 57 Fig. 4. A view of the type specimen, about x 4/5 Latomeandra lata Coryell, new species, 59 Fig. 5. The surface view of the type, about x 4/5 Scientific Survey of Porto Rico and the Virgin Islands Vol. Ill, Plate XXXIII 4 -*_ ^^ "5^. ^W^*rf^/^.^ "i-'-^V %. >:^'% « * ^ >1i v. w[L|^ ^ % ^^- ,# ■5?!^, .<>■ -v?' I >«N* i. Plate XXXIV PAGE Leptorla areolata hispida (Verrill), 60 Fig. 1. Surface of a small corallum, about x 4/5 Leptoria phrygia (Ellis and Solander), 61 Fig. 2. A fragment of a massive head, about x 4/5 Maeandra antiguensis Vaughan, Q'2 Fig. 3, Surface of a small corallum, about x 4/5 Maeandra lahyrinthiformis (Linnaeus), 63 Fig. 4. Surface of a fragment of a head, about x 9/10 Scientific Survey of Pokio Uico and the Virgin Islands Vol. Ill, Platk XXXIV Plate XXXV PAGE Manicina willoughbiensis Vaughan, 64 Fig. 1. A portion of the surface of a large specimen, x 4/5 Metastraea planulata Coryell, new species, 66 Fig. 2. A view of about one-third of the surface of the type specimen which is circular-like in expansion, x % Scientific Suuvey of Poino Kico axd the ViR'Jin Islands Vol.. Ill, Plate XXXV 3j9wi*»»ii'>f-'^s*""**; •"^y^^-^^ '™»»'T|?!*'^. ^' ^ \ "» ^ ' 'V t >. ^ ,A > Plate XXXYl PAGE Metastraea planulata Coryell, new species, 6Q Fig. 1. A portion of the surface of the type specimen, x 4/5 Agaricia agaricites crassa Verrill, > 67 Fig. 2. Surface of an undulating corallum, x 4/5 SCIEN-TIFIC SlRVEY OF PORTO RiCO AXD THK VlRi;i\ ISLANDS VOL. III. Pr.ATE XXXVI Plate XXXVII PAGE Agaricm, irregularis Coryell and Ohlsen, new species, 68 Fig. 1. A view of the type, which appears commonly as moulds, about x 5/6 Agaricia sinuata Coryell and Ohlsen, new species, 69 Fig. 2. A view of the type specimen, commonly appearing as moulds, about x 4/5 Pironastraea anguillensis Vaughan, 70 Fig. 3. A view of a portion of the corallum, about x 4/5 Scientific Sdrvet of Porto Rico and the Virgin Islands Vol. in, Platk XXXVII "t i^nv Plate XXXVIII PAGE Pironastraea antiguensis Vaughan, 70 Fig. 1. A surface view, a complete specimen, about x % Sidemstrea conferta (Duncan), 72 Fig. 2. A surface view of a small corallum, about x 5/6 Cyathomorplia antiguensis (Duncan), 74 Fig. 3. A surface view of a typical specimen, about x 5/6 Fig. 4. A specimen with large corallites, about x 4/5 Fig. 5. A fragment with one end polished, about x 9/10 SCIENTU-IC SlRVEY Or I'OliTO UlCO AND THE VlRlIN' ISLANDS Vol. III. Pr.ATE XXXVIII r ■ ■.i*?->^ ^^~,, ■ -;:^;^^,^^ Plate XXXIX PAGE Cyathomorpha tenuis (Duncan), 75 Fig. 1. A view of a small corallum, about x 4/5 Diploastrea crass olamellata (Duncan), 78 Fig. 2. The surface of a fragment of a massive corallum, about X % Acropora crassa Coryell, new species, 81 Fig. 3. Lateral view of a slightly weathered specimen, x 1 Fig. 3a. View of the entire specimen, about x I/2 Scientific Survey of Torto Rico and the Virdin Islands Vol. nr. Plate XXXIX Vf ■ **• • • • «r • « /"." -"■,' ^ Plate XL PAGE Acropora palmata (Lamarck), 81 Fig. 1. Lateral view of a fragment of a branch, about x % Acropora panamensis Vaughan, 83 Fig. 2. A fragment of a branch showing the prominent coral- lites, about x % Astreopora antiguensis Vaughan, 84 Fig. 3. A view of the end of a branch, about x 4/5 Goniopora canalis Vaughan, 85 Fiff. 4. A surface view of the end of a branch, about x 4/5 Goniopora cascadensis Vaughan, 86 Fig. 5. A fragment of a stem, about x 4/5 Goniopora clevei Vaughan, 86 Fig. 6. A fragment of a branch, about x 5/6 Goniopora decaturensis Vaughan, 87 Fig. 7. A surface view of a corallum, about x % Goniopora decaturensis silicensis Vaughan, 88 Fig. 8. Surface of a weathered specimen with a small portion polished, about x 4/5 Scientific Sukvky oi' Pouro IJico and tiik Viroix Islands V(H,. Ill, I>r,ATK Xfi ,K N i^^Sr^ 8 5 Plate XLI PAGE Goniopora hilli Vaughan, 88 Fig. 1. Surface of one-half of a corallum, about x % Goniopora imperatoris Vaughan, 89 Fig. 2. Surface of a slightly weathered specimen, about x % Goniopora jacohiana Vaughan, 89 Fig. 3. Surface of a fragment of a weathered specimen, about x3/4 Goniopora panamensis Vaughan, 90 Fig. 4. A portion of a dome-shaped undulating plate, about X 4/5 Goniopora portoricensis Vaughan, 91 Fig. 5. Fragment of a branching corallum, about x % Scientific Survf.y of roitTo Rico amp tiik Virgin Islands Vol. Ill, Plate XLF 1**1. ^ Plate XLII PAGE Goniopora regulans (Duncan), 91 Fig. 1. Surface of a dome-shaped mass, about x % Fig. la. Diagram showing typical Gonioporid arrangement of septa. Pontes anguillensis Vaughan 93 Fig. 3. The surface of a lamellose mass, about x 4/5 Porites astreoides Lamarck, 93 Fig. 3. Surface of a weathered specimen, about x 4/5 Fig, 3a. Diagram showing typical Poritid arrangement of septa. Porites baracoaensis Vaughan, 94 Fig. 4. A fragment of a branching corallum, about x 4/5 Porites douvillei Vaughan, 95 Fig. 5. A fragnuMit of a nodose branch, about x % Fig. 6. A weathered fragment, about x % Fig. 7. A typical branch, about x 5/6 < i Scientific Survey of Porto liico and the Vircin I.seand.s \uh. Ill, Plate XLII la 5a ■**-/.%5cy Plate XLIII PAGE Pontes (Synaraea) macdonaldi Vaughan, 95 Fig. 1. A view of lobed surface of a domed corallum, x % Pontes panamensis Vaughan, 96 Fig. 2. A complete corallum, x 2/5 i Scientific SritvEY of 1'okto Itico and tiik Viri;i\ Islands Vol. III. 1'latk XLIII Plate XLIV PAGE Porites panamensis Vaughan, 96 Fig. 1. A portion of the surface of a hemispherical mass, X 5/6 Pontes porites (Pallas), 97 Fig. 2. A branching colony, x % ( Scientific Sirvey of I'orto Rico and the Virgin Islands Vol. Ill, Plate XLIV \ ] THE NEW YORK ACADEMY OF SCIENCES SCIENTIFIC SURVEY OF PORTO RICO and the VIRGIN ISLANDS VOLUME III— Part 4 The Tertiary Foraminifera of Porto Rico J. J . Galloway and Caroline E. Heminwoy NEW YORK Published by the Academy Aphu, 21, 1941 Editor ERICH MAREN SCHLAIKJER Associate Editor (Porto Rico Publications) ROY WALDO MINER SCIENTIFIC SURVEY OF PORTO RICO AND THE VIRGIN ISLANDS This natural history survey of Porto Rico and the Virgin Islands, conducted by The New York Academy of Sciences, xras established in WIS. Continuous pub- lication of the restdts of this survey is made possible through contnbutions from the Department of Agriculture and Commerce of Porto Rico, and the University of Porto Rico. s ■<*- "^ 'LIBRAR' t^**** \ # > THE TERTIARY FORAMINIFERA OF PORTO RICO By J. J. Galloway* and Caroline E. Heminwayj CONTENTS PAGE Introduction 275 Characteristics of the Faunas 276 Age of the Formations 285 Acknowledgments 298 Systematic Description of Species 298 List of New Forms 445 Fossil Localities 446 Explanation of Plates 449 Index 487 INTRODUCTION On the island of Porto Rico, rocks of Tertiary age outcrop in two east-west belts on the north and south sides of the large area of Cretaceous igneous and metamorphic rocks which compose the greater part of interior Porto Rico. The northern area of Tertiary rocks is made up of five formations which strike nearly east-west and dip gently northward. The younger formations overlap to the east so that at the eastern end of the north shore of the island only the Quebradillas, the uppermost Tertiary formation, is present, while in the northwestern part of the island all five formations are found. In order of descending age, these formations are the Quebradillas, the Los Puertos, the Cibao, the Lares, and the San Sebastian, the basal part of which, sometimes called the Collazo shale, lies unconformably on the Cretaceous basement rocks. The Southern area of Tertiary rocks includes two formations, an older one, the Juana Diaz, which probably corresponds in age to the San Sebastian of the north shore, and a younger formation, the Ponce, which probably is the equivalent of several of the north shore formations. The exact position of these various formations within the Tertiary is still not well substantiated, although their local relationships are * Professor of Geology, Indiana University. 1 Associate Professor of Geology, Smith College. (275) 276 SCIENTIFIC SURVEY OF PORTO RICO clear. For the present, we are accepting the age assignments made by Meyerhoff in his recent work on the geology of Porto Rico.^ North Shore Thickness Lower Miocene Quebradillas limestone 700-1500 Los Puertos limestone 600-1000 Upper Oligocene Cibao limestone 250-1000 Lares limestone 1200-1300 Middle Oligocene San Sebastian formation 700 The Ponce formation of the south shore is considered upper Oligocene and lower Miocene in age. Its thickness is about 2,000 feet, while the underlying Juana Diaz is reported as up to 3,000 feet in thickness. CHARACTERISTICS OF THE FAUNA The total foraminiferal assemblage from the Tertiary of Porto Rico represents 25 families, 99 genera with 275 species and varieties,— 88 of which are new. With the exception of the very small fauna from the Lares formation, the north shore formations are all of about the same size. The Ponce formation of the south shore is thought to represent a greater stratigraphic interval than any single one of the north shore formations and this idea is supported by the greater size of the Ponce fauna, — both as to number of families and as to number of genera and species. Table 1 No. of species No. of No. of No. of restricted to For mation families genera species the formation San Sebastian 15 34 75 23 31% Lares 5 6 10 2 20% Cibao 15 34 64 29 45% Los Puertos 11 24 48 10 21% Quebradillas 16 34 66 13 20% Ponce 21 60 180 114 63% An examination of the number of species in each formation which is restricted to that formation illustrates again the greater time interval represented by the Ponce formation, and demonstrates the closer relationship between the various north shore formations than between any one of these formations and the south shore formation. The accompanying check list (table 2) shows the occurrence of the various species in the north shore and south shore formations. ' Meyerhoff, H. A. MonoK. Univ. Porto Rico. ser. B. no. 1. 1933. GALLOWAY AND JIEMINWAY, TERTIARY FORAMINIFERA 211 Table 2 Check List of the North Shore and South Shore Formations a a O TO North Shore c5 1 TO f O -t s- South Shore ® PJ fl> 9» 13 CO (» o" p 09 g- B Lagynidae Pseudarcella patella n. sp Astrorhizidaf; Rhabdammina irregularis W. B. Carpenter. . . . Spirillinidae Spirillina vivipara Ehrenberg Patellina corrugata Williamson MiLIOLIDAE Spiroloculina antillarum d'Orbigny Spiroloculina elongata d'Orbigny Spiroloculina oculina n. sp Quinqueloculina akneriana d'Orbigny X Quinqueloculina apiculata n. sp X Quniqueloculina audacula n. sp Quinqueloculina laevigata d'Orbigny X Quinqueloculina maculata n. sp X Quinqueloculina philippi Reuss Quinqueloculina ponceana n. sp Quinqueloculina pygmaea Reuss X Quinqueloculina seniinulum (Linne) X Quinqueloculina vulgaris d'Orbigny X Trlloculina austriaca d'Orbigny X Triloculina brongniartiana d'Orbigny Triloculina cervicula n. sp X Triloculina elliptica n. sp X Triloculina gibba d'Orbigny X Triloculina inflata d'Orbigny X Triloculina laevigata d'Orbigny X Triloculina longissima n. sp X Triloculina oblonga (Montagu) X Triloculina quadrilateralis (d'Orbigny) X Triloculina tricarinata d'Orbigny MUiolinella circularis (Bornemann) Pyrgo bougainvlllei (d'Orbigny) X Pyrgo clypeata (d'Orbigny) X Pyrgo inornata (d'Orbigny) X Pyrgo lunula (d'Orbigny) X Pyrgo oblonga (d'Orbigny) X Pyrgo peruviana (d'Orbigny) X Pyrgo subspherica (d'Orbigny) X Prygoella globulus (Bornemann) MassUina decorata Cushman Massilina inaequalls d'Orbigny Hauerina sansebastianensis n. sp X SOKITIDAE Dendritina preelegans n. sp Peneroplis carinatus d'Orbigny X Peneroplis proteus d'Orbigny X Archaias aduncus (Fichtel & Moll) X Archaias angulatus (Fichtel & Moll) X Archaias compressus d'Orbigny X p 9> cr p o X X o c a -i O X X X X o c a* 9= X o D n X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X 278 SCIENTIFIC SURVEY OF PORTO RICO Table 2 {Continued) o o5' North Shore d •a ens o a •t South Shore rt aj T 5" (r' p o » e» CO <-»■ t-l* 9> a TnOCHAMMINlDAE Trochamminoidcs approximatus n. sp LiTUOLlDAE Cyclamniina acutidorsata (Hantken) Ataxophrarmiidae Valvulina oviedoiana d'Orbigny X Valvulanimina cornucopia n. sp X Vernt'uilina mexicana Nuttall Doiothia caribaea Cushman Dorothia cylindrica (Nuttall) Dorothia praelonga (Karrer) Gaudryina asiphonia Andrea Gaudryina glabrata maxima n. var Gaudryina karreriana Cushman Gaudryina puortoricana n. sp X Gautirvina (Pseudogaudryina) atlantica (Bailey) LiebuseJla byramensis (Cushman) Listerella cf. conmiunis (d'Orbigny) Clavulina tricarinata d'Orbigny X Calvulinoides polygonalis n. sp Clavulinoides triangularis (Nuttall) Textulariidae Textularia agglutinans d'Orbigny X Tfxtulaiia articulata d'Orbigny X Textularia broussardi ITowe & Wallace Textularia candeana d'Orbigny X Textularia gramen d'Orbigny X Textularia grenadana Iledberg Textularia indenta n. sp Textulariella l)arrettii (P. & J.) Vulvulina pachylieilus Iladloy Vulvulina i)ennatula (Batscli) Nodohahiidak Saracenaria arcuata (d'Orliigny) Astacolus insolitus (Schvvagor) Astacolus ovatus n. sp Astacolus sublituus (Nuttall) Memicristellaria fragraria (Giimbel) \ aginuliiia faba n. sp \'aginulina mexicana (Nuttall) Vaninuliiia siluiuoidea n. sp Margiimlina iiisulensis n. sp Margiiiuliiia subciassa Schwager Amiihicorj ne obliciua n. sp (il;nnhiliii;i coiiiatula (Cusliman) (ilanihiliiia gallowayi (Cusliman) (ilaiidulina mauricensis Howe & Roberts. . . . Dentaliiia advena (Cusliman) Deiilaliiia lialk.vardi Cusliman Dentaliiia liaJUxardi ixmceana n. var Dentalina liillaeformis n. sp Di'utalina multilineata BorncMiiann o cr o o 01 C X o c a u- 1-1 P a o 13 n a X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X GALLO^yAY AND HEMINWAY, TERTIARY FORAMINIFERA 279 Table 2 {Continued) North Shore South Shore <> r^ r f d t-^ o o « p- W ^ ^ •d a to 1 01 P 'Ji O o S S^2 (S' crq p o IJ5 K M O r O ►tJ P 13 p P o en CO w O 'U •-J CD CD e p G- (^ a P e+ t-d U p P CA P NoDGSARiiDAE — Cotitinucd Dentalina semUaevis Hantken Nodosaria halkyardi antillana P. & B Nodosaria longiscata d'Orbigny Nodosaria obhquata (Batsch) Nodosaria pariana Hedberg Nodosaria raphanistrum (Linne) Nodosaria scalaris (Batsch) Nodosaria simplex Silvestri Lagena ampulla n. sp X Lagena bullosa n. sp Lagena impressa n. sp Lagena nuttalli n. sp Lagena strumosa Reuss Robulus calliferus (Stache) Robulus chambersi Garrett '. Robulus cibaoensis n. sp Robulus convergens (Bornemann) Robulus falcifer (Stache) Robulus iota (Cushman) Robulus occiden talis torridus (Cushman) .... Robulus planulus n. sp Robulus plummerae Cole Robulus protuberans (Cushman) Robulus subpapillosus (Nuttall) Lingulina ponceana n. sp Lingulina semicostata n. sp Fissurina laevis Seguenza Fissurina marginata (Montagu) POLYMORPHINIDAE Polymorphina terquemiana (Fornasini) Guttulina basalis n. sp Apiopterina cylindroides (Roemer) Raphanulina gibba globosa (von Miinster) ... X NONIONIDAE Nonion chapapotense Cole X Nonion dilatatum n. sp Nonion multiporatum n. sp Nonion nicobarense Cushman Nonion pompUioides (Fichtel & Moll) Nonion subgrateloupi n. sp X Pseudononion papillatum n. sp. . . Nonionella modesta n. sp X Pullenia bulloides (d'Orbigny) Elphidium lanieri (d'Orbigny) Elphidium lens n. sp X Elphidium lobatum n. sp Elphidium nautiloideum n. sp X Elphidium owenianum (d'Orbigny) X Elphidium poeyanum (d'Orbigny) X Elphidium puertoricense n. sp X Elphidium sagrai (d'Orbigny) X Elphidium sagrai crassum n. var X Antillesina marielensis (Palmer) X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X 280 SCIENTIFIC SURVEY OF PORTO RICO Table 2 (Continued) North Shore South Shore 2 r ? « a ■d ^ S- -3 a a d p 2 ►1 •1 n^ o g 2 S CO c (R O o aq (W CO M O r' o ^ p p o c o s s ^ CA cc cc CO o *tJ (t tn c p o* IT CL p •-J on <^^ ^ p' i p X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X ACERVULINIDAE Rupertia verrucosa n. sp (^arpenteria bulloides n. sp Carpentaria proteiformis Goes Gvpsina discus (Goes) Sphaerogypsina globulus (Reuss) X X Sphaerogypsina pilaris (Brady) ASTERIGEHINIDAE Amphistegina angulata (Cushman) Amphistegina floridana Cushman & Ponton. . X X Chilostomellidae Chilostomella czizeki Reuss Cliilostomella globata n. sp Chilostomella ovoidea Reuss Chilostomella urceolus n. sp Sphaeroidina bulloides d'Orbigny Orbulinidae Globigerina bulloides d'Orbigny X X X Globigerina dutertrei d'Orbigny X X Globigerina inflata d'Orbigny Globigerina ouacliitaensis Howe & Wallace. . X (ilobigerina pachyderma (Ehrenberg) Globigerina pseudotriloba White X X (ilobigerina trilocularis d'Orbigny X X Orbullna universa d'Orbigny X X Pegidiidae Sphaeroidinella seminulina (Schwager) X Heterohelicidae Pavonina advena Cushman X Bolivina byramensis Cushman X Bolivina elongata Hantken Q Bolivina heineae n. sp ^ Bolivina jacksonensis Cushman & Applin. ... X Bolivina matanzana convexa n. var X X Bolivina me.xicana aliformls Cushman x Bolivina tectiformis Cushman Q Bolivina ventricosa n. sp ^ Loxostomum hiwanneense Howe x Lo.xostomum normale n. sp x Plectofrondicularia trinitatensis Cushman & Plectofrondicularia vaughani Cusiiman X BULIMINIDAE Reussella glabrata (Cushman) X X X X Bulimina socialis Borneniann x Cassidtjlinidae Cassidulina laevigata d'Orbigny XX Cassidulina subglobosa Brady Q Cassidulina tricamerata n. sp Q Ehrenbergina caribbea n. sp Q Ehrenbergina serrata gibbera n. var x 282 SCIENTIFIC SURVEY OF PORTO RICO Table 2 (Continued) UVIGERINIDAE Uvigerina bulbacea n. sp Uvigerina elongata Cole Uvigerina fusiformis n. sp Uvigerina gallowaj i Cushman Uvigerina gardnerae Cushman Uvigerina mantaensis Cushman & Edwards. . Uvigerina mexicana Nuttall Uvigerina mexicana bulhosa n. var Uvigerina mexicana ranunculus n. var Uvigerina postica n. sp Uvigerina vicksburgensis Cushman & Ellisor. Siphogenerina costostriata n. sp Siphogenerina cumingsi n. sp Siphogenerina hubhardi n. sp Siphogenerina mexicana Cushman Siphogenerina multicostata Cushman & Jarvis Angulogcrina cibaoensis n. sp Angulogerina cooperensis Cushman Angulogerina decorissima n. sp Angulogerina ponceana n. sp Trifarina bradyi Cushman Pleurostomellidae I'leurostomella bierigi Palmer & Bermudez. . . I'leurostomella elliptica n. sp I'leurostomella gerontica n. sp Nodosarella constricta granulifera n. var Nodosarella crassielegans Nuttall Nodosarella paucistriata Galloway & Morrey Nodosarella verneuili (d'Orbigny) Kllipsoglandulina exponens (Brady) Ellipsoidina ellipsoides abbroviata Seguenza.. Camerinidae Operculinella sinuata n. sp lleterostegina antillea Cushman X OuBITOIDIDAE Lepidocyclina cf. j)arvula Cushman X Lepidocydina (Nephrolepidina) cf. tournoueri L. & D X Miogypsinoides complanata (Schlumberger). . X North Shore South Shore > ci c f* d n o O .Q p. •o ^ ^ V a n (T> n> T 1 -^gt O o % gao « f" o O (W K f o r r) ^ p !-»• o c o D U" CO TJ O o o o o o B CD o n B o o D o B OOOQOO og ® a. R R Siphonina advena R R R Sphaerogypsina globulus C R C A C Amphistegina floridana R Glooigerina bulloides X X R Globigerina dutertrei R Globigerina pseudotriloba X R Globigerina trilocularis R R Orbulina universa R Bolivina jacksonensis R Bolivina matanzana convexa n. var. C Reussella glabrata R R Heterostegina antillea R Lepidocyclina cf. parvula R Lepidocyclina (Nephrolepidina) cf. tournoueri C R R A A C Miogypsinoides complanata X X o X X X X X ^ ^ 2 C1 ■d ^ o. -a 1-1 1-1 J- T (5 X XXX X X X X X X X XXX X X X XXX X X X X X X X X X X 7 X Table 4 (Cmitinued) LARES FAUNA Ijocality >■ C Pseudarcella patella n. sp. C Triloculina austriaca R Triloculina oblonga R Pyrgo clypeata C Pyrgo oblonga A Pyrgo subspherica R Elphidiuin lobatum n. sp R Elphidium poeyanum .... R Siphonina advena R Pavonina advena a o 2 2 2 5* f^ 3!< n o o § o a a n § ft n § o B B » f t> r; f y r^ f g r; o h^ T1 o W o •o s( p. n 3! C. •rt ^ D. ■o o a n (t c r> o C (t -t a n n (S 1 T X T X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 289 Table 4 {Continued) CIBAO FAUNA r r o o o o p p '<1 << > > l«^ re CO 05 p R R W o f T) ►ri V o M n K o o a f5 (D O a a> CD P P P CD a> (t p CD o !? M T) Tl o d T1 5! o. « s? a T) si & rt fC p. et IT & (T fD U fD *i a> •s X X 01 1 I X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X Spirillina vivipara Quinqueloculina akneriana R Quinqueloculina seniinulum R Pyrgo clypeata R Pyrgo oblonga R Pyrgo peruviana C Gaudryina glabrata maxima n. sp. . . C Textularia agglutinans R Textularia indenta n. sp R Textulariella barrettii C Vulvulina pachyheilus X X R Astacolus ovatus n. sp R Astacolus sublituus X X R Glandulina gallowayi X R Nodosaria halkyardi antillana X C Robulus chambersi XXXX R Robulus cibaoensis n. sp R Robulus convergens XXX X C Robulus planulus n. sp R Nonion dilatatum n. sp R Nonion multiporatum n. sp R R Nonion nicobarense XXX X X X R Nonion pompilioides X XXXXXXX A Pseudononion papillatum n. sp R C Elpliidium owenianuni X R Elpliidium sagrai X XXXXX R Valvulineria maclureaformls n. sp R Valvulineria paucilocula X R Eponides advena X X R R Eponides ornatissimus n. sp C Eponides pulvinus n. sp A Eponides ventricosus n. sp C Eponides vortex n. sp R Gyroidina stellifera n. sp R Discorbis oligospiratus n. sp X R Anomalina pompilioides n. sp R Cibicides americanus C Cibicides choctawensis A Cibicides floridanus R R Cibicides lobatus R Cibicides scalenus n. sp R Cibicides sinistralis C Planulina zigzag n. sp C C Siphonlna advena A Planorbulinella larvata C Sphaerogypsina globulus C Amphistegina floridana C R Globigerina bulloides X X R C Globigerina dutertrei C Globigerina ouachitaensis X C Globigerina pseudotriloba X A Globigerina trilocularis C Bolivina byramensis C Bolivina matanzana convexa n. var. . . R Loxostomum hiwanneense X R Loxostomum normale n. sp R Cassidulina laevigata XXX C Uvigerina bulbacea n. sp R Uvigerina gallowayi X C Uvigerina postica n. sp C Angulogerina cibaoensis n. sp R Angulogerina cooperensis C Nodosarella paucistriata C Nodosarella verneuili C Miogypsinoides complanata X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X 290 SCIENTIFIC SURVEY OF PORTO RICO Table 4 (Continued) LOS PUERTOS FAUNA o o p > CO c R R R R C C R R R R R R C R R C R R C R R C C R R A A C R C C R C R A A C A R R R O R A o o > «o R R R R R R R M O >> *TJ o t5 OQ O .5 ^ - fl rt 03 +iX2 cS cs c a 52£OOOOOL.bo3o3aj .3^^^^^3300t- --. tH >■.>■.>»>■, >iaj©t^tHi»ca,S tH piH (1^ PL, CL, PkflH Ah <5 <: > O O cS 3-3 ■03 u 3 '- „ tao 03 w O ftO Bo3g £ 3 §|g5il»iliP|iiiiillillisli llill?! .3 "S '^«3 S23 35«0.„- cS - IT? r. t, c«-e """ 03-3 .::§" csssg 3 3 3 3 3 —■ — — '-> ?^_^_5— M y -rt T-: Ti 7-1 33 3 3.: uXi-g 03 l>t- i2£ ^'3 3 3:i,a.S ^SX3.0-W ft3 3 0) o O 3 c^ O 03 O o A 93 A15 tf tf A64 A20 A21 F358 F358(2) F359 A46 F64 «« « M « « « P? «« « o «« p? PhO PS 292 SCIENTIFIC SURVEY OF PORTO RICO s s o w n < Recent X X XXX X XXX XXX XX Pleistocene XX XX Pliocene Upper X X X XX X XX X Miocene Middle XX xxxx XXX X XX X Lower X X XX X X X xxxx X Upper X XX X X X X XX Oligocene Middle X X X XX X X X Lower X XX X XX xxxx X X Upper X XX X X X X X Eocene Middle Lower X X X a fa iJ Q < K W a |»:3 3 . h t- cs t. 2 C.2 C t- S O u 5? c cs > ca t; 2 = 3 c« ^ •-;— O.^ i— r- M rt ca 3 , i^ l; fi _:3 aas: >-= o<^ _3QSia^aj22 — ^ ^ ^ ^ •!? ■? cj ei ao o o o o 3ft a"' O ce 1) c. ^ii n ft a a-- •« c 3 ooc C — :S a S 5 o o a V. s o °« 3 £ « ^a3 .5 .5 -2 C C C aS-n° »3 C r; re a-5 .5 c P X •/: O C w 111! 3 3 B't — ~ C w c« :« sj .•« C C C C o. n 9J Cd fi- ^ ™ j^ .— - •'« !: ~ X 3= -^ T 3j 3:= aai si :i ci '■^- a. jS aaj*. ajja ;: ^ SI at it sc c_c_o_o - - 3 - 5 ^ i O 3 r— i X— ci 3 -J X 3 t. _2— 3 SX O u a ^ 3 a o A93 A 15 AG4 i1« P4 o A20 o O OOtf d; 33« « A21 o<; «« o opsoh «*;<;* « F61 « K GALLOWAY AND HEM I NW AY, TERTIARY FORAMINIFERA 293 s s o O n < Recent XXXX X XX XXXXXXXX XXXXX X X XX Pleistocene X X X X X X Pliocene X X XX X X X X Upper X X X X X X X XX Miocene Middle Lower Upper XX X X XX X XX X X X X XX X X XXX XX Oligocene Middle X X X X X X X Lower X XXX X X XXX X XX XX Upper X X X X Eocene Middle Lower X • < < w Q o u d "3 O; * Co u Ui d c« as S-?8 S d cS eS C C (3 _ fi w 3 f^ «« iSos w = tt>.S ■^ t- C * d +i 355 c o n Its O 3 S 3 gggo a's 3'E d =i TO 13 d _ d d d d cS d .S .9 .S -3.9.9 3 3S333 o o o o o o O O O O O O 't. 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U !-i ^ b ( >i >. >* >• 3.9 3 O 03 3 "^"^ O u d i- e w (C O SS C 3 o 3 2 £ S =* _ =* 2 ■ft o «*3g .2gSd.S§S-c ® o'S Si S.3 oj M ^ .^^ .1—1 -f^ .^H t^ ,^ 35 is 3 jn J3 — "^"^ Jl '' r t- t. fc. 3 3 ■d cs ®555 ? ? 3 ?."« be m 3 O Dc •C l* 3 d.2.2 3 bD oj a C d 'E''« O CO d O .9.9 3 3 > > Co o P433 P434 A432 P432 0505 P431 P259 P258 P255 P254 P253 P2 K P3 O P4 P251 tf « o «« «o p; « p; tf « « 294 •.SI o H -J m < Recent Pleistocene Pliocene Miocene Oligocene Eocene SCIENTIFIC SURVEY OF PORTO RICO XXX Upper Middle Lower Upper Middle Lower Upper X Middle Lower X X XXX X XX X XX X XX X X X xxxxx X X X X X X X X X XX X X X X X X X X X •X X X XXX X X X X X xxxx X X XX XXX XX =^ X X X X X XX XXXX XX X 6 < w a o O C Oj'^ u rt t, J £ O SU," rt c^ c^ ro 3 3 3 3 +^ 4J +i> -M OJ O) © "li >,o3 3 " o ^ aftcs 3 0"* ■5'g 0) r ? u ^> cd ,3 3 cS 3 3 OO <«5 OS Sis C w c^ cd C^ .s.s.s 3 3 3 c a a 'SL'S'El cd ctS c<3 2 =« !I s^ii 01 cd cd C '^ C fi >-.cd 3 3g=:: c^ ct C rt >^ CT cd Cd'*' C cd cd ■-.s.s •c'cd'3 Sen J» a, oj a o . o ^2 go. o3 a c^ :€ c^ cti 3 3 3 3 s '■" 5.9 =* '3. CS h 0.cd o-£ 5 "i " x a X .c 5(3 03 »=dSg 3 2* iZ- O t- te3: ft cdo3edcdcdcd3£ u t- u ;^ ^ I cd cd cd cd cd < « «3 JJ X X O O O O O I 00000 0030'2'5'5'2'2'S'^'* ai»®a;OOOOOOjj£^ Cd 3 S .„ X t.o = 2.2; 3 3i3So •=« XX cd 03 3 3 "^ 1/ S s 03 cd 05 cS o o H-1 P433 P434 A432 P432 P431 P259 P258 P255 P254 P253 P2 P3 P4 [ P251 « « « ««« K « « « « «« GALLOWAY AND HEMINWAY, TERTIARY FORAMJNIFERA 295 o n < Recent XXX xxxx xxxx X X Pleistocene XXX XX X X Pliocene XXX XX X X Upper X X X X X X X Miocene Middle XX X XXX XX X X X X X Lower X X X X X X X Upper X X X X XX Oligocene Middle X X X X X X X Lower XX XX XXX X X X XXX XX X Upper X X XX x'- Eocene Middle Lower XX X s 6 < < w D O CO a . .ii * fl m M * rt ® 1; ^'^ te ft _ - w <* O ■ SSi U.^ O ftft cc W W W EB « C/) 3 3 3 3 3 3„„ S 2 !« 5.9 o 3 3 3 3 3 3 3 3 ri.rl3 3 oooooo9"JS <4 u, ® 0) 3 oi S 3 .5 i^ to 0.5 S o oi » '- 3.rt O 3^ c«'E CI ft to .92Gfle-2 t- o o o o « ^ 3 Qi 3C S ,„ 03 g ©.„.„„-■ - o3 > g 3 c3 t» ^ W (D ^ s^ O 3 on (=" a* g> CO « g « S£ 3 3 c« rt 3 aa SoaatotcC^rtcsce 333CCS!*C'^-- 3 3 3 3 3 ^-Sy-S-S-S ft ft "^ to . p, . 3 to t/j - '-' 3 3 w:- Oi tor; 0--J 03 to -3 y-S a, o a^ ^ CO 3 (A ■d CO a a> o a> a cSto g'^ft '"ce CO 2o3c«« . O O Lh en <«'ao o 03 'C'CC'C'CC a^ cS u o P433 P434 A432 P432 P431 P259 P258 P255 00 « P254 O « P253 P2 «0 « «« P3 00 «tf P4 P251 ft a ft ft ft ft^ ^ > > cO c^ c^ >» to CO to to ^ dj 0.1 1- ^ *^ .ii.ii Hi 3 Qi rtj rtj O , ^ ^ Ji ^ Xj'C'o'r^ c^cSctc^c^ OC5aocSc3o3o3cS O O O O t--^*i4J-w-*^ ftaaa>. 00000 « < o o o o ooo« « « P5 « 296 SCIENTIFIC SURVEY OF PORTO RICO Recent XX XX X XXX X XXXX X Pleistocene X X XX X Pliocene X X XX Upper X X X X Miocene Middle X XX X X XXX X X X Lower X XX X X XX XX Upper X X X X X X X X XX X Oligocene Middle X X X X X Lower X XX XXXX XXXX X X XXXX Upper Middle XX xxxx XXX X X XX Eocene X XX Lower X X XX n < a o o Oh 3 CI d 3 o o o o w to QQ « C cS a cs « ftjS 3 CtJ w cfl 3 ft to »]-3 . C ^ o rf S S a^ - c O J- 03 3 ee ™ 3 3- c3 tcft P=o fS O^^ — r >53i;ScC^^ — 33_3ajx— its '* n u. a3.i:.t; a :i ^ ^ ^ ^ ^j:-C.5 C Gft'r c e ^Z-c >>>-'?''-^p^ p p~ c c a c G c c^- fli ^^ ^r ^ 1^ G H £ C^ ^ .,-. -^ -^ •« — t — .t; 3 3 3 o .2 rt -M 3f cs .25 ft ft.: =* 03 "„ 3 C ■ ■ ^ 3^ i* w-f, 3y^» y-S'S'""S ^ ° ° ..— — .— .*- c^ c^ J^ ° ° i; ftS S S ftftO o o o SZS232 -■? > '- o o _o _o _o o "o •; P433 P434 A432 P432 P431 P259 P258 P255 P254 P253 P2 P3 P4 P251 O O «tf o P4 O O P5 O cS .^ ^ • of? flisSt- 5 o c ^ CS c« .2 2 i2 cs c« c« i ^ !* fl c r X X XXX xxxx X XX XX X'^X X XXX X X ftcS 1 cS O '°*^ c c g-S =3 rt a3 2 a; o g 2 g g ^£ 2 =« CO CO---- 3 a CO CO 03 tri Lri .S.S.S'So P433 P434 A432 P432 P431 P259 P258 Ph P255tf tf O «« « P254« « tf ««0 «««« P253 P2 0« tf <«««« « ««« P3 O «« OP5« tf (^« P4 P251 „ 3 3— ■ S-SS 2.2.2 ''oaSo ^ D W) iS " S (S "v-,!^ ,, ~^So32S3 "^^ ^ 03 cS 03 O !. 5 — 3? ^ ^i; |'^gci303c«'gcS.S5o o o o OJ ? *^s ^,>,-;j ^<^+i ;- U I 3 3 3'O'C'C.S.SnS.O O) 0; 1; O O 03:1; © 05 P50 «« « O Ph OOPSPh OOP5P3 Ofi< p^op; «« « OP? « P3 P3 o« 298 SCIENTIFIC SURVEY OF PORTO RICO ACKNOWLEDGMENTS We wish at this time to acknowledge our great indebtedness to Professor H. A. Meyerhoff, who, by his careful collection of material from the many localities enumerated above, has made possible the prosecution of this work. To him we owe, also, our knowledge of the relative stratigraphic position of the various collecting localities. The figures were drawn by the senior author, J. J. Galloway. Valuable assistance in shading the figures was furnished by Frances Beede, Frederick P. Schweers, and J. Bernard Martindale. Drafting of the Porto Rican locality map was done by Frederick P. Schweers. We are grateful for the substantial grants from the research funds of Smith College and Indiana University for assistance in making the drawings. All of the types are catalogued and deposited in the Paleontological laboratory at Indiana University. SYSTEMATIC DESCRIPTION OF SPECIES Family LAGYNIDAE Schultze 1854 PSEUDARCELLA Spandel 1909 Pseudarcella patella NEW species Plate 1, figures la, b Test unilocular, discoidal, concavo-convex; edge narrowly rounded; dorsal side convex with a small, inconspicuous round knob in the center of the dorsal surface; wall calcareous, readily and completely soluble in dilute acid, hyaline in appearance, pores not observable; wall material not chitinous or arenaceous but calcareous with pre- cisely the same appearance as specimens of hyaline, finely perforate forms such as the Nodosariidae and the Polymorphinidae; aperture in the center of a circular depression on the concave side of the test, circular in shape. Diameter, from 0.25 to 0.55 millimeter; maximum height, 0.16 millimeter. Holotype No. 3950, locality A6, Lares formation. Common at A6 of the Lares formation, from which half a dozen specimens were obtained. P. patella is larger in diameter and thinner than P. arcnata Cush- man (1930. Fla. Geol. Surv. Bull. 4). GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 299 Family ASTRORHIZIDAE Brady 1881 RHABDAMMINA M. Sars 1869 Rhabdammina irregularis Carpenter Plate 1, figure 2 Rhabdammina irregularis Carpenter (1869) Proc. Roy. Soc. London 18: 60. — Cushman (1918) Bull. U. S. Nat. Mus. 104 (1) : 17. pi. 8, fig. 1. Recent, Bay of Biscay, Pacific Ocean along western coast of America as far nortli as Gulf of California, East Indies. Rhabdammina abyssorum Brady (1884) Rep. Voy. Challenger Zool. 9: 268. pi. 21, fig. 9 only. Recent, northwest of Ireland. Test unilocular, consisting of a dichotomously or irregularly branch- ing tubular chamber of nearly uniform diameter; wall of firmly cemented sand grains, exterior rather rough, interior more smoothly finished; ends of the tube serving as apertures. Average diameter, 0.83 millimeter; length of figured specimen, about 4 millimeters. Plesiotype No. 3972, locality P3, Ponce formation. Common at P3 ; rare at P2 of the Ponce formation. Family SPIRILLINIDAE Reuss 1861 SPIRILLINA Ehrenberg 1843 Spirillina vivipara Ehrenberg Plate 1, figures 3a, b Spirillina vivipara Ehrenberg (1841) Abh. k. Akad. Wiss. Berlin 422. pi. 3, sec. 7, fig. 41. Recent, off coast of Mexico. — Brady (1884) Rep. Voy. Challenger Zool. 9: 630. pi. 85, figs. 1-5. Recent, world wide.— Cushman (1915) Bull. U. S. Nat. Mus. 71 (5): 3. pi. 1, figs. 1, 2. Recent, north Pacific; (1931) Bull. U. S. Nat. Mus. 104 (8): 3. pi. 1. figs. 1-4. Recent, West Indies. Test discoidal, slightly biconcave, planispiral, composed of a single tubular chamber in several coils, slowly increasing in diameter and • only slightly involute; coils flattened and both sides of test some- what flattened ; periphery rounded ; spiral suture somewhat depressed ; aperture simple at the end of the tubular chamber. Diameter, 0.27 by 0.30 millimeter; thickness, 0.04 millimeter. Plesiotype No. 3993, locality A86, Cibao formation. Rare at A86 of the Cibao formation and at P432 of the Ponce formation. PATELLINA Williamson 1858 Patellina corrugata Williamson Pl.ATE 10, FIGURES 5a-c Patellina corrugata Williamson (1858) Rec. Foram. Great Britain 46. pi. 3. figs. 86-89. Recent, off Great Britain.— Cushman (1930) Contr. Cushman Lab. Foram. Res. 6: 15 pi. 3. fig. 5. Tertiary to Recent; (1931) BulL U. S. Nat. Mus. 104 (7) : 11. pi. 2, figs. 6. 7. Recent, east and west borders of North Atlantic. 300 SCIENTIFIC SURVEY OF PORTO RICO Test conical, apical angle nearly 90 degrees; ventral side nearly- flat; several whorls of chambers visible on the dorsal side, early tubular chamber obscure but apparently comprising 2 or 3 whorls; later chambers shortening to 2 to a whorl; lobate ornamentation on dorsal side of the chambers, usually attributed to presence of second- ary septa or septula, lobes sometimes simple, but more frequently double near the outer edge of the chamber ; lower side with secondary deposits in knobs and meandering lines; wall thin, translucent, not definitely perforate; aperture a slit on the ventral side between the last 2 chambers with valvular lip. Diameter, 0.31 millimeter; height, 0.16 millimeter. Plesiotype No. 3944, locality P432, Ponce formation. Rare at P432 of the Ponce formation. P. advena Cushman (1922. U. S. Geol. Surv. Prof. Pap. 129F) is closely related to P. corrugata but differs, as stated in the original description, in the much finer division by internal septa and in a lower spire which makes a broad flaring test. Family MILIOLIDAE d'Orbigny 1839 SPIROLOCULINA d'Orbigny 1826 Spiroloculina antillarum d'Orbigny Plate 1, figures 6a-c Spiroloculina antiUarum d'Orbigny (1839) in De la Sagra, Hist. Pliys. Pol. Nat. Cuba Fora- miniffires 166. pi. 9. flgs. 3. 4. Recent, off Cuba. — Brady (1884) Rep. Voy. Challenger Zool. 9: 155. pi. 10. flgs. 21a, b. Recent, south Atlantic. — Cushman (1921) Proc. U. S. Nat. Mus. 59: 63. pi. 14. flgs. 14, 15. Recent, off Jamaica; (1932) Bull. U. S. Nat. Mus. 161 (1): 36. pi. 9, flgs. 3-5. Recent, tropical Pacific. Test oval in side view, biconcave, narrowly oval in end view; chambers subcircular in transverse section, rather thick; surface marked by numerous longitudinal striae; aperture terminal at the end of a neck with a plate-like tooth. Length, 1.07 millimeters; width, 0.63 millimeter; thickness, 0.22 millimeter. Plesiotype No. 3994, locality A79, Los Puertos formation. Rare at F64 of the Quebradillas formation, and P258 of the Ponce formation. Common at A79 of the Los Puertos formation. Spiroloculina elongata d'Orl)igny Pl.\TE 1, FIOURE.S 5a-c Spiroloculina elongata cVOrhifsjty (1826) Ann. Sci. Nat. 7:298, No. 11. Pliocene. Italy.— Forna- sini (1904) Mem. R. Accad. Sci. Istit. Bologna Sez. Sci. Nat. VI. 1: 5. pi. 1, flgs. lOa-c. (After d'Orbigny 's unpublished flgurc of 1826.) GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 301 Test elongate, elliptical in side view; biconcave; back of the test broad, gently rounded; chambers elongate, with rounded back and concave sides in transverse section, tapering from the broadly rounded peripheral back of the chamber toward the center of the test, with an angled or nearly carinate contact between the back and sides of the chambers; aperture round, with plate-like tooth, at the end of a neck. Length, 1.41 millimeters; width, 0.75 millimeter; maximum thickness, 0.61 millimeter. Plesiotype No. 3995, locality A79, Los Puertos formation. Rare at A79 of the Los Puertos formation. Spiroloculina oculina NEW species Plate 1, figures 4a-c Test regularly oval in side view; oval in end view, sides slightly concave, with carinae at the junction between back and sides of the chambers; chambers closely appressed, 5 visible on one side, 6 on the other side of the test, with portions of the keeled edge of earlier chambers visible on both sides; aperture terminal, round, with a short neck and T-shaped tooth. Length, 1.52 millimeters; width, 0.85 millimeter; thickness, 0.41 millimeter. Holotype No. 3996, locality A79, Los Puertos formation. Very rare at A79 of the Los Puertos formation. S. oculina has fewer chambers and is thicker than *S. excavata d'Orbigny from the Vienna Miocene. QUINQUELOCULINA d'Orbigny 1826 Quinqueloculina akneriana d'Orbigny Plate 2, figures la-c Quinqueloculina akneriana d'Orbigny (1846) Foram. Foss. Vienne 290. pi. 18, figs. 16-21. Middle Miocene, Vienna. Quinqueloculina seminulum Cusliman (1918) Bull. U. S. Nat. Mus. 103: 78. pi. 28, figs, la-c (not figs. 2, 3). Middle Miocene, Panama Canal Zone. Test elongate, ovate in side view, width % length; % as thick as wide; edge rounded; chambers distinct, of uniform diameter, nearly circular in cross section, successive chambers enlarging regularly in size; sutures depressed; apertural end without a definite neck; aper- ture terminal, circular, with a simple tooth. Length, 0.68 millimeter; width, 0.47 millimeter; thickness, 0.33 millimeter. Plesiotype No. 3961, locality L6C, San Sebastian formation. Rare at L2C; common at L6C of the San Sebastian formation. Rare at A86 of the Cibao formation. 302 SCIENTIFIC SURVEY OF PORTO RICO We are following the generally accepted usage of restricting the name Q. semmulum (Linne) to those short, ovate quinqueloculines with subangular edge. Quinqueloculina apiculata NEW species Plate 2, figures 2a-c Test about twice as long as wide, % as thick as wide; periphery rounded; chambers distinct, lengthening rapidly, somewhat inflated, distal end of last chamber with a short blunt apicule ; sutures distinct, moderately depressed; surface of holotype poorly preserved; aper- tural end of last chamber extended into a neck; aperture terminal, circular, with a narrow, simple tooth. Length, 0.49 millimeter; width, 0.23 millimeter; thickness, 0.14 millimeter. Holotype No. 3962, locality L6C, San Sebastian formation. Very rare at L6C of the San Sebastian formation. This species has resemblance to Q. laevigata, but has a definite, distinctive neck. Quinqueloculina audacula NEW SPECIES Plate 1, figures 7a-c Test somewhat longer than wide; chambers polygonal in transverse section, with conspicuous angles or carinae; back flat or slightly con- cave between the carinae; sutures distinct, slightly depressed; aper- ture round, with simple tooth, at the end of a short, round neck. Length, L60 millimeters; width, LOO millimeter; thickness, 0.67 milli- meter. Holotype No. 3963, locality A79, Los Pucrtos formation. Rare at A79 of the Los Puertos formation. This species has more pronounced peripheral carinae and is less regularly oval in side view than Q. badenensis d'Orbigny from the Vienna Miocene. Quinqueloculina laevigata (d'Orbigny) Plate 1, figures 8a-c Quinqueloculina luevioalu d'Orbigny (1826) Ann. Sci. Nat. 7: 301. No. 6. Middle Eocene, Paris Basin; in Barker, Webl). & Berthelot (1839) Hist. Nat. Isles Canaries 2 (2): Fora- minif6r«!s 143. pi. 3, ilgs. 31-33. Itecent, Canaries.— Cushman (1922) Carnegie Inst. Wash. I'ubl. 311: 05. pi. 13. flg. 2. Recent. Tortugas; (1935) U. S. Geol. Surv. Prof. Pap. 181: 11. pi. 2, ngs. 13-15. Upper Eocene. Mississippi. Test elongate, subovate in side view, about 21/2 times as long as wide, % as thick as wide; periphery rounded; chambers distinct, somewhat inflated, enlarging gradually; sutures depressed; surface GALLOWAY AND HEM IN WAY, TERTIARY FORAMINIFERA 303 poorly preserved; apertiiral end slightly extended; aperture terminal, circular, with a narrow, simple tooth. Length, 0.81 millimeter; width, 0.38 millimeter; thickness, 0.25 millimeter. Plesiotype No. 3964, locality L6C, San Sebastian formation. Common at L6C, LlC, and L2C; rare at L4C and L5C of the San Sebastian formation. Although named by d'Orbigny in 1826, this species was a nomen nudum until 1839, when it was figured by d'Orbigny in his paper on the Foraminifera of the Canaries. In 1905, d'Orbigny 's original figure of the species was published by Fornasini. This figure differs from that of Q. laevigata in the Canaries paper, but the earlier pub- lished figure, that of the Canaries paper, is necessarily taken as the type figure. Quinqueloculina maculata new species Plate 2, figures 3a-c Test oval in side view, subquadrate in end view, length nearly twice breadth, thickness % breadth; periphery angled; chambers distinct, subtriangular in cross section; sutures only slightly depressed; wall of all chambers finely punctate ; apertural end not produced ; aperture oval. Length, 1.03 millimeters; width, 0.63 millimeter; thickness, 0.45 millimeter. Holotype No. 3965, locality L6C, San Sebastian formation. Rare at L4C; common at L6C of the San Sebastian formation. This species is distinctive in its ornamentation. It much resembles Massilina jacksoncnsis Cushman (1933. Contr. Cushman Lab. Foram. Res. 9) but it has no spiroloculine stage and, in any case, is less compressed. Q. jamaicensis Cushman and Jarvis which was described from the Eocene of Jamaica (1931. Contr. Cushman Lab. Foram. Res.) is more compressed and has pits arranged in definite nearly longitudinal lines. Quinqueloculina philippi Reuss Plate 2, figures 4a-d Quinqueloculina philippi Reuss (1856) Sitz. k. Akad. Wiss. Wien 18: 252. pi. 9. figs. 87a-d. Upper Oligocene, Germany. Test very slender and elongate, compressed, narrowly oval in end view; edge angled with the suggestion of a keel; length nearly three times the width; width more than twice the thickness; chambers tubular, of fairly uniform diameter; aperture terminal, round, at the end of a definite neck, tooth not seen. Length, 0.73 millimeter; width, 0.26 millimeter; thickness, 0.11 millimeter. 304 SCIENTIFIC SURVEY OF PORTO RICO Plesiotype No. 3966, locality A79, Los Puertos formation. Very rare at A79 and A91a of the Los Puertos formation. This species is close to Q. angustissima which was described by Reuss from the middle Miocene and which occurs also with Q. philippi in the upper Oligocene. It differs from Q. angustissima in having a distinctly angled edge. The Porto Rican form is slightly more slender than the original figure of Q. philippi. Quinqueloculina ponceana NEW SPECIES Plate 2, figures 5a-c Quinqueloculina angusta Galloway & Morrey (1929) Bull, Am. Paleont. 15 (55): S. pi. 3, flgs. 3a. b. Probably lower Oligocene, Ecuador. Test small, compressed, elliptical in side view; edge sharply rounded; both apical and apertural ends slightly produced; aperture at end of a short neck, round with a small, simple tooth. Length, 0.42 milli- meter; width, 0.21 millimeter; thickness, 0.11 millimeter. Plesiotype No. 3967, locality P2, Ponce formation. Very rare at P2 and P254 of the Ponce formation. Q. ponceana differs from Q. anguina Terquem (1878. Mem. Soc. Geol. France III. 1) in that it is shorter, it lacks the conspicuous, rather long neck, and the edge is more sharply rounded. Quinqueloculina pygmaea Reuss Pl.\TE 2, FIGURES 7a-c Quinqueloculina pygmaea Reuss (18.59) Denksch. k. Akad. Wiss. Wien 1: 384. pi. 50, flg. 3. Middle Miocene, Austria. Miliolina pygmaea Brady (1884) Rep. Voy. Challenger Zool. 9: 165. pi. 113. flgs. IGa, b. Recent, south Pacific and west coast of South America. Quinqueloculina seminulum Cushman (1930) Fla. Geol. Siu-v. Bull. 4: 19. pi. 2, flgs. 2a-c (not fig. 1). Middle and upper Miocene, Florida. Test very slender and elongate, oval in end view; three times as long as wide; thickness two-thirds of the width; ijorijihcry rounded; chambers distinct, tubular, of uniform diameter; sutures depressed; apertural end with a slight neck; aperture terminal, circular with a simple tooth. Length, 0.54 millimeter; width, 0.18 millimeter; thick- ness, 0.12 millimeter. Plesiotype No. 3968, locality L6C, San Sebastian formation. Rare at L6C of the San Sebastian formation. Common at A79 of the Los Puertos formation. Rare at F64 of the Quebradillas for- mation. GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 305 Quinqueloculina setninulum (Linne) Plate 2, figures 8a-c Serpula setninulum Linne (1768) Sys. Nat. ed. 10. 786. Recent. Adriatic Sea. Quinqueloculina seminulum d'Orbigny (1826) Ann. Sys. Nat. 7: 303. no. 44. Recent, north Atlantic coast of Europe, Adriatic Sea, Mediterranean (.Pliocene, Italy. — Cusliman (1929) Bull. U. S. Nat. Mus. 104 (6) : 24. pi. 2, figs. 1. 2. Recent, northeast coast of North America; (1929) Contr. Cushman Lab. Foram. Res. 5: 60. pi. 9, figs. 16-18; (1933) U. S. Geol. Surv. Prof. Pap. 175A: 9. pi. 2, flgs. 2a-c. Middle and upper Miocene, Atlantic coastal plain. Test short ovate in si(ie view; in end view subovate, edge narrowly rounded to subangular; chambers distinct, of uniform diameter, subtriangular in cross section, successive chambers enlarging regularly in size; sutures distinct, very little depressed; apertural end without a definite neck; aperture terminal, oval, with a simple narrow tooth. Length, 0.90 millimeter; width, 0.67 millimeter; thickness, 0.42 milli- meter. Plesiotype No. 3969, locality L6C, San Sebastian formation. Rare at L6C of the San Sebastian formation, and at A43a of the Cibao formation. Quinqueloculina vulgaris d'Orbigny Plate 2, figures 6a-c Quinqueloculina vulgaris d'Orbigny (1826) Ann. Sci. Nat. 7: 302. no. 33. Recent, Mediter- ranean, Adriatic. Antilles.— Fornasini (1902) Mem. R. Accad. Sci. Istit. Bologna Sez. Sci. Nat. V. 10: 21. fig. 13. (d'Orbigny's unpublished figure of Recent, Adriatic Sea speci- men.)— Cushman (1922) U. S. Geol. Surv. Prof. Pap. 129F: 142. pi. 32, figs. 9, 10. Lower Oligocene. Mississippi; (1929) Bull. U. S. Nat. Mus. 104 (6) : 25. pi. 2. fig. 3a-c. Recent, eastern Atlantic. Test small for the genus, short and rather stout, subcircular in side view, rounded triangular in end view; edge narrowly rounded; chambers distinct, subtriangular in cross section, successive chambers increasing rather rapidly in size; sutures distinct, depressed; aper- tural end not produced; aperture oval, elongate parallel to the plane of coiling of the chamber, with a simple tooth (bifid in d'Orbigny's figure in Fornasini). Length of figured specimen, 0.29 millimeter; width, 0.25 millimeter; thickness, 0.11 millimeter. Average specimen length, 0.56 millimeter; width, 0.43 millimeter; thickness, 0.19 milli- meter. Plesiotype No. 3970, locality L6C, San Sebastian formation. Rare at LlC, L4C, and L6C of the San Sebastian formation. Com- mon at A79 of the Los Puertos formation. Rare at A15, A93, and F359 of the Quebradillas formation. Rare at P259; common at P258 and P431 of the Ponce formation. 300 SCIENTIFIC SURVEY OF PORTO RICO TRILOCULINA d'0rbign3' 1826 Triloculina austriaca dOrbigny Plate 3, figures la-c Triloculina austriaca d'Orbigny (184G) Forani. Fo&s. Vienno 275. pi. IG, figs. 25-27. Middle Miocene, Vienna. Test irregularly oval in front view, subtriangular in apertural view ; chambers unevenly convex, somewhat narrower toward apertural end of each chamber so that only the lower part and one side of the last formed chamber are visible in front view; sutures deep; aperture sub- circular with a broad bifid tooth. Average specimen length, 0.83 millimeter; width, 0.58 millimeter; thickness, 0.70 millimeter. Plesiotype No. 4006, locality L6C, San Sebastian formation. Rare at LlC, L2C, L4C; common at L6C of the San Sebastian formation. Common at A6 of the Lares formation. Rare at F64 of the Quebradillas formation, and at P431 of the Ponce formation. Triloculina brongniartiana d'Orbigny Plate 3, figures 2a-c Triloculina bronyniarliunu d'Orbigny (1839) in Dc la Sagra, Hist. IMiys. I'ol. iSat. Cuba Foraniini feres 152. pi. 0, figs. 5-7. Recent, ofT Cuba. Test rather heavy in appearance, oval in side view, with the last chamber projecting apically beyond the earlier chambers; little com- pressed in end view; chambers distinct, marked by numerous fine longitudinal striae which extend up onto the neck, about 7 striae on each chamber on each side of the test; aperture at the end of a distinct neck, round with simple, narrow tooth. Length, 1.05 milli- meters; width, 0.75 millimeter; thickness, 0.60 millimeter. Plesiotype No. 4007, locality P258, Ponce formation. Common at P258 of the Ponce formation. Bermudez (1935) reports that this species is common in the Recent off northern Cuba but unfortunately gave no figure. Triloculina cervicula NEW SPECIES Plate 3, figures 3a-c Test elongate, si)indle shaped in side view; chambers smoothly rounded in end view; sutures depressed; surface smooth; aperture terminal, at the end of a slender neck, circular, with a simple tooth. Length, 0.43 millimeter; width, 0.21 millimeter; thickness, 0.16 milli- meter. Holotype No. 4008, locality L6C, San Sebastian formation. GALLOWAY AND IJEMJNWAY, TERTIARY FORAMINIFERA 307 Very rare at L6C of the San Sebastian formation. This species differs from T. gracilis in its greater width and from T. oblonga in the presence of a conspicuous neck. This species is not as slenderly proportioned as T. gracilis which was described by d'Orbigny from the Recent of Cuba. The presence of a conspicuous neck distinguishes it from T. oblonga (Montagu) . The specific name cervicula means "a small neck." Triloculina elliptica NEW species Plate 2, figubes 9a-d Test elongate, slightly tapering, subovate in side view with a slight neck, subtriangular in end view ; chambers elongate, with width greater than the thickness; sutures slightly depressed; aperture terminal, nearly circular with lip and small simple tooth. Length, 0.52 milli- meter; diameter, 0.22 by 0.23 millimeter. Holotype No. 4009, locality LlC, San Sebastian formation. Rare at LlC of the San Sebastian formation. Triloculina gibba dOrbigny Plate 3, figures 4a-c Triloculina gibba d'Orbigny (1826) Ann. Sci. Nat. 7: 299. no. 3. Recent, Adriatic near Rimini; Pliocene, Ca&tel-Arquato ; (1846) Foram. Foss. Vienne 274. pi. 16, figs. 22-24. Middle Miocene, Vienna. Test ovate in front and side views, approximately a rounded equi- lateral triangle in end view ; chambers curved and symmetrical toward both ends, last chamber wider than preceding chambers so that in front view the last chamber is visible on all sides of the two earlier chambers; outside of chambers flattened; chamber angles rounded; sutures distinct, depressed; surface smooth; aperture crescentic, ter- minal, without a neck. Average length, 0.74 millimeter; breadth, 0.49 millimeter; thickness, 0.53 millimeter. Plesiotype No. 4010, locality L4C, San Sebastian formation. Rare at LlC, L3C, L4C; common at L2C; and abundant at L6C of the San Sebastian formation. Rare at A79 of the Los Puertos formation, at A21 of the Quebradillas formation, and at P255 of the Ponce formation. Triloculina inflata d'Orbigny Plate 3, figures 5a-c Triloculina inflata d'Orbigny (1846) Foram. Foss. Vienne 279. pi. 17, figs. 13-15. Middle Miocene, Vienna. 308 SCIENTIFIC SURVEY OF PORTO RICO Test somewhat elongate, subovate in side and end views; chambers tapering a little toward the apertural end, with thickness and width nearly equal; sutures distinct, somewhat depressed; aperture terminal without definite neck or lip; nearly circular with a simple tooth. Length, 0.49 millimeter; width, 0.29 millimeter; thickness, 0.21 milli- meter. Plesiotype No. 4011, locality L3C, San Sebastian formation. Very rare at L2C of the San Sebastian formation. Rare at A79 of the Los Puertos formation, at F358 and F359 of the Quebradillas formation, and at P431 of the Ponce formation. Triloculina laevigata d'Orbigny Pl.\TE 3, FIGURES 6a-c Triloculina laevigata d'Orbigny (1826) Ann. Sci. Nat. 7: 300. no. 15. Recent, Mediterranean. — Schlumberger (1893) Bull. Zool. Soc. Fiance 18: 63. pi. 1. flgs. 45-47. Recent, Gull of Marseille. — Fomasini (1905) Mem. R. Accad. Sci. Istit. Bologna Sez. Sci. Nat. VI. 2: 61. pi. 1, fig. 10 (from d'Orbigny 's unpublished figure). Test elongate, subovate in side view, roundly subovate in end view; chambers elongate, tapering toward the proximal end, with width and thickness nearly equal; sutures distinct, depressed; aper- ture terminal, without a neck, semicircular in shape with a simple tooth. Length, 0.95 millimeter; width, 0.50 millimeter; thickness, 0.33 millimeter. Plesiotype No. 4013, locality L4C, San Sebastian formation. Rare at L2C and L4C of the San Sebastian formation, and at A79 of the Los Puertos formation. Very rare at P251 of the Ponce for- mation. Triloculina longissima new SPECIES Plate 3, figures 7a-c ? Triloculina oblonya Cushman (not Montagu) (1922) U. S. Geol. Surv. Prof. Pap. 129E: 104. pi. 28, flgs. 3, 4. Middle Oligocene, Byram marl, Mississippi. Test very small, narrowly elongate, subcylindrical; subovate in end view; chambers narrow, slightly larger at the distal end, with thick- ness and width about equal; sutures distinct, depressed; surface smooth; aperture terminal at end of a very slight neck, semicircular with a bifid tooth. Length, 0.57 millimeter; width, 0.22 millimeter; thickness, 0.12 millimeter. Holotypc No. 4012, locality Ti6C, San Sebastian formation. Rare at L4C; common at T^GC of the San Sebastian formation. GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 309 Triloculina oblonga (Montagu) Plate 3, figures 8a-c Vtrmiculum oblongum Montagu (1803) Test. Brit. 522. pi. 14, fig. 9. Recent, off British Isles. Triloculina oblonga d'Orbigny (1826) Ann. Sci. Nat. 7: 300. no. 16. Recent, Adriatic, Mediterranean, North Atlantic. Antilles; middle Eocene to Pliocene, France and Italy; Modeles, no. 95. Calcaire grossier de Paris. — Cushman (1917) Bull. U. S. Nat. Mus. 71 (6) : 69. pi. 26, fig. 3, text figs. 35, 36. Recent, north Pacific; (1929) Bull. U. S. Nat. Mus. 104 (6) : 57. pi. 13. figs. 4. 5. Recent, widely distributed in Atlantic. Test olive-shapeci, slightly compressed, oval in si(ie view, short oval in end view; chambers elongate, wider aborally; sutures only slightly depressed, those bounding the third chamber frequently oblique to general axis of the test; aperture terminal, without a neck, nearly circular in shape with a narrow simple tooth. Length, 0.43 milli- meter; width, 0.20 millimeter; thickness, 0.12 millimeter. Plesiotype No. 4014, locality L6C, San Sebastian formation. Rare at L2C and L6C of the San Sebastian formation, at A6 of the Lares formation, at A91a of the Los Puertos formation, at F358 of the Quebradillas formation, and at P431 of the Ponce formation. Triloculina quadrilateralis d'Orbigny Plate 3, figures 9a-c Triloculina quadrilateralis d'Orbigny (1839). in De la Sagra. Hist. Phys. Pol. Nat. Cuba Foraminiferes 179. pi. 10. figs. 15-17. Recent, off Cuba. — Cushman (1921) Proc. U. S. Nat. Mus. 59: 71. text fig. 11. Recent, off Jamaica.— Cushman & Ponton (1922) Fla. Geol. Surv. Bull. 9: 53. pi. 7, fig. 1. Lower Miocene, Florida. Test irregularly oval in front view, width less than the thickness, subquadrate in apertural view; axes of chambers not parallel to each other; chambers rather square in section, narrower and less convex at the apertural end of each chamber; chambers flattened but not con- cave on the back, especially the last two, thus producing the quad- rangular outline; angles not carinate; sutures deep; preservation as internal molds, surface smooth ; aperture oval with distinct short neck, probably with a bifid tooth. Average specimen length, 0.80 milli- meter; width, 0.38 millimeter; thickness, 0.58 millimeter. Plesiotype No. 4015, locality L6C, San Sebastian formation. Rare at L2C, L4C, L5C, and L6C of the San Sebastian formation, and at A79 of the Los Puertos formation. Triloculina tricarinata d'Orbigny Plate 3, figures 10a, b Triloculina tricarinata d'Orbigny (1826). Ann. Sci. Nat. 7: 299. no. 7; Modfiles. no. 94. Re- cent, Red Sea.— Cushman (1917) Bull. U. S. Nat. Mus. 71 (6): 66. pi. 25. figs. 1. 2, text fig. 32. Recent, north Pacific; (1932) Bull. U. S. Nat. Mus. 161 (1) : 59. pi. 13, figs. 3a, b. Recent, tropical Pacific. 310 SCIENTIFIC SURVEY OF PORTO RICO Test of symmetrical proportions, the edges of the test sharply angled, whole test somewhat longer than wide; in transverse section an equilateral triangle; chambers little inflated, shape of chambers subordinate to that of the test; sutures slightly depressed; aperture without a neck, round with a narrow tooth. Length, 0.43 millimeter; width, 0.29 millimeter; thickness, 0.26 millimeter. Plesiotype No. 4016, locality P253, Ponce formation. Rare at P431 and P253 of the Ponce formation. MILIOLINELLA Wiesuer 1931 Miliolinella circularis (Bornemann) Plate 4, figures 8a-c Triloculina circularis Bornemann (1855) Zeitschr. deutsch. geol. Ges. 7: 349. pi. 19, flg. 4 Middle Oligocene, Germany. — Cushman (1929) Bull. U. S. Nat. Mus. 104 (6) : 58. pi. 13 flgs. 6, 7, pi. 14, figs. 1, 2. Recent, widespread. Test circular in side view; in end view broadly oval, back rounded; chambers inflated; sutures distinct, depressed; aperture a narrow crescentiform slit. Length, 0.63 millimeter; diameter, 0.57 by 0.65 millimeter. Plesiotype No. 3997, locality P2, Ponce formation. Very rare at P2 and P431 of the Ponce formation. Rare at A79 of the Los Puertos formation. The Porto Rican form is more perfectly circular in side view than is the original figure of this species. PYRGO Defrance 1824 Pyrgo bougainvillei (d'Orbigny) Pl.ATE 4, FIGURES la-C BiJoculina bougainvillei d'OrV)igny (1839) Voy. Anifir. M6rid. 5 (5): Foraniinif&res 67. pi. 8, figs. 22-24. Recent, off west coast of South America. Test small for the genus, elongate, oval in front and side views, broadly oval in end view with width about twice the thickness of the test; edge shouldered near the suture; chambers symmetrical, com- pressed; sutures slightly curved in side view; aperture oval, tooth bifurcate. Length, 0.40 millimeter; width, 0.24 millimeter; thickness, 0.18 millimeter. Plesiotype No. 3953, locality L6C, San Sebastian formation. Rare at L4C and L6C of the San Sebastian formation. GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 311 Pyrgo clypeata (d'Orbigny) Plate 4, figures 2a-c Biloculina clypeata cl'Orbigny (1846) Foram. Foss. Vienne 263. pi. 15, flgs. 19-21. Middle Miocene, Vienna. Test somewhat compressed, oval in front view, narrowly oval in edge and end views; edge shouldered; chambers little inflated; sutures slightly depressed, slightly concave toward the penultimate chamber in edge view; aperture oval with narrow bifid tooth. Length, 0.45 millimeter; width, 0.35 millimeter; thickness, 0.29 millimeter. Plesiotype No. 3954, locality A79, Los Puertos formation. Rare at L3C, L4C, L5C, and L6C; common at L2C of the San Sebastian formation. Rare at A6 of the Lares formation, at A86 of the Cibao formation, at A79 of the Los Puertos formation, and at A64 and F358 of the Quebradillas formation. Pyrgo inornata (d'Orbigny) Plate 4, figures 3a-c Biloculina inornata d'Orbigny (1846) Foram. Foss. Vienne 266. pi. 16. figs. 7-9. Middle Miocene, Vienna. Test rotund, oval in front and edge views, subcircular in end view, thickness of test equal to or slightly greater than the width; edge rounded; chambers inflated; sutures slightly depressed, curved in side view with the aboral end of the last chamber overlapping and covering the preceding chamber; aperture round with bifid tooth partly filling the opening. Length, 0.48 millimeter; width, 0.33 millimeter; thick- ness, 0.36 millimeter. Plesiotype No. 3955, locality A79, Los Puertos formation. . Rare at L4C of the San Sebastian formation and F359 of the Quebradillas formation. Common at A79 of the Los Puertos for- mation. Pyrgo lunula (d'Oibijiny) Plate 4, figures 4a-c Biloculina lunula d'Orbigny (1846) Foram. Foss. Vienne 264. pi. 15. flgs. 22-24. Middle Miocene. Vienna. Test biconvex, of average size for the genus, nearly circular in front view; biconvex in side view, edge angled; chambers somewhat in- flated; aperture narrow and elongate with a broad flat tooth nearly filling the opening. Length, 0.74 millimeter; width, 0.67 millimeter; thickness, 0.45 millimeter. Plesiotype No. 3956, locality P258, Ponce formation. 312 SCIENTIFIC SURVEY OF PORTO RICO Rare at L2C and L6C; common at L3C; abundant at L4C of the San Sebastian formation. Rare at A91a and A79 of the Los Piiertos formation. Rare at A93 and F358(2) ; common at F359 of the Que- bradillas formation. Rare at P254 and P432; common at P255 and P258 of the Ponce formation. P. depressa of d'Orbigny (Model no. 91) is more compressed than P. lunula, although the two species have many points of similarity. Pyrgo oblonga (d'Orbigny) Plate 4, figures 5a-c Biloculina oblonga d'Orbigny (1839) in De la Sagra, Hist. Phys. Pol. Nat. Cuba Foraminiferes 163. pi. 8, figs. 21-23. Recent, Cuba. Test small for the genus, moderately elongate, oval in front and side views, subcircular in end view; shoulder near suture rounded; cham- bers only slightly compressed, wider at the aboral than at the oral end; sutures slightly depressed, curved in side view; aperture oval with a broad tooth. Length, 0.37 millimeter; width, 0.21 millimeter; thick- ness, 0.26 millimeter. Plesiotype No. 3957, locality P251, Ponce formation. Rare at LlC, L2C, L4C; common at L6C of the San Sebastian for- mation. Common at A6 of the Lares formation. Rare at A43a of the Cibao formation, at A79 of the Los Puertos formation, at F359 of the Quebradillas formation. Rare at P251; common at P431 of the Ponce formation. P. oblonga is considerably less elongate than P. elongata from the Eocene at Girondc, a species with which P. oblonga seems to have been confused by many authors. P. oblonga also has less symmetrical chambers, which arc wider at the aboral than at the oral end. In this characteristic and in the curvature of the suture in edge view, P. oblonga differs also from P. bougainvillei which was found by d'Orbigny off the west coast of South America. Pyrgo peruviana (d'OrbiKny) Plate 4, figures 6a-c Biloculina peruviana d'Orbigny (1839) Voy. Amfir. Mfirid. 5 (5) : Foraminiferes 65. pi. 9, flgs. 1-3. Recent, off Peru. Test ovoid; oval in front and apcrtural views; width of test only .slightly greater than the thickness; broadly oval or subcircular in side view; shoulder next to suture rounded; chambers strongly in- flated, sutures depressed; in side view the line of contact between the GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 313 two chambers curving forward due to broadening of the last chamber so that only the last formed chamber is visible at the distal end of the test; last formed chamber noticeably larger and broader than next to last chamber, being wider near apertural end of the test, tapering toward the distal end, and of an inverted pear shape as seen from the front of the test; surface smooth; aperture broadly oval with wide, bifid tooth which is infrequently and poorly preserved. Length, 0.70 millimeter; width, 0.57 millimeter; thickness, 0.54 millimeter. Plesiotype No. 3958, locality L4C, San Sebastian formation. Common in the San Sebastian formation at L4C and L6C. Rare at A86 in the Cibao formation, at A91a of the Los Puertos formation, and at P258 of the Ponce formation. Pyrgo subspherica (d'Orbigny) Plate 4, figures 7a-c Biloculina subspherica d'Orbigny (1839) in De la Sagra, Hist. Phys. Pol. Nat. Cuba Fora- miniferes 162. pi. 8, figs. 25-27. Recent, Cuba and Jamaica. Pyroo subspherica Cushman (1929) Bull. U. S. Nat. Mus. 104 (6): 68. pi. 18, figs. 1, 2. Re- cent, Atlantic, Jamaica, Cuba, Porto Rico, Tortugas, Florida, and Bahamas. Test small, rotund, broadly oval in front view, slightly longer than wide; width of test only a little greater than the thickness; edge rounded; chambers inflated, more so at posterior end; sutures deep, sinuous in side view; in apertural view chambers semicircular; sur- face smooth; aperture broadly oval with large bifid tooth that does not fill the aperture. Length, 0.37 millimeter; width, 0.29 millimeter; thickness, 0.29 millimeter. Plesiotype No. 3959, locality L2C, San Sebastian formation. Rare at LlC, L3C, L4C, and L5C; common at L2C and L6C of the San Sebastian formation. Abundant at A6 of the Lares forma- tion. Rare at A91a; common at A79 of the Los Puertos formation. Rare at A64, A21, and F359; common at F64 of the Quebradillas formation. Rare at P251 and P434; abundant at P258 of the Ponce formation. This form is very close to P. bulloides (d'Orbigny) from the Recent of Cuba, but is less elongate, has a sinuous suture, broader aperture and characteristic posterior inflation of the chambers as seen in side view. PYRGOELLA Cushman & White 1936 Pyrgoella globulus (Bornemann) Plate 4. figures 9a-c Biloculina globulus Bornemann (1856) Zeitsch. deutsch. geol. Ges. 7: 349. pi. 19. figs. 3a, b. Middle Ollgocene, Hermsdorf. 314 SCIENTIFIC SURVEY OF PORrO RICO Test globular, circular in front, side and end views; shoulder rounded near suture; chambers strongly inflated, last chamber larger than the preceding in all dimensions; sutures slightly depressed; con- cave toward the penultimate chamber in side view; aperture tri- angular, nearly filled by a triangular tooth. Length, 0.48 millimeter; width, 0.45 millimeter; thickness, 0.45 millimeter. Plesiotype No. 3960, locality P255, Ponce formation. Very rare at P255 of the Ponce formation. P. sphaera d'Orbigny (1839. Voy. Amer. Merid. 5 (5) Foramini- feres) is very similar to this species but differs in proportions of the triangular aperture and in relative size of the two chambers. Both P. glohuhis and P. sphaera are rare species. MASSILINA Schhimberger 1893 Massilina decorata Cushman Plate 5, figure 1 Massilina decorata Cusliniaii (1922) U. S. Geol. Surv. Prof. Pap. 129F: 143. pi. 34, Jig. 7. Mint Spring Marl, lower Oligocene, Mississippi.^ — Howe& Wallace (1932) La. Dept. Cons. Geol. Bull. 2: 20. pi. 2, fig. 6. Jackson, upper Eocene, Louisiana. — Cushman (1935) F. S. Geol. Surv. Prof. Pap. 181: 13. pi. 3, figs. 14-16. Jackson, upper Eocene, Mississippi. Test subovatc in side view, slightly longer than wide with apical and apertural ends projecting a little; in edge view much compressed with sides parallel; edge rounded; early milioloid stage small, about 5 chambers in the planispiral portion of the test; surface ornamented by "very fine pits giving a finely granular matte appearance to the test"; ajierture terminal, with short, round neck, tooth not known. Length, 0.90 millimeter; width, 0.03 millimeter. Plesiotype No. 3932, locality A79, Los Puertos formation. Very rare at A79 of the Los Peurtos formation. Massilina inaequalis d'Orbigny Plate 5, figures 2a-c Massilina inaequalis Cushman (1921) Proc. U. S. Nat. Mus. 59: 72. pi. 17. figs. 12. 13. Recent, off Jamaica.— Cushman & Ponton (1932) Fla. Geol. Surv. Bull. 9: 46. pi. 3, figs. 4a-c. Cliipola formation, lower Miocene, Florida. Test elongate, biconcave, slender with both apical and apertural ends tapering; in end view with flat back forming the thickest part of the test; acute angle between the back and sides of the chambers; aperture at the end of a neck, quadrate in outline with a T-shaped tooth. Length, 1.25 millimeters; width, 0.60 millimeter; thickness, 0.32 millimeter. GALLOWAY AND HEM I NW AY, TERTIARY FORAMINIFERA 315 Plesiotype No. 3933, locality A79, Los Puertos formation. Rare at A79 of the Los Puertos formation. This species might equally well be placed under SpirolocAilina. HAUERINA d'Orbigny 1839 Hauerina sansebastianensis NEW SPECIES Plate 5, figures 3a-c, and 4a-c Test discoidal, nearly equally biconvex and biumbilicate, immature specimens relatively more convex; periphery subacute; 4 chambers in last whorl of the adult, 3 in immature forms; chambers subtriangular in shape, increasing rapidly in size, occasionally involute, more com- monly with a few milioline chambers showing in the umbilical region ; sutures sharply depressed; surface smooth; aperture in the young an elongate slit with bluntly serrate margin extending the full length of the last septal face, in the adult a tuberculate trematophore. Adult specimen diameter, 0.84 by L05 millimeters; maximum thickness, 0.37 millimeter, Holotype No. 3925a, Paratype 3925b, locality L6C of the San Sebastian formation. Common at L6C of the San Sebastian formation. This species resembles H. coinpressa d'Orbigny from the Miocene of the Vienna Basin, but is thicker and differs in character of the aper- ture and in shape of the chambers. It differs from H. hradyi Cush- man (1917. Bull. U. S. Nat. Mus. 71 (6)) in its greater thickness and different shape of the chambers. Family SORITIDAE Ehrenberg 1840 DENDRITINA d'Orbigny 1826 Dendritina preelegans NEW SPECIES Plate 5, figures 5a, b Test compressed, biumbilicate; subcircular to oval in side view, close coiled in the early part tending to become evolute in the later part; edge rounded; chambers about 12 in the last whorl, short and wide, enlarging gradually; sutures distinct, little depressed; wall ornamented with fine longitudinal striae which are not continuous from chamber to chamber; aperture an elongate, oval dendritic open- ing in the middle of the last septal face. Diameter, 1.00 by 1.33 milli- meters; thickness, 0.33 millimeter. 316 SCIENTIFIC SURVEY OF PORTO RICO Holotype No. 3889, locality A79, Los Puertos formation. Common at A79 of the Los Puertos formation. This new species is closely related to Peneroplis elegans d'Orbigny from the Recent of Cuba, but differs in the character of the aperture which is dendritic and not perforate. It differs from Dendritina antillarum d'Orbigny, also from the Recent of Cuba, in that it is less involute and the aperture is different. The aperture of D. preelegans is more regularly dendritic and it is restricted to an oval area in the center of the apertural face, not occupying the entire apertural face as it does in D. antillarum. PENEROPLIS Moatfort 1808 Peneroplis carinatus d'Orbigny Plate 5, figures 6a, b Peneroplis carinatus d'Orbigny (1839) Voy. Amer. Merid. 5 (5) : Foraminiferes 33. pi. 3. figs. 7, 8. Recent. — Cushman (1930) Bull. U. S. Nat. Mus. 104 (7) : 36. pi. 12, figs. 7-10. pi. 14, flg. 1. Recent, Atlantic Ocean, West Indies. Peneroplis pertusus carinatus Cusliman (1917) Bull. U. S. Nat. Mus. 71 (6): 87. pi. 37, flg. 4. Recent, north Pacific. Test subcircular or oval, lenticular, asymmetrical, usually bium- bonate, closely coiled and completely involute; periphery slightly lobulate, edge sharply angled; chambers short, 14-18 in the last whorl; sutures strongly recurved, scarcely at all depressed, appear- ing limbate; surface smooth; aperture a group of pores on the aper- tural face. Average specimen diameter, 1 millimeter; thickness, 0.40 millimeter. Plesiotype No. 3945, locality L6C, San Sebastian formation. Common at LlC, rare at L2C, L3C, L4C, and L6C of the San Sebas- tian formation. Rare at F358 of the Quebradillas formation and at P258 of the Ponce formation, P. carinatus differs from P. proteus in that the chambers are involute and do not become cvolute and flaring. Peneroplis proteus d'Orbigny Plate 5, figures 7a, b Peneroplis protea d'Orbigny (1839) in De la Sagra, Hist. Phys. Pol. Nat. Cuba Foraminiferes 60. pi. 7, flgs. 7-11. Recent, oflf Cuba and Jamaica. Peneroplis dubia d'Orbigny (1839) in Do la Sagra. Hist. Phys. Pol. Nat. Cuba Foraminiffires 62. pi. 6, flgs. 21. 22. Recent, ofif Cuba. Orbiculina adunca Brady (in part) (1884) Rep. Voy. Challenger Zool. 9: pi. 14. flgs. 3. 4. Recent, off Bermuda. Peneroplis proteus Cushman (1930) Bull. U. S. Nat. Mus. 104, (7): 37. pi. 13. flgs. 1-17. Recent. Atlantic Ocean: off north coast Jamaica. Tortugas. Florida. Bermuda. Bahamas. Porto Rico. — Cushman & Ponton (1932) Fla. Geol. Surv. Bull. 9: 71. pi. 10. flgs. 7-11, 14. Lower Miocene, Florida. GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 317 . Test oval or irregular in outline, ovate in edge view, moderately compressed, thickest portion in umbonal region, peripheral portions thinner, often quite flat and compressed; periphery bluntly rounded; chambers numerous, in early involute stage about 9 chambers, short and wide, later chambers becoming semi-annular, evolute and flaring; sutures distinct, slightly depressed, strongly curved; surface smooth; aperture a single row of pores on the apertural face. Diameter of coiled portion, 0.83 millimeter; thickness of coiled portion, 0.43 milli- meter; thickness of evolute chambers, 0.10 millimeter. Plesiotype No. 3946, locality L6C, San Sebastian formation. Rare at LlC and L6C of the San Sebastian formation. Rare at A79 of the Los Puertos formation, and at A64 of the Quebradillas formation. ARCHAIAS Montfort 1808 Archaias aduncus (Fichtel & Moll) Plate 5, figures 8a-c Nautilus aduncus Fichtei & Moll (1803) Test. Micr. 115. pi. 23. figs. a-e. Recent, Arabian Sea. OrbicuHna adunca d'Orbigny (1839) in De la Sagra, Hist. Phys. Pol. Nat. Cuba Foraminiferes 64. pi. 8, figs. 10-16 (not 8. 9). Recent, ofif Cuba. Archaias angulatus Cushman (1930) Bull. U. S. Nat. Mus. 104 (7): 46. pi. 16, 17. Recent, Porto Rico, Tortugas, Bermuda. Test biconvex near the center, rest of the test explanate, very thin ; edge truncate; chambers numerous, short and very wide, early ones involute to the center, later ones tending to become annular, the arc of the chamber being around 120 degrees in the young chambers and 180 or more in the adult chambers; aperture consisting of numerous large, closely but irregularly arranged pores covering the entire aper- tural face. Diameter, 1.56 by 2.06 millimeters; thickness, 0.17 milli- meter. Plesiotype No. 3868, locality P258, Ponce formation. Abundant at L6C of the San Sebastian formation. Rare at A79 of the Los Puertos formation, and at F359 of the Quebradillas forma- tion. Rare at P251 and P434; common at P258 and P259 of the Ponce formation. Many authors have discarded this species and have considered these forms as adult forms of the species A. angulatus. The Los Puertos material shows specimens of two types, each of which is sufficiently distinct to merit specific rank. /rvrTP fclLlBRAR . 318 SCIENTIFIC SURVEY OF FORTO RICO A. aduncus differs from .1. anyulatus in the following ways: 1. larger diameter 2. thinner test, both relati\ ely and actually 3. general appearance: angulatus is nautiloid with angled edge, aduncus is cx- planate with truncate edge 4. the arc of chamber width is about 180° (120° minimum) in aduncus and only 90° in angulatus 5. the last chambers of angulatus are involute to the umbilicus It seems obvious that these cannot be adult forms of A. angulatus, particularly on points 2-5 above. Archaias angulatus (Fichtel & Moll) Plate 5, figures 9a, b Nautilus angulatus Fichtel & Moll (1803) Test. Micr. 113. pi. 22. figs. a-e. Recent, Arabian Orbiculina adunca d'Orbigny (1839) in De la Sagra, Hist. Phys. Pol. Nat. Cuba Foraminiferes 64. pi. 8, figs. 8, 9. Recent, ofif Cuba. Test lenticular, nearly bilaterally symmetrical; periphery bluntly rounded ; chambers involute ; sutures limbate, with transverse connect- ing ridges corresponding to the secondary septa which are slightly offset in successive chambers and spiral outward toward the peri- phery; wall calcareous, imperforate; aperture a series of pores in several rows on the apertural face. Diameter, 0.41 to 1.36 milli- meters; thickness, 0.25 to 0.70 millimeter. Plesiotype No. 3869, locality LlC, San Sebastian formation. Rare at LlC, L4C, and L6C of the San Sebastian formation. Com- mon at A79 of the Los Puertos formation. Rare at F359 of the Quebradillas formation. Common at P258 of the Ponce formation. Archaias compressus (d'Orbigny) Plate 5, figures 10a, b Orbiculina ampressa d'Orbigny (1839) in Do la Sagra, Hist. Phys. Pol. Nat. Cul»a Foraniini- fferes 73. pi. 8, ligs. 4-7. Recent. West Indies. — Cushman (1919) Carnegie Inst. Wasli. Publ. 291: 70. pi. 7. flg. 6. Bowden formation, Miowne. Jamaica. Archaias cnnipressus Cushman (1930) Bull. U. S. Nat. Mus. 104 (7): 48. pi. 17. figs. 1. 2. Recent. West Indies; late Tertiary. Jamaica, Florida, Panama, Test compressed, discoidal, subcircular in outline; chambers involute in early portion, later portion annular; earlier involute portion about twice the thickness of annular part of test; chambers in nucleoconch simple, later chambers with chainberlcts; wall calcareous, imperforate; aperture consisting of irregularly arranged pores on the periphery. Diameter of adult form, L83 millimeters; thickness, 0.17 millimeter. Plesiotype No. 3870, locality LlC, San Sebastian formation. Common at LlC; rare at L6C of the San Sebastian formation. GALLOWAY' AND IIEMINWAY, TERTIARY FORAMINIFERA 319 Family TROCHAMMINIDAE Sclnvagcr 1877 TROCHAMMINOIDES Cushman 1910 Trochamminoides approximatus Niow species Plate 6, figures la, b Test small, subcircular in side view, the two sides nearly exactly alike, biconcave, asymmetrically planispiral, composed of about 5 volutions, each volution embracing about half of the previous volu- tion, and increasing gradually in thickness; periphery moderately lobulate; edge regularly rounded; chambers about 12 in the last volution, increasing gradually in size; sutures distinct, slightly de- pressed, narrow, mostly nearly straight, some curved slightly outward and backward, discernible from the proloculum outward; the concamer- ation is not due to irregular constrictions but due to normal septation; wall white, very finely arenaceous, consisting of very fine, angular grains of calcite bound together with a small amount of white, cal- careous cement; surface finely granular; aperture a narrow arch at the base of the septal face. Diameter, 0.66 millimeter; thickness, 0.12 millimeter. Holotype No. 4064, locality A91a, Los Puertos formation. Rare at A91a of the Los Puertos formation. This species differs from T. proteus (Karrer) , from the Cretaceous of the Vienna Basin, in lacking an early tubular portion, in having a greater number of whorls, more nearly planispiral symmetry, definite septation and small aperture. It also differs from Recent specimens identified with T. proteus Brady (1884. Rep. Voy. Challenger Zool. pi. 40) in the planispiral symmetry of the early spire, more regular concameration, and the smaller aperture, as well as the smaller size and greater number of volutions. It is quite unlikely that the large number of forms from Jurassic, Cretaceous, Tertiary, and Recent ages which have been identified with T. proteus (Karrer) actually belong to that species. Family LITUOLIDAE Reuss 1861 CYCLAMMINA Brady 1876 Cyclammina acutidorsata (Hanlken) Plate 6, figures 2a, b Haplophragmium acutidorsatum Hantken (1868) Magyar, foldt. tarsulat munkalatai 4: 82. pi. 1, flgs. la, b; (1876) Mitth. Jahrb. k. ungar. geol. Anstalt 4: 12. pi. 1. figs. la. b. Upper Eocene and lower Oligocene. Hungary.— Andreae (1884) Abh. geol. Specialkarte Elsaas- Loth. 2 (3) : 197. pi. 7, flgs. 6a. b. Middle Oligocene, Alsace. 320 SCIENTIFIC SURVEY OF PORTO RICO Test planispiral, compressed, nearly completely involute, biumbili- cate; periphery bluntly angled, very slightly lobulate; 10-11 chambers in the last whorl; septal face triangular in apertural view; sutures very slightly depressed; wall coarsely arenaceous, made of calcareous sand grains; aperture crescentiform, at base of the last septal face, on the periphery. Diameter, 0.95 millimeter; thickness, 0.32 millimeter. Plesiotype No. 3887, locality P3, Ponce formation. Very rare at P3 of the Ponce formation. Family ATAXOPHRAGMIIDAE Schwager 1877 VALVULINA d'Orbigny 1826 Valvulina oviedoiana d'Orbigny Plate 6, figures 3a, b Valvulina oviedoiana d'Orbigny (1839) in De la Sagra, Hist. Phys. Pol. Nat. Cuba Foramini- feres 103. pi. 2, figs. 21. 22. Recent. Cuba. — Cushman (1921) Proc. U. S. Nat. Mus. 59: 51. pi. 11. flgs. 11-14. Recent, north coast Jamaica; (1922) Carnegie Inst. Wash. Publ. 311: 29. pi. 2. flgs. 7. 8. Recent. Tortugas; (1922) Bull. U. S. Nat. Mus. 104 (3): 64. pi. 11. flgs. 2-5. Recent. Florida. Bahamas, etc.; (1937) Cushman Lab. Foram. Res. Spec. Publ. 8: 10. pi. 2. flgs. 4, 5. Recent. Bermuda, Florida, and West Indies. Test large for the genus, tetrahedral, an inverted three-sided pyramid in shape, initial end pointed, the sides rapidly diverging, losing the three-sided form so that the apertural end is subcircular in shape; chambers triserially arranged, rapidly increasing in size; sutures depressed; wall coarsely arenaceous with much calcareous cement; aperture along the inner margin of the last chamber with a large projecting tooth or valve. Height of test, 1.2 millimeters; diam- eter of apertural end, 1.01 by 1.20 millimeters. Plesiotype No. 4026, locality L6C, San Sebastian formation. Common at L6C ; rare at L4C of the San Sebastian formation. Rare at P258 of the Ponce formation. VALVULAMMINA Cushman 1933 Valvulammina cornucopia NEW SPECIES Pl..M'E 6, FIGURES 4a-C Test shaped like a small, twisted cornucopia, widening very rapidly from the jiointed initial end; spire of medium height, last whorl con- stituting about half the height of the test; 4 to 5 chambers in the last whorl (in no case as many as 6), chambers loosely appressed, in- flated, increasing rapidly in size in width, but not in length or height; sutures distinct, depressed; wall mcdiuni coarsely arenaceous, smoothly GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 321 finished with much calcareous cement; aperture large, at the base of the last septal face, with short, wide, valvular tooth; apertural face large, flaring, constituting about half of the last whorl on the ventral side of the test. Height, 0.70 millimeter; diameter of apertural end, 0.90 by 1.13 millimeters. Holotype No. 4025, locality L6C, San Sebastian formation. Rare at L4C and L6C of the San Sebastian formation. Common at P258 of the Ponce formation. This species is very close to Valvulammina deformis (d'Orbigny) as figured by Fornasini (1904. Mem. R. Accad. Sci. Istit. Bologna Sez. Sci. Nat. VI. 1) from the Eocene of France, but the test is higher, the chambers are fewer in each whorl and do not increase as rapidly in width, the last chamber is not as large, and the apertural valve is less well formed. It differs from V. affinis Cushman and Bermudez from the Eocene of Cuba (1937. Contr. Cushman Lab. Foram. Res. 13) in having fewer chambers in the last whorl, and more depressed sutures. VERNEUILINA d'Orbigny 1840 Vemeuilina mexicana Nuttall Plate 6, figure 5 Vemeuilina mexicana NuttaU (1932) Jour. Paleont. 6: 6. pi. 2, figs. 1, 2. Lower Oligocene. Tampico Embayment, Me.\ico. Karreriella mexicana Cushman (1937) Cushman Lab. Foram. Res. Spec. Publ. 9: 130. pi. 15. figs. 13, 14. Test short, stout fusiform in shape, broadest across the middle of the last three chambers; apertural end rounded, initial end acute; nearly circular in cross section; chambers indistinct, last whorl con- stituting the major portion of the test; sutures obscure, flush with the surface; wall of medium arenaceous texture; aperture a short, straight or slightly curved oval slit located slightly above and parallel to the base of the last septal face; apertural face slightly concave. Length, 0.77 millimeter; maximum diameter, 0.60 millimeter. Plesiotype No. 4031, locality P2, Ponce formation. Rare at P2 and P3 of the Ponce formation, and at A93 of the Quebradillas formation. We have several complete adult specimens of this species, none of which show an early 4 or 5 chambered stage or an adult 2 chambered stage. It seems probable, therefore, that this is a true Vemeuilina. 322 SCIENTIFIC SURVEY OF PORTO RICO DOROTHIA Pluniiiier 1!)31 Dorothia caribaea Cu.-^hnum Plate 6, figuhks 6;i, b Doruthia curibtica Cu.'-lunun (193G) Cushman Lab. Forani. Rus. Spue. I'ubl. 6: 31. pi. o, ligs. 3a-c. Recent, C'ul)a. Test short and stout, broadly rounded at both ends, sides ahiiost l)arallel, nearly round in cross section; chambers closely appressed, early whorls with 4 or 5 chambers in a whorl, 2 to 3 pairs in biserial ])art of the test; sutures slightly depressed, somewhat oblique; wall coarsely arenaceous made of calcite grains with calcareous cement so that the surface is fairly smooth; aperture a high arched opening constricted at the base and extending up onto the last septal face. Length, 1.00 millimeter; width. 0.60 millimeter; thickness, 0.50 milli- meter. Plesiotype No. 3901, locality P255, Ponce formation. Rare at P25o of the Ponce formation. Dorothia cylindrica (Nnltall) Plate 6, figures 7a-c (laudryina cylindrica Nuttall (1932) Jour. Paleont. 6: 7. pi. 2, fig. 7. Lower Oligocenc, Mexico. Dorothia cylindrica Cushman (1937) Cushman Lab. Foram. Uvs. Spec. Publ. 8: SS. pi. 9. fig. 25. Lower Ohgocene, Mexico. Test elongate, compressed cylindrical, with triserial part of test slightly trihedral in shape; triserial portion constituting only about Mi or less of the test; chambers closely appressed, 4-5 pairs in the biserial part of the test; sutures narrow, slightly deiiressed, rather indistinct; wall arenaceous with fine to medium size grains and much cement; aperture a high arch extending up onto the last septal face with low upper lip. Length, up to 1.82 millimeters; diameter, 0.6 millimeter. Plesiotyjie No. 3916, locality P2, Ponce formation. Rare at P2 and P255 of the Ponce foiinatiou. Dorothia praelonga (KancM) Pl.'VTE 6, FiGUUES 8a, b daudri/ina praelonga Karrer (1877) Abh. k. k. geol. Reichs. 9: 371. pi. IGa, llg. G. Middle ^Miocene, V'ienna Basin. Dorothia pruclonqa Cusliman (1937) Cushman Lab. Koram. Res. Spec. Publ. 8: 90. pi. 10, figs. 2, 3. Miocene, Vienna Basin, and Bulgaria. Test small, elongate, cylindrical except for a very brief trihedral initial end. nearlv circular in cross section; chambers numerous, 6 GALLOWAY AND 11 EM IN WAY, TERTIARY FORAMINIFERA 323 pairs or more in biscrial part of test, closely appresscd; sutures dis- tinct, slightly depressed, nearly straight and transverse to axis of test; wall finely arenaceous with much cement, surface smooth; aper- ture a low arch at the base of the septal face. Length, 0.88 milli- meter; breadth, 0.30 millimeter; thickness, 0.34 millimeter. Plesiotype No. 4046, locality P2, Ponce formation. Rare at P2 of the Ponce formation. GAUDRYINA d'Oibigny 1839 Gaudryina asiphonia Andreae Plate 7, figures la, b Gaudryina siphonella asiphonia Andreae (1884) Abh. Gecl. Specialkarte Elsass-Loth. 2 (3): 200. pi. 7, fig. 7. Middle Oligocene, Alsace. Gaudryina asiphonia Nuttall (1932) Jour. Paleont. 6: 7. pi. 2, flg. 2. Lower Oligocene, Tampico Embaynient, Me.xico.— Hadley (1934) Bull. Am. Paleont. 20 (70 A) : 8. pi. 1, flg. 6. Oligocene, probablj- upper, north coast ol Cuba. Test of average size for the genus, elongate, slightly tapering, oval in cross section; triserial portion very short constituting only about 1/4 of the length of the test; 4 pairs of chambers in the biserial por- tion of the test, these chambers subglobular in shape, more inflated than the chambers in triserial part of the test; sutures distinct, de- pressed in biserial part of test; wall finely arenaceous, surface rather smooth; ai^erture an arched opening on the inner margin of the last septal face. Length, LIO millimeters; breadth, 0.45 millimeter; thick- ness, 0.35 millimeter. Plesiotype No. 4045, locality P2, Ponce formation. Rare at P2 and P3 of the Ponce formation. Specimens of G. asiphonia from Porto Rico are larger than those reported by Nuttall from Mexico, which were only 0.60 millimeter long. Gaudryina glabrata maxima jsew vakiety Plate 7. figukes 2a, b Test large, stout, early triserial portion very short, later biserial portion constituting most of the test; test lozenge shaped in cross section; chambers numerous, quite distinct, with the lower side exca- vated so that the test has a transversely rugose appearance ; early and middle portion without the transverse rugae but with papillae which more or less obscure the sutures; sutures depressed, nearly horizontal or slightly convex toward the aperture in the middle ; wall arenaceous with much cement; surface moderately rough; aperture an elongate, low arch at the base of the last septal face. Length of figured speci- 324 SCIENTIFIC SURVEY OF PORTO RICO men, 1.11 millimeters; breadth, 0.7 millimeter; thickness, 0.45 milli- meter. Plesiotype No. 4050, locality A43a, Cibao formation. Common at A43a of the Cibao formation. This variety differs from G. glabrata (Cushman) (1937. Cushman Lab. Spec. Publ. 7) in the much larger size and the presence of papillae. Gaudryina karreriana Cushman Pl.^te 7, FIGURES 6a, b Gaudryina karreriana Cushman (1936) Cushman Lab. Foram. Res. Spec. Publ. 6: 8. pi. 1, figs. 18a, b. Miocene, Hungary. Test large for the genus, elongate, tapering, enlarging abruptly at the initial end which is rather sharp, enlarging more gradually toward the apertural end; the early triserial portion of the test comprising about 2/5 of the test, trihedral in shape with rounded angles, the remaining biserial portion quadrangular to subcircular in cross sec- tion; 2 or usually 3 pairs of chambers in the biserial part of the test; sutures narrow, slightly depressed, nearly horizontal; wall of medium arenaceous texture with much cement; surface moderately rough; aperture a low arched opening on the inner margin of the last septal face. Length, 2.21 millimeters; breadth, 1.20 millimeters; thickness, 1.18 millimeters. Plesiotype No. 3917, locality P2, Ponce formation. Rare at A20 of the Quebradillas formation. Rare at P2 and P255 of the Ponce formation. Gaudryina puertoricana NEW SPECIES Pl.'^te 7, FIGURES 7a, b Test small, flattened triangular in the early portion, becoming thicker and rounded in the later portion; triserial portion about half the length of the test; two sides flat, the third rounded; angles sharp but not keeled; chambers indistinct, not inflated except the last two; sutures flush with the surface, inconspicuous, thickened with calcite in the triserial portion; wall very finely arenaceous, composed of calcare- ous grains and calcareous cement, white in color; surface rather smooth; aperture a small, low arch at the base of the last septal face. Length, 0.33 millimeter; width, 0.20 millimeter. Holotype No. 3918, locality L6C, San Sebastian formation. Very i-are in the San Sebastian formation at L6C only. This species differs from d'. athmtica (Bailey) as figured by Bailey GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 325 and also by Cushman (1937. Cushman Lab. Foram. Res. Spec. Publ. 7) in being much smaller, having a relatively shorter triserial portion, less inflated chambers, more inconspicuous sutures, finer wall material, flat apertural face, and absence of keels on the angles of the test. Gaudryina (Pseudogaudryina) atlantica (Bailey) Plate 7, figures 3a, b Textularia atlantica BaUey (1851) Smiths. Contr. 2: 12. pi., figs. 38-43. Recent, off northeast coast USA. Gaudryina (.Pseudogaudryina) atlantica Cushman (1937) Cushman Lab. Foram. Res. Spec. Publ. 7: 95. pi. 14, flgs. 4, 5. Recent, off northeast coast of USA; Miocene, Jamaica. Test large, elongate, tapering gradually from the pointed initial end; triangular in section; sides concave, edges angled; chambers dis- tinct, not inflated although last chamber or two may be slightly rounded; sutures slightly depressed; wall coarsely arenaceous, surface rather smoothly finished; aperture an elongate arch in a reentrant of the inner margin of the last septal face. Length 1.52 millimeters; diameter, 0.7 millimeter. Plesiotype No. 4085, locality P3, Ponce formation. Very rare at P3 of the Ponce formation. G. jacksonensis Cushman (1926. Contr. Cushman Lab. Foram. Res. 2) seems quite close but has more concave sides, stronger carinae, more chambers, and is made of finer material than G. atlantica. LIEBUSELLA Cushman 1933 Liebusella byramensis (Cushman) Plate 6, figures 9a, b Clavulina byramensis Cushman (1922) U. S. Geol. Surv. Prof. Pap. 129-E: 92. pi. 16, fig. 1; (1923) U. S. Geol. Surv. Prof. Pap. 133: 22. pi. 2, fig. 3. Oligocene, Mississippi. ? Clavulina cylindrica Palmer & Bermudez (1936) Mem. Soc. Cubana Hist. Nat. 10: 245. pi. 14, figs. 3-5, pi. 19, flgs. 1, 2. Lower Oligocene, Cuba. Liebusella byramensis Cushman (1937) Cushman Lab. Foram. Res. Spec. Publ. 8: 164. pi. 19, flgs. 16-18. Lower Oligocene, coastal plain U. S. and Mexico. Test elongate subcylindrical, circular in transverse section; early portion of test triserial and slightly bulbous, major portion of test uniserial with 2-4 chambers which constitute about two-thirds of the length of the test; wall of the test very thick, composed of agglutinated calcareous grains which are mainly small hyaline foraminifera such as Bolivina and Globigerina, not labyrinthic or vesicular but appear- ing so on eroded surfaces; interior of the chambers in transverse sec- tion shows indefinite radial projections extending inward a short distance but chamber interiors are not labyrinthic; surface arena- 326 SCIENTIFIC SURVEY OF PORTO RICO ceous but smoothly finished with a great deal of calcareous cement; aperture round, terminal and central at the end of a tapering neck-like projection of the last chamber. Length, 1.81 to 2.71 milli- meters; maximum diameter, 0.71 to 0.86 millimeter. Plesiotype No. 4132, locality P25o, Ponce formation. Rare at P3 and P255 of the Ponce formation. This Porto Rican species has decided resemblances to Cuhanina alavensis Palmer (1936. Mem. Soc. Cubana Hist. Nat. 10) and it is uncertain just what the significant differences may be. These speci- mens are more like the topotypes figured by Cushman (1937. Monog, Valvulinidae) than like the original figure of L. byramensis;. LISTERELLA Cushman 1933 Listerella cj. communis (d'Orbigny) Plate 6, figures 10a, b Clavulina communis d'Orbigny (1826) Ann. Sci. Nat. 7: 268, No. 4. Recent, Adriatic; Miocene, France: Pliocene, Italy: (1846) Forani. Foss. Vienne 196. pi. 12, flgs. 1, 2. Middle Miocene, Vienna.— Cushman & Jarvis (1930) Jour. Paleont. 4: 356. pi. 32, flg. 4. Miocene, Jamaica. Listerella communis Cushman (1937) Cushman Lab. Foram. Res. Spec. Publ. 8: 148. pi. 17, figs. 4-9. Miocene to Recent, Mediterranean region. Test elongate, subcylindrical ; chambers few, height in uniserial part of test about % the diameter; sutures distinct, slightly depressed; wall finely arenaceous, smoothly finished; aperture terminal, round, occasionally with a very short neck. Length, up to about 1.50 iiiilli- meters; average diameter, 0.27 millimeter. Plesiotype No. 4056, locality P255, Ponce formation. Rare to common at P2 and P255 of the Ponce formation. Our specimens do not have the early part jireserved. CLAVULINA (I'Orhifiny lcS26 Clavulina tricarinata d'Orhinny Pl.\te 7, FIGURES 8a. b Clavulina tricarinata d'Orbigny (1839) in De la Sagra. Hist. Phys. Pol. Nat. Culm Fora- miniffires 111. pi. 2. figs. 16-18. Recent, Cuba. — Cushman (1937) Cushman Lab. Foram. Res. Spec. Publ. 8: 22. pi. 3, figs. 1-3. Recent, West Indian region; Miocene, Florida. Clavulina c.f. C. tricarinata Cushman & McGlamery (1938) U. S. Geol. Surv. Prof. Pap. 189D: 104. pi. 24. ttg. 3. Middle Oligocenc, Mississippi. Test elongate, tapering, increasing in diameter toward the aper- tural end, triangular in cross section, sides concave; chambers numer- ous, triserial portion short, 3 to 7 chambers in uniserial part of test; GALLOWAY AND lIE.\nN\\ AY, TERTIMiY FOliAMINIFERA 327 chambers of triserial part of test obscure, those of the imiserial part more distinct, each extended posteriorly over the preceding chamber on the angidations of the test; sutures in uniserial part distinct, slightly depressed with a goniatitic outline due to the aboral deflection of the chambers; wall arenaceous but smoothly finished; aperture terminal, nearly circular, without a neck, with large valvular tooth. Length, up to 1.52 millimeters; maximum diameter, 0.54 millimeter. Plesiotype No. 3885. locality L3C, San Sebastian formation. Rare at all San Sebastian localities, except L2C. Common at A79 of the Los Puertos formation. Rare at r64 and F359 of the Quebra- dillas formation. Rare at P254, P255, P259, and P432; abundant at P258 of the Ponce formation. CLAVULINOIDES Cushman 1936 Clavulinoides polygonalis new species Plate 7, figures 4a, b Test small, consisting of a triangular early third, a middle poly- gonal portion of variable length, and a terminal round portion; cham- bers closely apjiressed, about 3 in the middle part and 1 to 3 in the apertural part; sutures obscure, nearly flush with the surface; wall very finely arenaceous with a moderate amount of cement; surface rather smooth but distinctly granular; aperture terminal, polygonal, oval or round, with raised rim. Length, 0.75 millimeter; diameter, 0.3 millimeter. Holotype No. 4091, locality F64, Quebradillas formation. Rare at F64 of the Quebradillas formation. This form resembles Cole's species C. guayabalensis (1927. Bull. Am. Paleont. 14) , but differs in lacking the concave sides, in having 3 to 6 sides, and in having a round terminal portion. Clavulinoides triangularis (Nuttall) Plate 7, figures 5a, b ClavuUna triangularis Nuttall (1932) Jour. Paleont. 6: 8. pi. 2, fig. 4. Lower Oligocene. Mexico. Clavulinoides jarvisi Cuslmian (1936) Cushman Lab. Foram. Res. Spec. Publ. 6: 23. pi. 3, figs. 18a. b. Miocene, Trinidad; (1937) Cushman Lab. Foram. Res. Spec. Publ. 7: 135. pi. 19, figs. 3-.5. Miocene, Trinidad, and Venezuela. Clavulinoides triangularis Cushman (1937) Cushman Lab. Foram. Res. Spec. Publ. 7: 133. pi. 18, figs. 31, 32. Lower Oligocene, Mexico. Test triangular in section throughout, sides flat to slightly concave, angles subacute; sides of test nearly parallel; triserial portion short, a 328 SCIENTIFIC SURVEY OF PORTO RICO little inflated, uniserial portion constituting most of the test, but con- taining few chambers, 2 to 4; chambers closely appressed, obscure; sutures indistinct, slightly depressed; wall finely arenaceous with much cement, surface rather smooth; aperture terminal, round or oval. Length, up to 1.7 millimeters; diameter, up to 0.65 millimeter. Plesiotype No. 4086, locality P3, Ponce formation. Rare at P2; common at P254; abundant at P3 and P255 of the Ponce formation. We see no reliable distinction between C. triangularis and C. jarvisi Cushman. Family TEXTULARIIDAE d'Orbigny 1846 TEXTULARIA Defrance 1824 Textularia agglutinans d'Orbigny Plate 8, figures 2a-c Textularia agqlutinans d'Orbigny (1839) in De la Sagra, Hist. Phys. Pol. Nat. Cuba Fora- mlniferes 144. pi. 1, figs. 17, 18. 32-34. Recent, Cuba.— Cushman (1922) U. S. Geol. Surv. Prof. Pap. 129E: 89. pi. 14, figs, la, b. Upper Oligocene, Mississippi; (1922) Bull. U. S. Nat. Mus. 104 (3): 7. pi. 1, flgs. 4, 5. Recent, West Indies. — Cushman & Caliill (1933) U. S. Geol. Surv. Prof. Pap. 175A: 7. pi. 1, flgs. 8a, b. Miocene, Florida, North Carolina, Virginia, Maryland. Test elongate, tapering, slightly compressed; ovate in apertural view; edge rounded; chambers inflated, increasing gradually in height toward the apertural end, 7-8 pairs of chambers in average tests; sutures distinct, depressed, except in earliest portion of test; wall rather coarsely arenaceous but smoothly finished, consisting in Porto Rican material of calcareous grains; aperture a flat arch at the base of the last septal face. Length, 0.82 millimeter; width, 0.44 millimeter; thickness, 0.32 millimeter. Plesiotype No. 3996, locality LlC, San Sebastian formation. Rare at L2C, L3C, L6C; common at LlC of the San Sebastian formation. Common at A43a of the Cibao formation. These Porto Rican specimens differ from typical T. agglutinans in being slightly more compressed. Nuttall makes the same comment on his specimens of T. agglutinans from the Lower Miocene of Trini- dad. The first few chambers are more compressed than the rest of the test, with an almost subangular edge. Li general there are fewer chambers than in typical T. agglutinans. GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 329 Textularia articulata , 6. I'ppcr Oligocene. Venezuela. Test lanceolate in side view, compressed in early part of test with biconvex cross section, becoming thicker and ovoid in cross section near the apertural end; edge subangular in early part of the test, narrowly rounded in later part; i)eriphery slightly lol)ulate; chambers closely appressed, If) or more jiairs, overlapping nearly half of their width; sutures distinct, slightly depressed, nearly horizontal near the periphery; wall very finely arenaceous, surface smooth; apertural face truncate; aperture a low. transverse arch. Length of largest GALLOWAY AND llEMINWAY, TERTIARY FORAMINIFERA 331 specimen, 1.6 millimeters; width, 0.6 millimeter; thickness, 0.4 milli- meter. Plesiotypes No. 4002a, b, locality P3, Ponce formation. Rare at P2, P3, P254, and P255 of the Ponce formation. Textularia indenta xi<:w SPECIES Pl.\TE 8, FIGURES 8a-c Textularia rugosa Brady (not Reuss) (1884) Rep. Voy. Challenger Zool. 9: 303. pi. 42, fig. 24 (not 23). Recent. Pacific. — Cushman (1921) Bull. U. S. Nat. Mus. 100 (4): 114. pi. 23, flg.s. 3, 4. Recent, Pacific. Test large, rather stout, sides and edges nearly parallel throughout most of the test; roughly hexagonal in cross section; chambers bi- serially arranged throughout, numerous, about 10 pairs, closely ap- pressed; chambers with distinct lobes at the basal border, one or two on each side of the test; sutures distinct, recurved adorally between the lobes; wall rather coarsely arenaceous with much cement, smoothly finished; surface irregular due to the apically projecting lobes which form a series of deep indentations; aperture a low, elongate opening at the inner margin of the last-formed chamber in a slight re-entrant. Length of holotype, 1.77 millimeters; width, 0.81 millimeter; thick- ness, 0.61 millimeter. Holotype No. 4051, locality A43a, Cibao formation. Rare at A43a of the Cibao formation. Forms similar to this species have been found many times and have usually been confused with Reuss's species, which he called Plecnnium rugosum (1869. Sitz. Akad. Wiss. Wien 59). Reuss shows a form with fairly straight sutures without the lobate extensions which characterize T. indenta. Brady (1884) gives two figures of a form which he calls Textularia rugosa (Reuss) , neither of them being T. rugosa as described and figured by Reuss. The forms figured by Brady, with the lobate projections and irregularly excavated cham- bers seem to be identical with that found by us in the Cibao forma- tion, for which we here propose the name indenta. Cushman proposed the name Gaudryina rugulosa (1932. Bull. U. S. Nat. Mus. 161 (1)) and placed both of Brady's figures in .synonymy, but it is evident from Cushman's type figure that he had in mind a form in which the test has conspicuous, nearly horizontal corrugations separated by "excavated" areas. It is probable that the genus is Textularia, since a section of the earliest part of the test showed nothing but biserial chambers. The sections made .show no indication of septula, so the species cannot 332 SCIENTIFIC SURVEY OF PORTO RICO be Textulariella, although it looks very much like Hofker's figure of Textularia rugosa. TEXTULARIELLA Cushman 1927 Textulariella barrettii (Jones & Parker) Plate 7, figures 9a, b Textularia barrdtii Jones & Parker (1863) Rep. Brit. Assoc. Newcastle Meeting, 80, 105. Recent, Jamaica; (1876) Ann. Soc. Mai. Belg. 11: 99. woodcut. Miocene, Jamaica. Textularia pseuetotrochus Cushman (1922) Bull. U. S. Nat. Mus. 104 (3): 21. pi. 5. flgs. 1-3. Recent, West Indies and southeast U. S. Textulariella barrettii Cushman (1937) Cushman Lab. Foram. Res. Spec. Publ. 8: 66. pi. 7, flgs. 5-8. Miocene, Jamaica, and Santo Domingo; Recent, Florida and West Indies. Test forming a low broad cone with height anci breadth nearly equal, the apical end bluntly angled, the apertural end flattened or concave; about 5 pairs of chambers in the biserial part of the test, the interior of the chambers divided by partitions which extend inward from the periphery; aperture a low elongate arch at the base of the last chamber. Width, 1.00 millimeter; thickness, 0.95 milli- meter; height, 0.95 millimeter. Plesiotype No. 4004, locality A86, Cibao formation. Rare at A86 of the Cibao formation, at A21 and A93 of the Quebra- dillas formation, and at P432 of the Ponce formation. The Porto Rican specimens are smaller than those typically re- ferred to as T. barrettii, although Cushman mentions small indi- viduals of this si)ecies from the Miocene of Venezuela. VULVULINA d'Orbigny 1826 Vulvulina pachyheilus Hadley Plate 7, figures 10a, b Vulvulina pachyheilus Hadley (1934) Bull. Am. Paleont. 20 (70A) : 7. pi. 1. flgs. 2-4. Oligocene. probably upper, Cuba. — Palmer & Bcrmudez (1936) Mem. Soc. Cubana Hist. Nat. 10: 241. pi. 14, flgs. 12, 13, 16. Lower Oligocene, Cuba. Test tapering from the pointed initial end with greatest breadth near the apertural end; in cross section diamond shaped thinning from the thickened central portion to an acute edge; chambers numer- ous, short and wide, biserial throughout except for the final chamber which i.'^ uniscrial; sutures nearly straight, forming an oblique angle with Ihc edge of llie test, slightly elevated, except for the last one which is depressed; wall made of fine arenaceous material with much cement; surface smooth; aperture terminal, a fissurine slit with thickened lip. Length of figured specimen, 0.92 millimeter; breadth, 0.60 millimeter; thickness, 0.20 millimeter. Maximum length, 1.75 millimeters. GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 333 Plesiotype No. 4032, locality A93, Qiiebradillas formation. Common at A86 of the Cibao formation. Rare at A93 of the Quebradillas formation. Rare at P3 and P254; common at F2 and P255 of the Ponce formation. Vulvulina pennatula (Batsth) Plate 7, figures Ua, b Nautilus {Orthouras) pennatula Batsch (1791) Conch. Seesandes no. 13, pi. 4, figs. 13a-d. Recent. Adriatic. Bigenerina capreolus Nuttall (1928) Quart. Jour. Geol. Soc. London 84: 73. pi. 3, fig. 2. Upper Eocene and lower Miocene, Trinidad. Bigenerina pennatula Nuttall (1928) Quart. Jour. Geol. Soc. London 84: 73. pi. 3, flg. 3. Lower Miocene, Trinidad. Vulvulina pennatula Cusliman (1932) Contr. Cushman Lab. Foram. Res. 8: 76. pi. 10, figs. 1-5. Recent, West Indies; Pliocene, Sicily. Test compressed, elongate, the microspheric form widening gradu- ally from a rather sharp apical end, the megaspheric form wider at the apical end; sides of the test biconvex outward in biserial part of the test, nearly parallel in uniserial part; chambers closely appressed, earliest part of the test planispiral, followed by numerous low, curved biserial chambers, then 1 to 4 higher uniserial chambers; early part of test thicker than uniserial part; edge angled; sutures in planispiral and biserial part of test limbate and raised, depressed in uniserial part of test; wall finely arenaceous, with much cement; surface smooth; aperture terminal, fissurine, parallel to the compression of the test. Dimensions of the figured specimen, which is one of the largest found in the Porto Ricari material: length, 2.13 millimeters; width, 1.23 millimeters; thickness, 0.33 millimeter. Plesiotype No. 4033, locality P2, Ponce formation. Common at P3 and P254; abundant at P2 and P255 of the Ponce formation. Family NODOSARIIDAE Schultze 1854 SARACENARIA Defrance 1824 Saracenaria arcuata (d'Orbigny) Plate 9, figures 6a-c Cristellaria arcuata d'Orbigny (1846) Foram. Foss. Vienne 87. pi. 3, figs. 34 36. Middle Miocene, Vienna. Cristellaria acutauricularis Brady (not Fichtel & Moll) (1884) Rep. Voy. Challenger Zool. 9: 543. pi. 114, fig. 17. Recent, ofif Australia. Test subovate in side view, triangular in transverse section, outer peripheral angle sharply rounded, not carinate; chambers closely appressed, early chambers coiled, later chambers evolute, extending 334 SCIENTIFIC SURVEY OF I'ORTO RICO back nearly to the i)oint of coiling of the earlier chambers; sutures flush with the surface, marked by clear shell material; wall smooth; apertural face broad, slightly convex; aperture radiate, at the outer margin of the last septal face. Length, 1.03 millimeters; width, 0.45 millimeter; thickness, 0.38 millimeter. Plesiotype No. 3982, locality P3, Ponce formation. Rare at P2, P3, and P254 of the Ponce formation. This species is much like many forms which have been referred to aaitavricidaris of Fichtel and Moll. It differs from the original of Fichtel and Moll in the absence of a conspicuous keel in the early stages, in the thinner, less inflated test, and in the shape of the aper- tural face, which is subtriangular in S. arcuata and oval in S. acutauricularis. ASTACOLUS Montl'ort 180S Astacolus insolitus (Schwagor) Platk S, figurks 9a, Ij CrisleUtiria hisoIUa Schwagor (1866) Novara-Expwl. fU-ol. 2: 2t2. pi. G, lig. 85. Pliocene, Kar Nikobar. Cristellaria crcpidula Brady (1884) Rep. Voy. Challenger Zool. 9: pi. 67, fig. 17 (not 1!>, 20). Test elongate oval, compressed; edge narrowly rounded; about 10 chambers in the test, last chambers increasing rapidly in length; chambers involute in varying amounts; sutures distinct, flush with the surface, narrowly limbate; aperture terminal, peripheral, round and radiate. Length, 1.05 millimeters; width, 0.42 millimeter; thick- ness, 0.12 millimeter. Plesiotyi)e No. 4037, locality P254, Ponce foi'iiiation. Rare at P2, P3, P254, and P255 of the Ponce formation. A. insolita differs from ^4. crcpidula of Fichtel and Moll (1798. Test. Micr.) in the irregular overlapping of the chambers in side view. Astacolus ovatus m:\\ srKciios Pl.\TE 8. FIGURKS 10a. 1) Test elongate, compressed, the early chambers coiled, later cham- bers becoming somewhat evolutc; about 8 chambers visible, increas- ing regularly in size; sutures distinct, not depressed; wall smooth; ai)erture terminal on the outer margin of the last chamber, radial. Length, 0.70 luillimeter; width, 0.38 millimeter; thickness. 0.24 milli- meter. Holotype No. 3871. locality A81), Cibao formation. ]{ai'e at A8G of the Cibao formation. GALLOWAY AM) IIEMINWAY, TEItriARY FOEAMINIFERA 335 Aatacolus ovatiis (lifters from ^4. crcpidula in being less elongate and thicker. Astacolus sublituus (Nuttall) Plate 8, figures 11a, b Cristellaria subliluus Nuttall (1932) Jour. Paleoni. 6:11. pi. 1, figs. 13. 14. Lower Oligoceno, Mexico. Test elongate, much compressed; edge subacute, sharper on the back especially in the early part; early part with a narrow keel; chambers distinct, enlarging gradually in length, the last 2 or 3 not reacliing back to the preceding whorl; sutures narrow, obliciue, flush with the surface, very slightly limbatc; aperture round and radiate, at the outer margin of the last chamber. Length, 0.71 millimeter; width, 0.26 millimeter; thickness, 0.07 millimeter. Plesiotype No. 3872, locality A86, Cibao formation. Rare at A86 of the Cibao formation and at P255 of the Ponce formation. HEMICRISTELLARIA Stache 1864 Hemicristellaria fragaria (Giimbel) Plate 9, figures 7a, b Marginulina fragaria Giimbel (1870) Abh. k. l>ay. Akad. Wiss. Miinchcn Math.-Pliys. CI. 10: 635. pi. 1, figs. 58a, b (not c). Eocene, northern Alps. Test elongate, planispirally coiled in early portion, most of the test evolute with sutures curved but tending to become at right angles to axis of the test; test oval in transverse section, angled on outer margin, carinate when one of the costae comes on the edge of the test; chambers closely appressed, not inflated; sutures flush with the surface, obscured by the ornamentation; test ornamented by longi- tudinal costae and elongate nodes which in places merge into con- tinuous costae, the nodes are also in transverse ranks marking the chambers ; aperture near the outer edge of the last chamber, produced, with short slit extending onto the septal face, and with vestigial radia- tions. Length, L42 millimeters; width, 0.6 millimeter; thickness, 0.32 millimeter. Plesiotype No. 4053, locality A21, Quebradillas formation. Common at A21; rare at A20 of the Quebradillas formation. VAGINULINA d'Orbigny 1826 Vaginulina faba >fEW species Pl.\te 9. figures 3a-c Marginulina sp. Howe & Wallace (1932) La. Dept. Cons. Geol. Bull. 2: 34. pi. 7, fig. 3. LTpper Eocene, Louisiana. 336 SCIENTIFIC SURVEY OF PORTO RICO Test large, elongate, bean shai)cd, oval in section; about 15 cham- bers in the test, rather short in comparison with their width, enlarg- ing very gradually; edges angled in early part of test, later narrowly rounded; sutures flush with the surface, narrow in early part becom- ing broad and limbate in evolute part of test; aperture large, round and radiate, at the outer edge of the septal face. Length, 1.81 milli- meters; width, 0.51 millimeter; thickness, 0.25 millimeter. Holotype No. 4024, locality P3, Ponce formation. Very rare at P3 of the Ponce formation. Vaginulina mexicana Xuttali Pl.\te 9, FIGURES 4a, b Vaginulina elegans mexicana Nuttall (1932) Jour. Paleont. 6: 16. pi. 3, figs. 12, 16. Lower Oligocene, Alazan, Mexico.— Palmer & Bermudez (1936) Mem. Cubana Hist. Nat. 10: 277. pi. 14, figs. 23, 24. Lower Oligocene, Cuba. Test large, elongate, compressed; test biformed, the early coiled portion being much smaller than the later evolute part of the test; rectilinear portion of the test sharply rounded on the outside, sharply angled on the inside, frequently with a narrow keel; sutures con- spicuously raised along the middle of each side of the test; chamber width greater than the length, thickness and length being about the same; aperture oval, vestigially radiate, located at the outer margin of the last septal face. Length of broken specimen, 2.0 millimeters; width, 0.9 millimeter; thickness, 0.56 millimeter. Plesiotype No. 4090, locality P3, Ponce formation. Very rare at P2 and P3 of the Ponce formation. Vaginulina siliquoidea NEW SPECIES Plate 9, figures 5a, b Vaginulina legumcn elegans Cushman (1923) U. S. Geol. Surv. Prof. Pap. 133: 30. pi. 4. fig. 8. Lower and Middle Oligocene, Mississippi and Alabama. — ?Cole (1927) Bull. Am. Paleont. 14 (51): 21. pi. 3, fig. 10 (not 11). Guayabal, middle Eocene, Mexico. Test small, elongate, oval in cross section; about 7 chambers in the test, enlarging gradually in size; sutures flush wath the surface, marked by clear shell material, wide, greatest width on apertural side of tlie test; sutures sligiitly convex upward, nearly at right angles to the axis of the test; last chamber shouldered on one side, with the aperture on the other side which is slightly produced; aper- ture small, round, obscurely radiate. Length, 0.70 millimeter; width, 0.20 millimeter; thickness, 0.10 millimeter. Holotype No. 4052, locality P2, Ponce formation. GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 337 Rare at P2 of the Ponce formation. V. siliquoidea is quite different from V. elegans d'Orbigny (Model no. 54) in that it lacks the raised sutures. It differs from V. legumen (Linne) in its smaller size, fewer chambers, and the absence of a spine. MARGINULINA d'Orbigny 1826 Marginulina insulensis NEW SPECIES Plate 9, figures la-c Test elongate with nearly parallel sides; transverse section sub- triangular, with the angles rounded; early part close coiled, last 3 or 4 chambers uncoiled, only those in the uncoiled part inflated; sutures gently curved in the early coiled part of the test, oblique in the rest of the test meeting the periphery at an angle of about 45 degrees, nearly flush with the surface in the early part of the test, slightly depressed between the uncoiled chambers; aperture nearly terminal or toward the outer margin of the last septal face, round and radiate. Length, 0.76 millimeter; width, 0.21 millimeter; max- imum thickness near apertural end of test, 0.16 millimeter. Holotype No. 4062, locality P2, Ponce formation. Rare at P2 of the Ponce formation. Marginulina subcrassa Schwager Plate 9, figures 2a, b Marginulina subcrassa Schwager (1866) Novara-Exped. Geol. 2: 240. pi. 6. fig. 82. Pliocene, Kar Nikobar. Marginulina eximia (?) Cushman (1939) Contr. Cushman Lab. Foram. Res. 15: 56. pi. 9, flgs. 27, 28. Eocene, submarine core off eastern North America. Test slightly curved, stout, circular in cross section; chambers in early part coiled, later part uncoiled, uniserial; chambers closely ap- pressed, the last chamber fairly elongate, tapering to the aperture; sutures oblique, slightly depressed, more so on the concave side of the test; surface smooth; aperture produced, radiate, terminal on convex side of the test. Length, 0.85 millimeter; diameter, 0.29 milli- meter. Plesiotype No. 4057, locality P2, Ponce formation. Rare at P2, P254, and P255 of the Ponce formation. 338 SCIENTIFIC SURVEY OF PORTO RICO AMPHICORYNE Sthlumbeiger 1881 Amphicoryne obliqua NKW SPECIES Plate 9, figures 8a, b Test bifonned, the outer edge sharply angled, the early part com- pressed consisting of several chambers arranged as in Vaginulina, later part nodosarian, not compressed, consisting of only one globular chamber; sutures somewhat depressed; surface ornamented by diago- nal costae, about 18 on the last chamber and fewer on the vaginuline part of the test; costae discontinuous across the sutures; a few costae extend up onto the lower part of the neck, the upper part of which is ornamented with a few obscure horizontal rings; aperture terminal, round, not definitely radiate. Length, 0.73 millimeter; width, 0.27 millimeter. 'Holotype No. 3861, locality P2, Ponce formation. Very rare at P2 of the Ponce formation. This species differs from A. jalx (Jones and Parker) (1860. Quart. Jour. Geol. Soc. London 16) particularly in the oblique and discon- tinuous costae, GLANDULINA d'Orbigny 1839 Glandulina comatula (Cuslunan) Platk 10, EitiUKES 6a-c; Plate 11, figukks lu, b Noclusariu comatula Cushman (1923) Bull. U. S. Nat. Mus. 104 (4) : 83. pi. 14, fig. 5. Kcccnt, Ciulf of Mexico, West Indies, Caribbean. — Nuttall (1928) Quart. Jour. Gcol. Soc. London 84: 84. pi. 5, flg. 3. Lower Miocene, Trinidad. Test short and stout, gradually enlarging from the first cham- ber, initial end broadly rounded, usually with a short, stubby spine, apertural end tapering to the aperture; chambers few, 2 to 5, slightly inflated, gradually enlarging; sutures obscure, slightly depressed; sur- face ornamented by numerous low, rounded, longitudinal costae, con- tinuous from chamber to chamber, close together, 35^5 on the last- formed chamber; in the adult, the apertural end of the last chamber may be smooth; ajierture terminal, central, distinctly radiate. Length, 1.20 millimeters; diameter, 0.35 millimeter. Plesiotypes No. 3919a, b, locality P2, Ponce formation. Common at P2; rare at P3 and P255 of the Ponce formation. Glandulina gallowayi (Cii.shin.ni) Pl.\TE 11, FIGURES 2a, b Glandulina comata Cialloway & Morrey (1929) Bull .'\ni. Paleont. 15 (55): 13. pi. 1, flg. 7a, b. lyower Oligocene, Ecuador. rsendoglanduUna yallnuayi Cushman (1929) Contr. Cushman Lab. hOraiii. Kes. 5: ST. pi. 13, flg. 13. Oligocene, or Miocene, Ecuador. (JALLOWAY AND JIEMJNWAY, TERTIARY FORAMINIFERA :VS9 Test pyriform, inaxinium diameter above the middle of the test, form ratio 1.4; initial end subacute, apertural end conical; chambers few, obscure, uniserial in arrangement; sutures obscure; surface orna- mented by 20 to 24 high, rather heavy, round costae which are con- tinuous from the apical end to near the apertural end; apertural face smooth; aperture terminal, slightly produced, round, with radiating slits. Length, 0.55 millimeter; diameter, 0.41 millimeter. Plesiotype No. 3951, locality A86, Cibao formation. Rare at A86 of the Cibao formation and P254 of the Ponce forma- tion. Although forms which are identified as Lagena strumosa Reuss occur in the Porto Rican material, they do not come from the same localities as the specimens of G. gallowayi and, in any event, it seems im- probable that they could be young specimens of G. galloivayi as was stated by Cushman in 1929 for the following reasons: 1. The shape of the apical end of the test is different in the two forms, that of G. galloivayi being much more tapering. 2. The aperture is distinctly radiate in G. gallowayi but is not in L. strumosa. Glandulina mauricensis Howe & Roberts Plate 10, figure 7 Pseudoglandulina mauricensis Howe & Roberts (1939) La. Dept. Cons. Geol. Bull. 14: 48. pi. 6, fig. 10. Middle Eocene, Louisiana. Test small, stout, circular in cross section, composed of two spheroidal chambers, last chamber overlapping about a third of the earlier one; suture depressed; initial chamber with an apical spine; aperture terminal, round and radiate. Length, 0.38 millimeter; diameter, 0.22 millimeter. Plesiotype No. 4047, locality P254, Ponce formation. Rare at P254 of the Ponce formation. DENTALINA d'Orbigny 1839 Dentalina advena (Cushman) Plate 9, figures 9a, b Nodosaria advena Cusliman (1923) Bull. LT. S. Nat. Mus. 104 (4) : 79. pi. 14, fig. 12. Recent. Gulf of Me.xico, Caribbean Sea, and off northea&tern U. S. Dentalina communis Cushman & Cahill (1933) U. S. Geol. Surv. Prof. Pap. 175A: pi. 5, fig. 2. Miocene, Florida. Test elongate, slender, slightly tapering, slightly curved, circular in cross section; chambers numerous, about 7 in the complete test, those near the apertural end inflated a little; sutures oblique, slightly 340 SCIENTIFIC SURVEY OF FORTO RICO depressed, more so on the convex edge of the test; surface smooth, un- ornamented; aperture radiate, eccentric, nearer the concave side of the test. Length, 1.55 millimeters; diameter, 0.28 millimeter. Plesiotype No. 3895, locality P254, Ponce formation. Rare at P2, P254, and P255 of the Ponce formation. Many forms which have been referred to D. communis (d'Orbigny) probably belong to D. advena. The features of d'Orbigny's species are well shown by Fornasini in his reproduction of d'Orbigny's un- published figure (1898. Mem. R. Accad. Sci. 1st. Bologna V. 7) in which the test has 12 chambers and an apical spine. D. advena bears a strong resemblance to D. debilis Hantken (1875. Mitt. Jahr. k. ungarn geol. Anstalt 4 (1)). Dentalina halkyardi Cushman Plate 9, figures 10a, b Dentalina halkyardi Cushman (1933) Contr. Cushman Lab. Foram. Res. 9: 9. pi. 1. flg. 20. Upper Eocene, South Carolina and Biarritz. Test elongate, very slightly tapering; chambers about 5, subglobular, the early ones more closely appressed than the later ones; early sutures obscure, nearly flush with the surface of the test, later sutures depressed; surface conspicuously ornamented, the early chambers ornamented by discontinuous longitudinal costae, about 7 visible on each side of the test, the next chambers with spinose projections developing at the apical ends of the interrupted costae, the last cham- ber with a series of fine pustules in longitudinal lines ; aperture round, terminal and slightly produced, not definitely radiate. Length, 1.00 millimeter; diameter of last chamber, 0.30 millimeter. Plesiotype No. 3892, locality P2, Ponce formation. Very rare at P2 of the Ponce formation. Dentalina halkyardi ponceana new variety Plate 9, figures 11a, b Test elongate, tapering, slightly arcuate, circular in cross section; apical end blunt, with a short, stout spine; about 8 chambers in the test, increasing slowly in size, early chambers slightly wider than long, last chamber nearly equidimensional; first few chambers very slightly inflated, sutures nearly flush with the surface; later chambers in- flated, widest near the basal part of each chamber, separated by de- pressed sutures; surface ornamented by 12-14 diagonal costae which are continuous on \hv first 3-4 chambers and which extend to the GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 341 final chamber; costae become discontimious after the first 4 chambers with apically pointing spines or prickles on the costae of penultimate chamber and finally break up into spines on the lower two-thirds of the last chamber, apertural third of the final chamber being un- ornamented; apertural end of final chamber slightly produced, aper- ture eccentric, round and radiate. Length, 2.37 millimeters; diameter, 0.53 millimeter. Holotype No. 3893, locality P3, Ponce formation. Rare at P2, P3, and P255 of the Ponce formation. This variety differs from D. halkyardi Cushman (1933. Contr. Cushman Lab. Foram. Res. 9) in the strongly oblique character of the longitudinal costae, the presence of an apical spine, and the presence of separate spines on the final chamber only. Dentalina hillaeformis new species Plate 9, figure 12 Nodosaria pauperata Cushman (not d'Orbigny) (1923) Bull. U. S. Nat. Mus. 104 (4) : pi. 14. flg. 13. Recent off northeastern TJ. S. Test stout, curved, with short apical spine; round in cross section; chambers few, 4 to 5, somewhat inflated; sutures narrow, slightly oblique, slightly depressed, more so on the convex side of the test; aperture terminal, radiate, eccentric. Length, 0.73 millimeter; diam- eter, 0.18 millimeter. Holotype No. 3890, locality P2, Ponce formation. Rare at P2 of the Ponce formation. D. hillaeformis (sausage-shaped) differs from D. pauperata d'Orbigny (1846. Foram. Foss. Vienne) in having fewer chambers, and short, stouter test. Dentalina multilineata Bornemann Plate 9, figure 13 Dentalina multilineata Bornemann (1855) Zeitsch. deutsch. geol. Ges. 7: 325. pi. 13, flg. 12. Middle Oligocene, Germany.— Cushman (1927) Jour. Paleont. 1: 152. pi. 24, figs. 7, 8. Lower Oligocene, Alazan, Mexico.— Nuttall (1932) Jour. Paleont. 6: 14. pi. 3. flg. 5. Lower Oligocene, Alazan, Mexico. Test elongate, slender, with apical spine; chambers elongate; sutures slightly depressed; surface ornamented by numerous continuous, rather fine, longitudinal costae which are oblique on the first few chambers; apertural end not observed. Length of broken specimen, 1.5 milli- meters; diameter, 0.25 millimeter. Plesiotype No. 4035, locality P254, Ponce formation. Rare at P254 of the Ponce formation. 342 SCIENTIFIC SURVEY OF PORTO RICO Dentalina semilaevis Hantken Plate 9, figure 14 Dentalina semilaevis Hantken (1876) Magyar kir. foldt. int. evkiin. 4: 32. pi. 4, fig. 6, pi. 12, flg. 13. Lower Oligocene, Hungary.— Xuttall (1932) Jour. Paleont. 6: 15. pi. 3, fig. 8. Lower Oligocene, Mexico. Test slender, elongate, slightly arcuate; chambers numerous, about 12 in the adult test; sutures oblique to the main axis of the test, lim- bate, flush with the surface in most of the test, being slightly depressed between the last few chambers; surface ornamented by about 14 oblique, longitudinal costae, which extend from the apical end as far as the lower half of the last or next to the last chamber; aperture not preserved on the Porto Rican specimens. Length of broken specimen, 1.7 millimeters; diameter, 0.3 millimeter. Plesiotype No. 3894, locality P3, Ponce formation. Rare at P3 of the Ponce formation. NODOSARIA Lamarck 1812 Nodosaria halkyardi antillana Palmer & Bermudez Plate 10, figure 2 Nodosaria halkyardi anlillana Palmer & Bermudez (1936) Mem. Soc. Cubana Hist. Nat. 10: 269. pi. 16, fig. 3. Lower Oligocene, Cuba. Test incomplete; last chamber top-shaped, tapering abruptly at the apertural end to a distinct, rather long neck, with ring-like costae ; surface of the test ornamented by about 15 sharp, longitudinal costae which extend from the base of the neck about half the length of the chamber, terminating in spines; surface of the lower half of the chamber irregularly papillate and hispid; aperture terminal, round. Diameter of the test, 0.37 millimeter. Plesiotype No. 4060, locality A86, Cibao formation. Very rare at A86 of the Cibao formation. The Porto Rican form has fewer costae and less globular chambers than the type figure of A^. halkyardi antillana. Nodosaria longiscata d'Orbigny Plate 11, figure 8 Nodosaria longiscata d'Orbigny (1846) Foram. Foss. Vienne 32. pi. 1, figs. 10-12. Middle Miocene, Vienna.— Nuttall (1928) Quart. Jour. Geol. Soc. 84: 81. pi. 4, flg. 13. Lower Miocene, Trinidad.— Hedberg (1937) Jour. Paleont. 11: 671. pi. 91. figs. 3, 4. Oligocene, Venezuela. Test slender, stem-like except for the inflated first chamber; first chamber globular, succeeding chaml)ers indistinct, much smaller in GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 343 diameter, cylindrical in shape; sutures obscure, flush with the sur- face, at right angles to the length of the test; surface smooth; aper- tural end not found. Length, 1.5 millimeters; diameter of globular chamber, 0.32 millimeter; diameter of stem-like portion of test, 0.2 millimeter. Plesiotype No. 3934, locality P3, Ponce formation. Rare at P3 and P254 of the Ponce formation. Nodosaria obliquata (Batsch) Plate U, figure 5 Nautilus (Orthoceras) obUquatus Batsch (1791) Conch. Seesands pi. 2, figs. 5a-d. Recent, Rimini. Nodosaria obliquata Cushman (1931) Contr. Cushman Lab. Foram. Res. 7: 65. pi. 8, figs. 15-19. Recent, Rimini.— Hadley (1934) Bull. Am. Paleont. 20 (70A) : 12. pi. 1, fig. 13. Oligocene, probably upper, Cuba. Test elongate, curved, blunt apically with a stout spine; tapering at the apertural end; 7 chambers, somewhat longer than wide, becom- ing slightly more elongate toward the apertural end of the test; sutures obscure, flush between the early chambers, depressed between the last few chambers; surface ornamented with 8-10 low, longitudinal, oblique costae, extending from the apical end to the lower part of the last chamber, the upper part of the last chamber being smooth; aperture terminal, round, not definitely radiate. Length, 1.4 milli- meters; diameter, 0.18 millimeter. Plesiotype No. 3935, locality P3, Ponce formation. Very rare at P3 of the Ponce formation. The Porto Rican form is similar to N. vertebralis, but it is nearer the topotype figure of N. obliquata given by Cushman (1931. pi. 8, fig. 19). It differs from the original figures by Batsch in the fewer costae. Nodosaria pariana Hedberg Plate 11, figure 7 Nodosaria longiscata Cole & Ponton (1930) Fla. Geol. Surv. Bull. 5: 33. pi. 6, fig. 4. Lower Oligocene, Florida. Nodosaria pariana Hedberg (1937) Jour. Paleont. 11: 672. pi. 91, fig. 5. Oligocene, Vene- zuela. Test small, elongate, consisting of a bulbous proloculum and from 2 to 5 keg-shaped chambers of smaller diameter than the proloculum ; sutures slightly depressed, obscure, transverse; surface smooth; the proloculum is provided with a large apical spine; aperture not seen in the fragmentary specimens. Length of incomplete specimen, 0.7 millimeter; diameter of later chambers, 0.2 millimeter. 344 SCIENTIFIC SURVEY OF PORTO RICO Plesiotype No. 4099, locality P254, Ponce formation. Pare at P254 of the Ponce formation. Nodosaria raphanistrum (Linne) Plate 11, figure 6 Nautilus raphanistrum Linnfi (1758) Sys. Nat. ed. 10. 710. Recent. Nodosaria raphanistrum Cushman (1918) Bull. U. S. Nat. Mus. 103: 59. pi. 21. fig. 10. Oligocene, Panama Canal Zone. — Nuttall (1928) Quart. Jour. Geol. Soc. 84: 82. pi. 4. fig. 21. Lower Miocene, Trinidad; (1932) Jour. Paleont. 6: 16. pi. 3, flg. 10. Lower Oligocene, Mexico. Test large, long and stout, tapering very slightly if at all; circular in cross section; chambers a little wider than high, slightly inflated, enlarging gradually; sutures depressed; surface ornamented by about 14 prominent, heavy ribs which are continuous across the sutures and extend up to the aperture; aperture terminal, round, slightly pro- duced. Length of broken specimen, 2.5 millimeters; diameter, 1.3 millimeters. Plesiotype No. 4097, locality P3, Ponce formation. Rare at P3 and P255 of the Ponce formation. Nodosaria scalaris (Batsoh) Plate 10, figure 4 Nautilus (Orthoceras) scalaris Batsch (1791) Conch. Seesandes 91. pi. 95, flgs. B-M. Recent, Adriatic. Nodosaria scalaris Brady (1884) Rep. Voy. Challenger Zool. 9: 510. pi. 63, flgs. 28-31. Recent, Pacific, north Atlantic. Test cylindrical, composed of few, nearly spherical chambers; apical end blunt, usually with an apical spine; chambers increasing rather rapidly in size; sutures depressed; surface ornamented by numerous longitudinal costae, 10-12 visible from one side of the test; aperture terminal, round, faintly radiate, at the end of a fairly long neck. Length, 0.85 millimeter; diameter, 0.38 millimeter. Plesiotype No. 3936, locality P255, Ponce formation. Rare at P2, P254, and P255 of the Ponce formation. Nodosaria simplex Silvestri Plate 10, figure 3 Nodosaria simplex Silvestri (1872) Atti. Accad. Gioenia Sci. Nat. III. 7: 95. pi. 11, figs. 268- 272. Pliocene, Italy.— Cushman (1923) Bull. U. S. Nat. Mus. 104 (4): 68. pi. 14, fig. 10. Recent, western Atlantic. Nodosaria sp. Cushman (1933) Bull. U. S. Nat. Mus. 161 (2): 14. pi. 3, flg. 11. Recent, tropical Pacific. GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 345 Test short, consisting of only two chambers, closely appressed; suture depressed; apical end with a short, slender spine; apertural end constricting rapidly to form a tapering neck ; apertural end of neck broken. Length, 0.62 millimeter; diameter, 0.30 millimeter. Plesiotype No. 4061, locality P2, Ponce formation. Very rare at P2 of the Ponce formation. LAGENA Walker & Boys 1784 Lagena ampulla new species Plate 11, figures 9a, b Ovulina tenuis Bomemann (1855) Zeitsch. deutsch. Geol. Ges. 7: 317. pi. 12, fig. 3*a, b (not fig. 3a, b which we here select as the type of Bornemann's species). Middle Oligocene, Hermsdorf. Test flask shaped, nearly circular in cross section, body of test subcylindrical; neck moderately long, gradually contracted from the body of the test; surface ornamented with 10 medium sized costae which begin at or near the apex and extend up onto the beginning of the neck; upper part of neck unornamented; apical end broadly rounded; aperture simple, round, terminal. Length, 0.42 millimeter; diameter, 0.17 millimeter. Holotype No. 3926, locality L6C, San Sebastian formation. Rare at LlC, L2C, and L6C of the San Sebastian formation, and at P2 of the Ponce formation. This species differs from L. amphora Reuss (1862. Sitz. k. Ak. Wiss. Wien 46 (1)) in, having fewer costae, a shorter neck, and a more abruptly tapering apertural end. L. raricosta (d'Orbigny) (1839. Voy. Amer. Merid. 5 (5) : Foraminiferes) is less elongate and has a shorter neck. L. tenuis (Bornemann) , as designated above, is similar neither in shape nor ornamentation. Lagena bullosa new species Pl.\TE 11, FIGURES 10a, b Test spheroidal with long neck and short, blunt apical spine; sur- face covered with round knobs of different sizes, the neck covered with short, transverse ridges; aperture terminal, round, without phia- line lip, but with radial petaloid depressions. Length, 0.7 millimeter; diameter, 0.47 millimeter. Holotype No. 4079, locality P2, Ponce formation. Rare at P2 of the Ponce formation. Although there are numerous spinose forms of Lagena the com- 346 SCIENTIFIC SURVEY OF PORTO RICO bination of the knobby surface, rugose neck, and apical spine seems not to have been recognized. It may have evolved from a Lagenono- dofiaria, such as the form figured by Brady in the Challenger Report (pi. 63, fig. 16) . Lagena impressa new species Plate 10, figures lOa-c Test small, globular with flat base and conical apertural end, broadly oval in end view; surface ornamented with a whorl of small costae on one side, suggestive of a finger print, and with small, vertical, dividing and anastomosing costae on the opposite side; aperture small, terminal, round, neither radiate nor with a neck or phialine lip. Length, 0.5 millimeter; breadth, 0.44 millimeter; thickness, 0.36 milli- meter. Holotype No. 4094, locality P2, Ponce formation. Rare at P2 of the Ponce formation. The ornamentation of this species is similar to that of L. curvUineata (Cushman (1923) Bull. U. S. Nat. Mus. 104 (4): pi. 2. figs. 5-7), but the species arc otherwise dissimilar. Lagena nuttalli new spkcies Plate 10, figures 8a, b Lagena sulcata Nuttall (not Walker & Jacob) (1928) Quart. Jour. Oeol. Soc. London 84: 70. pi. 4, flg. 3. Oligoceno and Miocene, Trinidad. Test subglobose, slightly longer than wide, with large, inflated nock, constricted at the base and tapering rapidly to the apertural end; apical end broadly rounded, with a few inconspicuous spinose i)ro- jections; surface of the test ornamented by about 26 fine, longitudinal costae which extend from the base of the test to the apertural end of the neck; sometimes decreasing by merging and sometimes increas- ing by implantation; aperture terminal, round, without lip. Length, 0.53 millimeter; diameter, 0.35 millimeter. Holotype No. 4054, locality P254, Ponce formation. Rare at P2 and P254; common at P255 of the Ponce formation. Lagena strumosa Roiiss Pi. ATE 10, FIOUHES Oil, b Laoena strumosa Reuss (18.58) Zeltsch. Keol. Ges. 434; (18fi3) Ritz. Akad. Wiss. Wien 46 (1): 328. pi. 4, flg. 49. Olinocene, Pietzpuhl, Germany. Laqrna striata strumosa Cushman (1913) Bull. U. S. Nat. Mus. 71 (.^) : 20. pi. 7, flps. 7-10. Recent, Paclflc; (1918) Bull. U. S. Nat. Mus. 10.^: ."JR. pi. 21, flg. 7. Tvowor CJatun forma- tion, middle Miocene, Panama Canal Zone. GALLOWAY AND IIEMINWAY, TERTIARY FORAMINIFERA 347 Test subglobiilar; apical end with a short stout spine; apertiiral end with an elongate neck; surface of the test ornamented with about 30 low, round costae, several of which extend up onto the neck; costae terminate near the base of the test with spinose projections; aperture terminal, round, at the end of an elongate neck. Length, 0.61 milli- meter; diameter, 0.36 millimeter. Plesiotype No. 4055, locality P255, Ponce formation. Common at P255; rare at P3 of the Ponce formation. ROBULUS Montfoit 1808 Robulus calliferus (Stache) Plate 11, figures 12a, b Crislellaria calUfera Stache (1864) Novara-E.xped. Geol. 1: 236. pi. 23, flgs. 15a, b. Middle Tertiary, Auckland. Test round, strongly biconvex, with a narrow, angled keel of clear shell material; 9 to 10 chambers in the last whorl, enlarging very gradually in size; sutures flush with the surface, marked by broad bands of clear shell material, curved, merging at the center with a small umbonal area of clear shell material; aperture peripheral, at the outer margin of the last septal face, a narrow slit extending down into the last septal face, finely radiate on the outer sides, not on the septal face. Diameter, 1.1 millimeters; thickness, 0.67 millimeter. Plesiotype No. 4096, locality P2, Ponce formation. Rare at P2 and P254 of the Ponce formation. This species is similar to R. arcuatositriatus carolinianus Cushman (1933. Contr. Cushman Lab. Foram. Res. 9) but differs in being thicker, with narrower keel, flush sutures, and non-protruding aperture. Robulus chambersi Garrett Plate 12, figures la, b Robulus chambersi Garrett (1939) Jour. Paleont. 13: 576 pi. 65, flgs. 8. 9a, b. Middle Tertiary, Texas. Robulus clericii Coryell & Rivero (1940) Jour. Paleont. 14: 332. pi. 43, flgs. 7a-b. Middle Miocene, Haiti. Test thick, lenticular, biumbonate ; edge angled with narrow keel of clear shell material ; 5 to 6 chambers in the last whorl ; sutures limbate, flush with the surface or slightly raised, sharply curved near the umbonal region, then oblique to the periphery; umbonal region filled with clear shell material; aperture radiate at outer margin of the last chamber with a short, narrow slit extending down onto the last 348 SCIENTIFIC SURVEY OF PORTO RICO septal face. Diameter, up to 0.73 by 0.85 millimeters; thickness, 0.45 millimeter. Plesiotype No. 3973, locality P254, Ponce formation. Common at A86 of the Cibao formation. Rare at A21; common at A20 of the Quebradillas formation. Rare at P3 and P254; common at P251 of the Ponce formation. As was stated by Garrett, R. chambersi differs from R. clericii (Fornasini) , "in having fewer chambers, very prominent sutures and prominent umbones". Robulus cibaoensis new species Plate 12, figures 8a, b Test subcircular tending to be pentagonal in side view, thick dis- coidal in edge view; 5 to 6 chambers in the last whorl, last chamber most inflated especially in apertural view; sutures flush with the surface except between the last two chambers where it is a little depressed ; edge with narrow keel ; aperture at the outer margin of the last septal face, elongate and radiate; apertural face very wide, with strongly convex sides. Diameter, 0.48 by 0.56 millimeters; thickness, 0.35 millimeter. Holotype No. 4036, locality A86, Cibao formation. Rare at A86 of the Cibao formation. R. cibaoensis is somewhat similar to Brady's figure of Cristellaria articulata (1884. Rep. Voy. Challenger Zool.) but it is thicker, has fewer chambers, and a wider apertural face. Robulus convergens (Borncinann) Plate 12, figures 3a, b CristeUaria convergens Bomemann (1855) Zeitsch. deutsch. geol. Ges. 7: 327. pi. 13, flg. 16. Middle Oligocene, Germany.— Brady (1884) Rep. Voy. Challenger Zool. 9: 546. pi. 69, flgs. 1-4. Recent. Atlantic and Pacinc Oceans. — Cushman (1923) U. S. Geol. Surv. Prof. Pap. 133: 28. pi. 4, flg. 2. Lower Oligocene, Missis.sippi and Alabama; (1927) Jour. Paleont. 1 : 152, pi. 23. flg. 12. Alazan, lower Oligocene, Mexico. Lenticulina convergens Cole & Gillespie (1930) Bull. Am. Paleont. 15 (57B) : 7. pi. 3, flg. 1. Middle Oligocene, Meson formation, Mexico. Test lenticular, biconvex; broadly oval in side view; edge sub- angular with narrow margin of clear shell material ; about 6 chambers in the last whorl, enlarging rather rapidly, with the lateral faces of the last chamber drawn together so as to form an extremely nar- row septal face; sutures limbate, rather oblique, flush with the surface of the test; aperture radiate at the outer margin of the last chamber with a slit extending down onto the last septal face. Diameter, 0.83 by 1.10 millimeters; thickness, 0.40 millimeter. GALLOWAY AND HEMINWAY, TERTIARY FORAMINIhERA 349 Plesiotype No. 3974, locality P3, Ponce formation. Rare at A86 of the Cibao formation. Common at P3, P254, and P255 of the Ponce formation. Robulus falcifer (Stache) Plate 12, figures 5 and 6 Cris3()) Mem. Soc. Cubana Hist. Nat. 10: 127. text figs. ]-.*}. Lower Oligocene, Cuba. Test free, i)lanispiral throughout, involute, non-umbilicate, becom- ing more embracing as more chambers are added; round on the back; chambers numerous, 4 to 7 in the last whorl, inflated; sutures de- pressed; wall calcareous, finely perforate; surface costate; aperture absent or represented by small pores on the septal face. Length, 0.24 to 0.50 millinieter. Shallow water marls. Eocene of Egypt, Oli- gocene of West Indies, Although we have about 40 specimens, many of them well preserved, not one shows a crescentic aperture and even the pores on the septal face are scarcely larger tlian the pores elsewhere on the test. GALLOWAY AND HEMINWAY, TEUTIARY FORAMINIFERA 307 This species and the form figured by Sch wager as Polystomella ? ob.scura from the Eocene of Egypt (1883. Palcontogr. 30 (2): 138. pi. 27, figs. 2a-c) are the only known members of this new genus. The significant characteristics of this genus, whereby it dift'ers from Nonion, are its aperture, its completely involute test, and its costate ornamentation. The most closely related genus is Pullenia, rather than Nonion. Antillesina differs from Pullenia only in its porous aperture and costate ornamentation. Antillesina marielensis (Palmer) Plate 10, figures 12a, b and 13a-c Nonion ? marielensis Palmer (19.36) IMem. Soc. Cuhana Hist. Nat. 10: 127. text figs. 1-3. ]..o\ver Oligocene, Cui^a. Test small, mature specimens nearly spherical; back broadly rounded; 4 to 6 chambers in the last whorl, increasing rapidly in size, inflated; the axial diameter of the last chamber varies from 3 to 5 times that of the earliest, smallest chamber in the last whorl; cham- bers planispirally arranged, involute to umbilicus, or more, the later chambers progressively more embracing with a smaller number in the latest whorl; in apertural view the maximum thickness of the final chamber is seen, and it is about three times the thickness of the earliest chamber of the last whorl; sutures depressed, nearly radial; apertural face convex, low and broad; aperture obscure, evi- dently consisting of the scattered pores on the last septal face; wall hyaline, finely perforate; surface of test ornamented by numerous fine, narrow, low, longitudinal costae which continue down onto the last septal face but which are not continuous across the sutures, 7 to 10 costae on each side of the test, more prominent on the last few chambers. Diameter, 0.24 by 0.31 millimeter; thickness, 0.24 milli- meter. Plesiotypes No. 4063, locality LlC, San Sebastian formation, and No. 3943, locality P254, Ponce formation. Rare at LlC of the San Sebastian formation. Abundant at P254 and P255; rare at P2, P3, and P258 of the Ponce formation. 368 SCIENTIFIC SURVEY OF PORTO RICO Family ROTALIIDAE Reuss 1860 GLOBOROTALIA Cushman 1927 Globorotalia menardii (d'Orbigny) Plate 16, figures 5a-c Rotalia menardii d'Orbigny (1826) Ann. Sci. Nat. 7: 273. no. 26; Model no. 10. Recent, Adriatic. rulvinulina menardii Brady (1884) Rep. Voy. ChaUenger Zool. 9: 690. pi. 103, figs. 1, 2. Recent, world wide. — Nuttall (1928) Quart. Jour. Geol. See. London 84: 101. pi. 7, flg. 20. OUgocene and lower Miocene, Trinidad. Globorotalia menardii Cusliman (1930) Fla. Geol. Surv. Bull. 4: 60. pi. 12, flg. 1. Middle and upper Miocene, Florida. — Nuttall (1932) Jour. Paleont. 6: 29. pi. 4, fig. 16. Lower OUgocene, Mexico.— Hadley (1934) BuU. Am. Paleont. 20"(70A): 25. pi. 3, figs. 12, 13. Oligocene, Cuba. — Coryell & Rivero (1940) Jour. Paleont. 14: 336. pi. 42, figs. 34, 35. Middle Miocene, Haiti. Test compresseci, moderately convex on the ventral side, less con- vex on the dorsal side; edge with a conspicuous, rounded keel; test generally oval in side view, periphery lobulate, the lobulation being more pronounced in the last few chambers; early whorls showing on the dorsal side, involute and umbilicate on the ventral side; 5 to 7 chambers in the last whorl, increasing rather rapidly in size; sutures distinct, gently curved on the dorsal side, nearly radial ventrally; slightly depressed on the ventral side, limbate and flush or slightly raised on the dorsal side ; ventral surface near aperture covered with granulations; aperture large, ventral, opening into the umbilicus with a slight valvular lip. Diameter, 0.68 by 0.88 millimeter; thickness, 0.36 millimeter. Plesiotype No. 4082, locality A21, Quebradillas formation. Common at A21; abundant at A93 of the Quebradillas formation. LAMARCKINA Bcrthelia 1831 Lamarckina echinata NEW SPECIES Plate 15, figures 5a-c Test minute, oval in side view, pyriform in edge view, biconvex, dorsal side moderately convex, ventral side deep, with large umbilicus; })eriphery lobulate and spinose; edge sharply rounded to broadly rounded; whorls about 2, rapidly enlarging; chambers about 5 in the last whorl, rapidly increasing in size in all dimensions as added, the last chamber making \\\) about half the test on the ventral side; sutures on both sides obscure, slightly curved and slightly depressed, not limbate; surface covered with small, close-set spines, variable in size, largest on the last chamber and on the margin and back, smallest on the ventral septal face; wall finely but conspicuously per- GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 369 forate, the perforations not covered by the spines but continuing between them as the spines are built; aperture a low arch at the base of the septal face, at one side of the umbilicus but opening into it, with small upper lip. Longest diameter, 0.36 millimeter; thickness of last chamber, 0.24 millimeter. Holotype No. 4111, locality P4, Ponce formation. Rare at P4 of the Ponce formation. This species is most like "Discorbina" tuberculata Balkwill & Wright, but differs in the larger umbilicus, more highly spinose sur- face ; the pores do not emerge at the ends of the spines, and the aper- ture is farther toward the periphery. VALVULINERIA Cushman 1926 Valvulineria maclureaformis new species Plate 16, figures la-c Test oval in side view, with nearly smooth periphery ; biconvex, the dorsal side nearly flat, ventral side very deep ; edge narrowly rounded to truncate in the last chambers; chambers not inflated, about 7 in the last whorl, enlarging rapidly radially and in thickness but not in length; less than 2 whorls; sutures on the dorsal side strongly and regularly curved, limbate and slightly raised, marked by broad bands of clear shell material; sutures on the ventral side slightly curved, slightly raised, marked by broad bands of clear shell material; ven- tral side with large, deep umbilicus; dorsal side with a slight amount of secondary wall material, thickest near the middle, slightly obscur- ing the spire; wall finely but distinctly perforate; aperture a slit opening into the umbilicus and extending less than half way towards the periphery, partly covered by a valvular lip. Diameter, 0.73 millimeter; thickness, 0.45 millimeter. Holotype No. 4113, locality A86, Cibao formation. Rare at A86 of the Cibao formation. This form somewhat resembles forms which have been included in Cibicides (1935. U. S. Geol. Surv. Prof. Pap. 181: pi. 23), but neither the aperture nor the mural pores are like those of Cibicides. The specific name was suggested because of the resemblance of this form to the gastropod genus Maclurites. 370 SCIENTIFIC SURVEY OF PORTO RICO Valvulineria nuttalli Palmer & Bermudez Plate 16, figures 2a-c 7 Cancris sagra Nuttall (1932) Jour. Paleont. 6: 27. pi. 6, figs. 6, 7. Lower Oligocene. Mexico. Valvulineria nuttalii Palmer & Bermudez (1936) Mem. Soc. Cubana Hist. Nat. 10: 300. pi. 19, figs. 3-5. Lower Oligocene, Cuba. Valvulineria inaequalis Coryell & Eivero (1940) Jour. Paleont. 14: 338. pi. 43, figs. 2Ca-c. Middle Miocene, Haiti. Test large, biconvex, elongate oval in side view, narrowly elliptical in edge view; periphery smooth; edge narrowly rounded; about 2 whorls in the test, earlier whorls visible on the dorsal side, involute on the ventral side with chambers terminating in slightly elevated lobes around the umbilicus; chambers enlarging rather rapidly in all three dimensions, closely appressed, 7 to 8 in the last whorl; sutures narrow, on dorsal side flush with the surface, slightly curved between early chambers, tending to become radial between the last few chambers, on ventral side radial, depressed near the umbilicus; aperture obscure due to poor preservation, opening into the umbilicus and covered by a valvular process. Length, 1.29 milli- meter; width, 0.73 millimeter; thickness, 0.49 millimeter. Plesiotype No. 4028, locality P3, Ponce formation. Rare at P3 of the Ponce formation. As stated by Palmer and Bermudez, V. nuttalli differs from V. inaequalis (d'Orbigny) (1839. Voy. Amer. Merid. 5 (5)) in having "fewer chambers in the final whorl and a more broadly oval outline in side view". Valvulineria palmarealensis (Nuttall) Pl.\TE 16, FIGURES 3a-c Globorotalia palmarealensis Nuttall (1932) Joiu-. Paleont. 6: 30. pi. 7, figs. 1-3. Lower Oligocene, Mexico. Test somewhat compressed, biconvex, ventral side more so than the dorsal; edge narrowly rounded; test broadly oval in side view, periphery smooth; dorsally with two whorls visible, ventrally um- bilicate; 9 chambers in the last whorl, enlarging rather rapidly; su- tures distinct, gently curved on the dorsal side, nearly radial on the ventral side; sutures narrowly limbatc, flush with the surface; wall finely perforate; aperture ventral, opening into the umbilicus and ex- tending part way to the periphery, with a large, lobate, overhanging valvular extension of the last chamber. Diameter, 0.60 millimeter; thickness, 0.27 millimeter. Plesiotype No. 4029, locality P3, Ponce formation. Very rare at P2 and P3 of the Ponce formation. GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 371 This species is very close to Valvulineria nuttalli, which was de- scribed by Pahiier and Bermudez from the Oligoccne of Cuba. Both forms were found by Nuttall in the Alazan of Mexico and it may be that better suites of specimens will show that they are the same species. V. nuttalli, "differs in having fewer chambers, a more oval outline in side view and a greatly enlarged final chamber". Valvulineria paucilocula Cushman Plate 16, figures 4a-c Valvulineria paucilocula Cushman (1935) Contr. Cushman Lab. Foram. Res. 11: 37. pi. 5, figs. 7a-c. Lower Oligocene, Mississippi. Test oval in side view, narrowly ovate in edge view; back broadly rounded; periphery lobate; chambers distinct, inflated, typically 5 in the last whorl, enlarging rapidly, last chamber constituting about one-third of the test, particularly on the ventral side; dorsally with the early whorls depressed below the final whorl; ventrally umbili- cate; sutures distinct, depressed, oblique and slightly curved on the dorsal side, nearly radial on the ventral side; wall conspicuously perforate except for a clear area above the aperture on each chamber on the ventral side of the test; aperture at the base of the last chamber opening into the umbilical vestibule with a slight valvular projection of the last chamber. Diameter, 0.54 by 0.70 millimeter; thickness, 0.40 millimeter. Plesiotype No. 4030, locality P4, Ponce formation. Rare at A79 of the Los Puertos formation and at A43a of the Cibao formation. Common at P431; abundant at P4 of the Ponce for- mation. This species differs from V. cubana Palmer and Bermudez from the Eocene of Cuba (1936. Mem. Soc. Cubana Hist. Nat. 10) in the open umbilicus and absence of fingerlike processes near the umbilicus ; also the last chamber is larger than in V. cubana. EPONIDES Montfort 1808 Eponides advena (Cushman) Plate 17, figures 5a-c Rotalia advena Cushman (1923) U. S. Geol. Surv. Prof. Pap. 133: 46. pi. 7, figs. 4-6. Lower Oligocene, Alabama. Eponides advena Cole & Ponton (1930) Fla. Geol. Surv. Bull. 5: 42. pi. 11, figs. 11, 12. Lower Oligocene, Florida. Test nearly circular in side view, unequally biconvex, the ventral side the deeper; periphery not lobulate; edge bluntly angled; 5 to 6 372 SCIENTIFIC SURVEY OF PORTO RICO chambers in the last whorl, not inflated, enlarging gradually; whorls about 3, the earlier ones on the dorsal side covered with secondary material, the last whorl barely discernible; ventral side with small umbilical depression, not covered with secondary material; sutures on the dorsal side narrowly limbate, flush with the surface, oblique and tangential to the previous whorl; sutures on the ventral side nearly straight and radial, slightly depressed; surface on both sides smooth; aperture a small arch midway between umbilicus and periph- ery, at the base of the septal face, with slight upper lip. Diameter, 0.8 millimeter; thickness, 0.5 millimeter. Plesiotype No. 4104, locality A86, Cibao formation. Rare at A86 of the Cibao formation. The form figured by Palmer and Bermudez from the Oligocene of Cuba (1936. Mem. Soc. Cubana Hist. Nat. 10: pi. 19, figs. 16, 17) as E. nana (Reuss), is almost identical with the Porto Rican form here referred to E. advena, the differences being the smaller size, and as stated in their description, "ventral surface umbonate", although this does not seem particularly conspicuous in their figure. Eponides ellisorae Garrett Plate 17, figures 4a-c Eponides ellisorae Garrett (1939) Jour. Paleont. 13: 579. pi. 66. figs. 6-8. Middle Tertiary, Texas. Test subcircular, with slightly lobulate periphery, biconvex, a little deeper on the ventral side, with small umbilicus; edge angled but not carinate; whorls 2 to 3, those on the dorsal side obscured by a slight secondary deposit; chambers about 6 in the last whorl, enlarging very little as added; ventral sutures nearly radial and slightly curved, very little depressed and not limbate; dorsal sutures slightly curved, tangential to the previous whorl, flush with the sur- face and marked by clear shell material which runs out onto the periphery; surface smooth; aperture an elongate slit extending from near the umbilicus to near the periphery, enlarging outwardly, with strong upper lip. Diameter, 0.67 millimeter; thickness, 0.41 milli- meter. Plesiotype No. 4100, locality P254, Ponce formation. Rare at P254 of the Ponce formation. This species bears obvious resemblances to "Pulvinulina" bijra- mensis Cushman (1922. U. S. Geol. Surv. Prof. Pap. 129E) and to Rotalia advena Cushman ((1923) U. S. Geol. Surv. Prof. Pap. 133), GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 373 but differs from both in the lobulate periphery, sharper edge, and less dorsal deposit. Eponides exiguus (Brady) Plate 17, figures la-c Pulvinulina exigua Brady (1884) Rep. Voy. ChaUenger Zool. 9: 696. pi. 103, figs. 13, 14. Recent, Atlantic and Pacific Oceans. Eponides exigua Cusliman (1931) BuU. U. S. Nat. Mus. 104 (8) : 44. pi. 10, flg&. 1, 2. Recent, north Atlantic. Test small for the genus, about equally biconvex; edge bluntly angled; slightly lobulate in dorsal view, usually 6 chambers in the last whorl, about 21/2 whorls in the test; chambers of uniform shape, gradually increasing in size as added, slightly inflated ventrally; spiral suture on dorsal side marked by clear shell material; sutures on the dorsal side obliquely curved, flush with the surface, marked by clear shell material, on the ventral side nearly radial and some- what depressed; aperture obscure, apparently a narrow slit at the inner margin of the last formed chamber on the ventral side. Diam- eter, 0.28 by 0.32 millimeter; thickness, 0.13 millimeter. Plesiotype No. 3912, locality L2C, San Sebastian formation. Rare at L2C and L3C of the San Sebastian formation. Eponides omatissimus new species Plate 17, figures 2a-c Test delicate, planoconvex, dorsal side convex, ventral side flat with relatively large round or oval umbilical depression; edge angled; test nearly circular in side view, periphery not lobulate but with minute irregularities where the costae reach the periphery; as many as 3 whorls in the test, all visible from the dorsal side, only the last whorl from the ventral side, 12 to 15 chambers in the last whorl, enlarging very slowly; sutures radial but sharply curved near the periphery on both sides of the test; wall finely perforate; dorsal side of test conspicuously ornamented by raised spiral sutures and radial costae which are not continuous from whorl to whorl and which approximate the position of the sutures and usually bifurcate near the periphery with one branch following the curvature of the sutures; aperture a low slit at the base of the last chamber on the ventral side extending from midway of the base of the septal face nearly to the periphery; aperture obscure in most of the specimens. Diameter, 0.23 by 0.28 to 0.38 by 0.41 millimeter; thickness, 0.11 millimeter. Holotype No. 3913, locality L2C, San Sebastian formation. Common at L2C of the San Sebastian formation. Rare at A43a 374 SCIENTIFIC SURVEY OF PORTO RICO and A86 of the Cibao formation. Common at A79 of the Los Puertos formation. Rare at P2 of the Ponce formation. There are strong resemblances between E. ornatissimus and "Plan- ulina" camagueyana Bermudez from the Eocene of Cuba (1927. Mem. Soc. Cubana Hist. Nat. 11). E. ornatissimus has fewer chambers, more regular ornamentation on the dorsal side, and is smaller in size. Eponides parantillarum new SPKCIES Plate 18, figures la-c Eponides antillarum Cushman & Jarvis (1930) Jour. Paleont. 4: 364. pi. 33, figs. 14a-c, pi. 34. fig. 2. Miocene, Jamaica. Test biconvex, usually with dorsal side more convex than the ven- tral; edge angled, with narrow keel, nearly circular in dorsal or ventral view, periphery smooth, or very slightly lobate in the last few chambers; about 3I/2 whorls in the test, early ones obscure on the dorsal side of the test, only those of the last whorl being dis- tinct; 6 to 7 chambers in the last whorl, enlarging slowly, closely appressed; sutures on the dorsal side flush with surface, oblique, nar- rowly limbate, on ventral side nearly radial, flush, or slightly de- pressed near the periphery, becoming limbate and raised near the umbilical region where they become fused into a ring surrounding a small but deep umbilical depression; aperture an elongate arch with lip at the base of the last chamber on the ventral side midway be- tween the umbilicus and periphery. Diameter varies from 0.36 by 0.41 to 0.84 by 0.93 millimeter; thickness, 0.30 to 0.54 millimeter. Holotype No. 3914, locality A21, Quebradillas formation. Rare at A93; common at A20; abundant at A21 of the Quebra- dillas formation. Rare at P3 of the Ponce formation. This species is related to E. antillarum (d'Orbigny) from the Recent of Cuba, but differs consistently in several respects. The sutures on the dorsal side are more oblique, the last whorl is narrower, and the periphery is not lobulate. Several other species from Eocene- Oligocenc formations of the Gulf Coast and West Indian region are similar in some respects. Most of these lack the characteristic raised limbate ornamentation of the ventral side. Those that have similar ornamentation differ in having radial, not oblique dorsal sutures, as E. mexicana Cu.<5hman (1925. Am. Assoc. Petr. Geol. 9) and E. guayahnlensis Cole (1927. Bull. Am. Paleont. 14), or in having more chambers as in E. byramensis cubensis Palmer and Bermudez (1936. Mem. Soc. Cubana Hist. Nat. 10). GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 375 Specimens at P3 which are referred to this species differ consistently in that they are much more fiat on the dorsal side. This may be a variety of E. parantillarum or may merely show the extreme indi- vidual variation of the species. Eponides pulvinus NEW species Plate 18, figures 2a-c Test subcircular in side view, unequally biconvex, slightly convex on the ventral side, strongly convex to conical on the dorsal side; pe- riphery not lobulate; edge sharply angled; whorls about 4, on the ventral side involute leaving a very small umbilicus, the earlier whorls on the dorsal side covered by smooth secondary wall ma- terial, obscuring the spire; 5 chambers in the last whorl, increasing slowly in size; sutures on the ventral side nearly radial, very slightly curved, flush with the surface excepting between the last few cham- bers where they are depressed; sutures on the dorsal side somewhat limbate, slightly curved and tangential to the earlier whorls; surface of both sides smooth; aperture a slit at the base of the septal face extending and widening from near the umbilicus to two-thirds of the way to the periphery, with prominent upper lip. Diameter, 0.65 millimeter; thickness, 0.35 millimeter. Holotype No. 4103, locality A15, Quebradillas formation. Common at A86 of the Cibao formation. Rare at A15 of the Que- bradillas formation. This species differs consistently from E. advena (Cushman) (1923. U. S. Geol. Surv. Prof. Pap. 133) in being deeper on the dorsal side and shallower on the ventral side. It has fewer chambers and is higher dorsally than E. jacksonensis (Cushman and Applin) (1935. U. S. Geol. Surv. Prof. Pap. 181). Eponides repandus (Fichtel & Moll) Plate 17, figures 3a-c Nautilus repandus Fichtel & Moll (1798) Test. Micr. 35. pi. 3. flgs. a-d. Recent, Mediter- ranean. Eponides repandus Montfort (1808) Conch. Syst. 1: 127. 32e genre. Recent, Mediter- ranean.— Cushman & Ponton (1932) Fla. Geol. Surv. Bull. 9: 92. pi. 13, flgs. 9a-c. Lower Miocene, Florida; Recent, off Florida and the West Indies. Pulvinulina repanda Brady (1884) Rep. Voy. Challenger Zool. 9: 684. pi. 104, flgs. 18ar-c. Recent, widespread. Test subcircular, with slightly lobulate periphery, biconvex, slightly deeper on the ventral side; edge sharp and narrowly carinate; cham- bers 6 to 7 in the last whorl, closely appressed, flat on the dorsal 376 SCIENTIFIC SURVEY OF PORTO RICO side, slightly inflated on the ventral side, the last chamber with a lobed extension; sutures strongly curved and raised on the dorsal side, moderately curved and depressed on the ventral side; wall finely perforate, with scattered large pores on the last septal face; aperture an arch at the base of the septal face, highest near the umbo, without upper lip. Diameter, 1.1 millimeters; thickness, 0.5 milli- meter. Plesiotype No. 4093, locality A21, Quebradillas formation. Rare at A21 of the Quebradillas formation. Our specimens lack the ventral limbate sutures usually ascribed to this species, and have an umbonal lobe not seen in Brady's figure in the Challenger Report. They are much like the Florida Miocene specimens. Eponides ventricosus NEW SPECIES Plate 18, figures 3a-c Test subcircular in side view, strongly and almost equally convex, the dorsal side smoothly hemispherical, the ventral side slightly con- vex just beneath the periphery; periphery smooth, but sometimes slightly lobulate in the last few chambers; edge sharply angled to sharply rounded, sometimes appearing keeled, whorls about 3, nar- row, only the last one or part of the last one visible on the dorsal side, which is covered with a smooth, mottled secondary deposit; chambers 11 to 13 in the last whorl, enlarging gradually; ventral side with strong, smooth, imperforate umbo; dorsal sutures slightly curved, oblique, broadly limbate but not raised, the spiral suture also limbate but not raised; ventral sutures curved, narrowly limbate, flush with the surface, except for the last few which may be broadly depressed; ventral surface finely mottled; wall finely perforate, the perforations seeming to continue through the secondary deposit on both sides of the test; appearing on the dorsal umbo as coarse puncta, so that the form might be mistaken for Cibicides ; aperture a low arch midway between umbo and periphery, with slight upper lip. Diam- eter, 0.5 millimeter; thickness, 0.33 millimeter. Holotype No. 4116, locality A86, Cibao formation. Abundant at A86 of the Cibao formation. Rare at A21 of the Quebradillas formation, and at P251 and P259 of the Ponce formation. Eponides vortex NEW SPECIES Plate 18, figures 4a-c Test biconvex, somewhat compressed, central portion of dorsal and ventral sides inflated with last whorl thinner, forming a somewhat GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 377 explanate border; ventral side umbonate; early whorls on the dorsal side obscure, 10 chambers in the last whorl, short and wide, enlarging very gradually in size; periphery not lobate; edge sharp; sutures strongly and nearly equally curved on both sides of the test, nearly flush with the surface and marked by clear shell material, the last few on the ventral side raised; wall very finely perforate; aperture a low arch near the periphery on the inner margin of the last chamber on the ventral side, with lip. Diameter, 0.83 millimeter; thickness, 0.36 millimeter. Holotype No. 3915, locality A86, Cibao formation. Common at A86 of the Cibao formation. This species is quite distinctive, but has some points of resemblance to "Anomalina" subtenuissima Nuttall (1928. Quart. Jour. Geol. Soc. London 84). It differs in being thicker, without depressed sutures, and with a narrower final whorl on the dorsal side of the test. GYROIDINA d'Orbigny 1826 Gyroidina cf. soldanii d'Orbigny Plate 15, figures 7a-c Gyroidina soldanii d'Orbigny (1826) Ann. Sci. Nat. 7: 278. no. 5, Modeles no. 36. Recent, Adriatic. Test small, subcircular in top view, dorsal side nearly flat, the last whorl flat or depressed, the earlier whorls slightly convex, ven- tral side unusually deep for the genus, with a moderately large um- bilicus; chambers about 6 in the last whorl; dorsal sutures slightly curved and moderately oblique, not depressed or raised, obscured by a slight deposit of secondary material ; ventral sutures slightly curved, nearly radial, flush with the surface, narrow; entire surface smooth; aperture a long slit at the base of the septal face midway between umbilicus and periphery, with small upper lip. Diameter, 0.64 milli- meter; thickness, 0.5 millimeter, Plesiotype No. 4114, locality P3, Ponce formation. Rare at P3 of the Ponce formation. This form is unusually deep ventrally and has fewer chambers than most other species of the genus. There are fewer chambers in the last whorl, and the dorsal surface of the final whorl is more flattened than in d'Orbigny's model of G. soldanii. 378 SCIENTIFIC SURVEY OF PORTO RICO Gyroidina stellifera NEW SPECIES Plate 15, figures 6a-c Test subcircular in side view, biconvex, thick dorso-ventrally, dorsal side moderately convex, ventral side very convex; periphery not lobulate; edge sharply angled; whorls about 3; chambers about 6 in the last whorl, enlarging gradually; sutures on the dorsal side curved and long, oblique, tangential to the earlier whorl, limbate but scarcely raised, the spiral suture limbate and raised; dorsal side covered with a smooth secondary deposit obscuring the sutures, thicker over the earlier whorls; ventral sutures slightly curved, scarcely depressed, nearly flush with the surface; ventral side smooth, with secondary deposit around the small umbilicus, raised, in the form of a star, with a lobe just posterior to each suture; aperture a small arch at the base of the septal face, midway between umbilicus and periph- ery, with slight upper lip. Diameter, 0.76 millimeter; thickness, 0.53 millimeter. Holotype No. 4105, locality A86, Cibao formation. Rare at A86 of the Cibao formation. ROTALIA Lamarck 1804 Key to the Porto Rican Species A. Whorls involute on dorsal side R. meyerhoffi, page 381. AA. Whorls not embracing on dorsal side B. Test domed on dorsal side R. tholus, page 382. BB. Test about equally biconvex C. Periphery strongly spinose R. byramensis, page 378. CC. Periphery not strongly spinose D. Dorsal surface nearly smooth R. mexicana, page 380. DD. Dorsal surface rough; edge not keeled E. Ventral ornamentation regular R. mexicana inccatepeccnsis, page 380. EE. Ventral ornamentation irregular R. chodawensis, page 379. Rotalia byramensis Cushman Plate 19, figures la-c Rotalia byratyiensis Cushman (1922) U. S. Geol. Surv. Prof. Pap. 129E: »9. pi. 23, fig. 1. Middle Oligocene. Mississippi. — Cushman & McGlamery (1938) U. S. Geol. Surv. Prof. Pap. 189D: 110. pi. 27, fig. 3. Middle or upper Oligocene. Alabama. Test unequally biconvex, ventral side the deeper; periphery lobulate, with a short peripheral spine on each chamber located slightly anterior to the midpart of the chamber; chambers rotaloid in GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 379 arrangement, not involute on the dorsal side, 6 to 7 in the last whorl; earlier whorls on the dorsal side covered by secondary material, making a prominent boss; only the last whorl visible on the ventral side; sutures on the dorsal side slightly curved, limbate and flush with the surface; sutures on the ventral side radial, narrow and de- pressed, particularly between the last few chambers; in the umbilical region there is a circular boss with the sutures ending in a depressed ring around it; surface finely papillate on the dorsal side, coarsely papillate near the periphery on the ventral side; aperture a low arch at the base of the septal face between the umbilicus and the periphery, without definite lip. Diameter, 0.5 millimeter; thickness, 0.24 milli- meter. Plesiotype No. 4106, locality P431, Ponce formation. Common at P431 of the Ponce formation. The Porto Rican specimens seem to differ somewhat from the original figures of R. hyramensis, particularly in the much smaller size (0.5 millimeter instead of 2 millimeters), fewer chambers, and less pointed chambers, but inasmuch as the Choctaw Bluff forms, which seem identical with the Porto Rican form, have been re- ferred to this species, we are for the present placing our material in this species. Rotalia choctawensis Cushman & McGlamery Plate 19, figures 2a-c Rotalia choctawensis Cushman & McGlamery (1938) U. S. Geol. Surv. Prof. Pap. 189D: 110. pi. 27, fig. 4. Middle Oligocene, Alabama. Test subcircular in side view, biconvex, the ventral side slightly deeper; periphery irregular; edge bluntly angled; ventral side with round, prominent umbo; 10 to 12 chambers in the last whorl, gradually enlarging, somewhat irregular in size; sutures on the dorsal side slightly curved, raised, obscure; sutures on the ventral side nearly radial, depressed; surface on both sides, more on the dorsal side, covered with secondary wall material in irregular, lumpy masses; aperture a small slit at the base of the septal face, about midway be- tween umbo and periphery, with slight upper lip. Diameter, 0.56 millimeter; thickness, 0.31 millimeter. i Plesiotype No. 4107, locality LlC, San Sebastian formation. I Rare at LlC of the San Sebastian formation. This species is scarcely distinguishable from R. mexicana, differing mainly in the rougher surface. It is noteworthy that R. choctawensis does not occur in the Ponce formation, which has at least five other 380 SCIENTIFIC SURVEY OF PORTO RICO species of Rotalia, nor does any other species of Rotalia occur in the San Sebastian formation, so far as our collections show. Rotalia mexicana Nuttall Plate 19, figures 3a-c Rotalia mexicana Nuttall (1928) Jour. Paleont. 2: 374. pi. 50, figs. 6-8. Upper Eocene, Mexico. Test subcircular in side view, biconvex, deeper on the ventral side; edge angled to sharply rounded; periphery even and papillate except for the last few chambers which are extended radially into blunt points, giving an angularly lobate outline to the test; whorls not involute on the dorsal side; 10 to 12 chambers in the last whorl, obscure on the dorsal side, distinct on the ventral side, not inflated; sutures slightly curved and flush dorsally, nearly radial and conspicuously depressed ventrally, meeting near the umbilicus to form a narrow, depressed ring around an umbonal boss of clear material; surface of the dorsal side smooth or granular; surface of the ventral side papillate near the pe- riphery and granulate with small, irregular depressions up to the central boss; aperture a narrow slit at the base of the septal face be- tween umbo and periphery, nearer the periphery, without definite lip. Diameter, 0.6 miUimeter; thickness, 0.45 millimeter. Plesiotype No. 4109, locality P253, Ponce formation. Common at P4, P253, and P432 of the Ponce formation. Our specimens are uniformly deeper ventrally than dorsally. Nuttall states, "the dorsal side strongly convex and the ventral flattened". He may have been using the words "dorsal" and "ven- tral" in the opposite sense from that of general usage, or have made an error. Otherwise, our form is very similar to Nuttall's. Rotalia mexicana mecatepecensis Nuttall Plate 19, figures 4ar-c Rotalia sp. Cushman (1927) Jour. Paleont. 1: 166. pi. 26, fig. 5. Alazan Clays, Eocene or lower Oligocene, Mexico. Rotalia mexicana mecatepecensis Nuttall (1932) Jour. Paleont. 6: 26. pi. 4, figs. 11, 12. Lower Oligocene, Mexico. Test large, biconvex, ventral side the deeper; subcircular in side view; periphery smooth but papillate in the early part of the last whorl becoming slightly lobate in the later part; edge subangular in young specimens to narrowly rounded in adult; 11 to 13 chambers in the last whorl, enlarging gradually; test involute to umbo on the ventral side, with a large round or oval central boss and several small GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 381 bosses of clear shell material ; dorsal side with some of the later cham- bers distinct, the last half whorl usually raised above the earlier whorls, the earlier whorls and much of the last whorl covered with secondary wall material in a rough and papillate manner; ventral side covered with secondary material, thickest near the periphery, making an in- tricate pattern of radial and concentric knobs; sutures on the ventral side distinct, radial, depressed particularly toward the umbo; sutures on the dorsal side shghtly curved, obscure except for the last few chambers where they are depressed; aperture a short slit at the base of the septal face, nearer the periphery than the umbo, obscured by the secondary deposit, without definite Up. Diameter, 1.15 milli- meters; thickness, 0.58 milUmeter. The largest specimens are nearly 2 millimeters in diameter. Plesiotype No. 4075, locahty P431, Ponce formation. Abundant at P431 of the Ponce formation. This variety differs from R. mexicana in the rougher surface, and from R. choctawensis (1938. U. S. Geol. Surv. Prof. Pap. 189D) in the regularity of the ventral ornamentation. Rotalia meyerhoffi new species Plate 20, figures la-c Test small, oval in side view, the later part of the last whorl being very broad radially; biconvex and biumbonate, the ventral side more strongly convex, involute more than half the width of a whorl on the dorsal side, tending to become evolute on the ventral side, exposing more than the last whorl to view; periphery not particularly lobulate but with a slight projection at the anterior periphery of the last few chambers where there is a short spine; edge angular in the last few chambers, somewhat rounded by papillate secondary wall tissue in most of the last whorl; whorls few; chambers 9 to 11 in the last whorl, increasing gradually in length, very broad radially, only those of the last whorl, or less, visible on the dorsal side due to secondary shell material which covers the earlier whorls and makes an umbo ; ventrally the um- bilical region is filled by a boss of clear shell material ; surface smooth on the ventral side, granulate on the dorsal side and papillate on and near the periphery on both sides of the test; sutures on the ventral side nearly radial, narrow and depressed, on the dorsal side slightly curved, limbate and more or less raised; aperture a narrow slit at the base of the last septal face between the umbo and the periphery, nearer the periphery, with small upper lip. Diameter, 0.7 millimeter; thickness, 0.32 millimeter. 382 SCIENTIFIC SURVEY OF PORTO RICO Holotype No. 4108, locality P431, Ponce formation. Common at P431 of the Ponce formation. This species differs from all associated species of Rotalia, and from nearly all other species of the genus, in being partially involute on the dorsal side, yet with very wide chambers radially. It most nearly resembles R. aculeata (d'Orbigny) (1846. Foram. Foss. Vienne pi. 8). We take pleasure in naming this species for Dr. Howard A. Meyer- hoff. Professor of Geology at Smith College, who has devoted many years to the study of the geology of Porto Rico, and who collected the foraminiferal material for the present study. Rotalia tholus NEW species Plate 20, figures 2a-c Test subcircular in side view, dome shaped in edge view, sUghtly convex ventrally, strongly convex, domal or even campanulate dorsally; edge angled but not keeled; periphery sHghtly lobulate, a little irregular but not erose; chambers 9 to 10 in the last whorl, increasing gradually in size, not inflated; whorls not involute on the dorsal side, the earlier ones obscured by strongly papillate or knobbed secondary shell material, most marked on the sutures; ventral side with a strong umbilical plug of clear shell material, separated by a groove from the chambers; sutures on the dorsal side gently curved, hmbate and flush with the surface or raised, only those of the last whorl distinct; sutures on the ventral side slightly depressed, more so at the inner ends, not limbate, nearly straight, radial; surface of the ventral side unornamentcd except for a crescentic addition of second- ary material on the inner ends of the chambers; aperture a slit at the base of the septal face, midway between umbo and periphery, without upper Hp. Diameter, 0.55 millimeter; thickness, 0.33 millimeter. Holotype No. 4110, locality P431, Ponce formation. Common at P431 of the Ponce formation. This species differs from R. alabamensis Cushman and McGlamery (1938, U. S. Geol. Surv. Prof. Pap. 189D) in the flatter ventral side, more domed dorsal side, more chambers, and smoother, non-keeled edge. The specific name refers to the cupola or dome-shaped dorsal side of the test, from the Latin noun, tholus. GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 383 DISCORBIS* Lamarck 1804 Discorbis floridanus Cushman Plate 21, figures 5a-c Discorbis floridana Cushman (1922) Carnegie Inst. Wash. Publ. 311: 39. pi. 5, figs. 11, 12. Recent. Tortugas: (1933) U. S. Geol. Surv. Prof. Pap. 175A: 29. pi. 9. figs. 12, 13. Middle and upper Miocene, southeastern U. S. Test planoconvex, dorsal side convex, ventral side flat, or a little concave; edge narrowly rounded; test oval in side view, somewhat lobulate; 5-6 chambers in the last whorl, increasing gradually in size; sutures on the dorsal side obliquely curved, shghtly depressed except for the earlier ones which are narrowly limbate, sutures on the ventral side nearly radial, depressed; wall finely but conspicuously perforate; aperture an elongate low arch at the base of the last chamber on the ventral side extending from the umbilicus nearly to the periphery. Diameter, 0.41 by 0.50 miUimeter; thickness, 0.16 miUimeter. Plesiotype No. 3897, locality L5C, San Sebastian formation. Rare at LlC and L5C of the San Sebastian formation, and at A79 of the Los Puertos formation. Discorbis havanensis Cushman & Bermudez Plate 20, figures 3a-c Discorbis havanensis Cusliman & Bermudez (1937) Contr. Cushman Lab. Foram. Res. 13: 19. pi. 2. figs. 15, 16. Eocene, Cuba. Test compressed, round in side view, with lobulate periphery; dorsal side moderately convex and umbonate, ventral side flat or slightly convex; edge acute with narrow keel; chambers distinct, 4 to 5 in the last whorl, increasing gradually in size as added, not inflated on the dorsal side, slightly inflated on the ventral side, the last chamber extending nearly a third of the circumference of the test; dorsal sutures distinct, strongly curved, limbate but little raised, distinct in the last whorl, the earlier whorls usually covered by a secondary deposit; ventral sutures narrow, depressed, sUghtly curved, usually bordered on one or both sides by thin lines of secondary tissue extending outward from the umbilicus; umbilicus modified by addition of secondary tissue, leaving several small, lobate depressions; the exact forms made by the ventral secondary tissue varies considera- bly from specimen to specimen; wall finely perforate on both sides; aperture a sht opening into the umbilicus, covered by a thin, valvular * The gender of Discorbis is masculine since the Latin word orbis is masculine. Adjectival specific names should therefore have a masculine ending, a point which has been generally overlooked heretofore. 384 SCIENTIFIC SURVEY OF PORTO RICO flap extending from the inner, middle edge of the last chamber. Diam- eter, 0.74 by 0.64 milUmeter; thickness, 0.24 milHmeter. Plesiotype No. 4112, locaUty P258, Ponce formation. Abundant at A79 of the Los Puertos formation. Common at F64 and F359 of the Quebradillas formation. Rare at P259; abundant at P258 of the Ponce formation. Perhaps this species belongs in the genus Glohorotolia. Discorbis multisectus NEW species Plate 20, figures 4a-c Test planoconvex, dorsal side convex forming a low cone, ventral side umbilicate, chambers only slightly convex, with general ventral surface slightly concave; edge bluntly angled; nearly 3 whorls in the test, 4 chambers in the last whorl, enlarging gradually, last chamber occupying more than a third of the periphery of the test; ventral margin of last chamber lobulate due to apertural recess, pre\'ious apertural recesses covered with a flap of shell material ; sutures on the dorsal side narrow, obliquely curved, not limbate, flush with the surface except for the last one which is slightly depressed, sutures on the ventral side shghtly depressed, primary sutures short, nearly radial, secondary sutures with sigmoid curve; wall distinctly, but not coarsely, perforate; aperture large, ventral, near the umbilicus. Average diameter, 0.31 by 0.36 millimeter; thickness, 0.16 miUimeter. Holotype No. 3898, locality L5C, San Sebastian formation. Common at L5C; rare at L3C of the San Sebastian formation. This species seems related to the D. mirus-orbicularis group. It differs from D. minis Cushman (1922. Carnegie Inst. Wash. Publ. 311) in having a smooth, not lobulate outline; nonlimbate, more oblique dorsal sutures; fewer chambers, and a smaller test. It differs from D. orbicularis (Terquem) (1876. Ess. Anim. Plage Dunkerque (2)) in being less circular, and in having less elongate chambers. The specific name multisectus refers to the secondary sutures on the ventral side. Discorbis oligospiratus NEW SPECIES Plate 21, figures la-c Discorbina globularis Flint (not d'Orbigny) (1899) Ann. Rept. U. S. Nat. Mus. 1897: 327. pi. 72, flg. 2. Recent, off Florida. Test of average size for the genus, planoconvex, nearly flat on the ventral side, convex on the dorsal side; on dorsal side somewhat involute, early whorls depressed; umbilicate on th(> ventral side, the GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 385 lobatc inner ends of the chambers forming a star-shaped umbiUcus; periphery smooth; edge subangular in early part of the test becoming broadly rounded and shouldered on the dorsal side in most of the last whorl of the test; six chambers in the last whorl, enlarging rapidly in thickness, the last chamber constituting nearly a third of the test on the ventral side; sutures distinct, marked by a filling of clear shell material, curved on both sides of the test, flush with the surface except between the last two chambers where it is depressed; aperture an arched opening near the umbilicus on the ventral side of the test, partially covered by a thin, plate-like extension of the chamber. Diameter, 0.64 by 0.80 millimeter; thickness 0.46 milhmeter. Holotype No. 3899, locahty A86, Cibao formation. Rare at A86 of the Cibao formation. Discorbis pelliculatus NEW SPECIES Plate 21, figures 6a-c Test small, nearly circular in side view; dorsal side low conical with concave slopes and prominent umbo; ventral side slightly convex to flat, not concave, with low, median umbo; periphery smooth, not lobulate; edge angled but neither sharp nor carinate; chambers very broad, 4 in last whorl, the last comprising more than one-third of the circumference of the test, each ventral chamber with an umbonal extension, similar to those seen in Tetrataxis; whorls about three, the early ones obscure ; dorsal sutures curved and tangential to the earlier whorl, narrow and flush with the surface, the last one or two slightly depressed, only those of the last whorl visible; ventral sutures obscure, consisting of two outward curves and one inward curve, narrow and flush with the surface; wall very finely perforate, as seen after removal of the pellicle by acid ; surface matt mostly covered on both sides by a thin smooth layer or pelhcle of opaque shell material, thickest on the dorsal umbo; aperture a small, thin arch in the front, outer curve of the ventral part of the last chamber, not under the umbilical exten- sion. Diameter, 0.69 millimeter; thickness, 0.30 milhmeter. Holotype No. 4141, locahty A21, Quebradillas formation. Common at locality A21 of the Quebradillas formation. This species is obviously similar to several forms as yet included in D. orbicularis (Terquem), particularly the Florida Miocene form (1932. Fla. Geol. Surv. Bull. 9), but it differs from more typical forms of D. orbicularis (such as Brady 1884. Rep. Voy. Challenger Zool. 9: pi. 88, fig. 5, from the Bermudas, and Cushman 1931. Bull. U. S. Nat. Mus. 104 (8): pi. 6, fig. 3, from Jamaica) in lacking the concave 380 SCIENTIFIC SURVEY OF PORTO RICO ventral side, the cariiiate edge ajid the eoarse perforations, and in the presence of the umbos and the pellicle of secondarj' shell material. Discorbis subaraucanus Cusliinan Plate 21, figures 2a-c Discorbis subaraucana Cuslunan (1922) Carnegie Inst. Wash. Publ. 311: 41. pi. 7, li^:^^. 1, 2. Recent, Tortugas.— Cole & Ciillespk- (1930) BuU. Am. Paleont. 15 (57B) : 11. pi. 3. flg.s. 2, 3. Middle Oligocene. Meson, Mexico. — Cusiiman (1935) U. S. Geol. Surv. Prof. Pap. 181: 43. pi. 18, figs. la-c. Upper Eocene, Alabama. Test planoconvex, very slightly convex on the dorsal side, and nearly flat on the ventral side except for the last chamber which is somewhat inflated and the concave depression in the umbilical region; edge sub- acute; two and a half whorls in the test, 6 to 7 chambers in the last whorl, early chambers enlarging gradually in size, last few chambers enlarging more rapidly, the last chamber in the adult form being arcuate on the dorsal side, and constituting nearly }i of the ventral side of the test; sutures on both .sides, and early part of spiral suture on the dorsal side marked by clear shell material, last few sutures more narrowly limbate; sutures obliquely curved on the dorsal side, flush with the surface except for the last few which are slightly depressed, sutures on the ventral side curved, more so in the earlier part of the test; wall finely but conspicuously perforate; aperture a .slit at the base of the last chamber on the ventral side extending from the umbilicus to near the periphery, with a slight lip. Diameter, 0.40 by 0.45 milHmeter; thickness, 0.16 millimeter. Plesiotype No. 3900, locality L5C, San Sebastian formation. Rare at L2C and L3C; common at L5C of the San Sebastian forma- tion. Abundant at A70 of the Los Puertos formation. Common at P258 of the Ponce formation. EPISTOMARIA Galloway 1933 Epistomaria pontifera NEW SPECIES Pl.\te 24, I'iGUKES 2a-c Discvrbina polustomelhides Heron-Allen & Earland (1915) Tians. Zool. Soc. London 20 (2): pi. 52, flgp. 19-23. Recent, Keriiuba Arcliipelago. Test rotaloid, convex on the dorsal side with depressed spire, concave on ihv, ventral si(l(>, broadly oval in side vi(>\v; pcMiphery lobulate; back smoothly rounded; chambers 7 to 8 in th(« last whorl, inflated, increasing rapidly in size as added; whorls about 2; early w^horls on the dorsal side separated from the last whorl by a depressed spiral suture; on tlu> ventral side secondary shell material forms GALLOWAY AND HEMINWAY, TEUriARY FORAMINIFERA 387 triangular places over part of each chamber converging at the um- bilicus; dorsal sutures nearly radial, distinct, depressed, marked by clear shell material; ventral sutures depressed, radial, narrow between the secondary deposits; between the chambers on the back of the test there are one or two retral processes or septal bridges similar to those in Elphidium; wall coarsely and plainly perforate, the perfora- tions continuing through the ventral secondary deposits; aperture a wide slit at the base of the septal face, usually closed by secondary material, and elongate secondary apertures between the chambers on both sides near the periphery, the accessory apertures being outlined by thin bands of clear shell material ; there are no accessory apertures at the outer edges of the triangular umbilical plates. Diameter, 0.87 by 0.62 millimeter; thickness, 0.33 miUimeter, for the medium-sized figured specimen. Holotype No. 4115, locality P258, Ponce formation. Common at P258 of the Ponce formation. This species differs from the Recent Arctic species, E. rimosa (Parker and Jones) (1865. Phil. Trans. Roy. Soc. London 155) in having septal bridges and lacking the openings at the outer ends of the triangular umbilical plates. The septal bridges are similar to those in D. polijstomelloides (Parker and Jones), but are fewer, there are fewer chambers, and the deposits are thinner and smoother. Our form seems to be identical with the form identified with Dis- corbina polystomelloides by Heron-Allen and Earland, from the Kerimba Archipelago, in an extraordinary provincial fauna. ANOMALINA d'Orbigny 1826 Anomalina alazanensis Nuttall Plate 22, figures la-c Anomalina alazanensis Nuttall (1932) Jour. Paleont. 6: 31. pi. 8, flgs. 4, 8, 9. Lower Oligo- cene, Mexico. — Palmer & Bermudez (1936) Mem. Soc. Cubana Hist. Nat. 10: 311. pi. 19, flgs. 11-13. Lower Oligocene, Cuba. Test plano-convex, dorsal side slightly convex, ventral side flat or slightly concave; periphery smooth, not lobate; edge rounded; cham- bers more involute on the ventral side, dorsal side showing 23/^ whorls, ventral side showing about IJ^ whorls; about 14 chambers in the last whorl, enlarging very slowly in size; sutures conspicuous, limbate, slightly raised, gently curved on both sides of the test; spiral sutures not raised; wall coarsely perforate; aperture a narrow arch at the base of the last chamber on the peripher}' and extending about one chamber 388 SCIENTIFIC SURVEY OF PORTO RICO length onto the dorsal side. Diameter, 0.67 by 0.55 millimeter; thickness, 0.20 millimeter. Plesiotype No. 3866, locahty P3, Ponce formation. Very rare at P3 and P254 of the Ponce formation. Anomalina nucleata (Seguenza) Plate 22, figures 2a-c Truncaiulina nucleata Seguenza (1880) Atti R. Accad. Lincei III. 6: 04. pi. 7, fig. 8. Lower Miocene, Italy. Truncaiulina trinitatensis Nuttall (1928) Quart. Jour. Geol. Soc. London 84: 97. pi. 7, figs. 3, 6, 6. Upper Eocene to Miocene, Trinidad. Cibicides nucleata Galloway & Morrey (1929) Bull. Am. Paleont. 15 (55): 31. pi. 4, fig. 9. Lower Oligocene, Ecuador. Cibicides trinitatensis Nuttall (1932) Jour. Paleont. 6: 33. pi. 7, fig. 9. Lower Oligocene, Mexico. — Hadley (1934) Bull. Am. Paleont. 20 (70A) : 29. pi. 4, figs. 10, 11. Upper Eocene and Oligocene, Cuba. Anomalina nucleata Coryell & Rivero (1940) Jour. Paleont. 14: 334. pi. 44, figs. 2a-c. Middle Miocene, Haiti. Test subcircular in side view, biconvex and biumbonate, nearly planispiral, dorsal side moderately convex, ventral side strongly con- vex; periphery smooth, not lobulate; edge sharply to evenly rounded; involute on both sides, more so on the ventral side; on the dorsal side the last whorl is wide, the early whorls covered by a thick, flat or convex boss of clear shell material; chambers 12 to 15 in the last whorl, enlarging gradually as added; dorsal sutures gently curved, strongly raised and marked by clear shell material between the early chambers of the last whorl, somewhat depressed between the last few chambers; ventral sutures gently curved, flush or slightly raised, merging into a thin or thick layer of transparent secondary material covering the early whorls and making an umbo; wall coarsely and conspicuously perforate, some of the pores showng in the limbate sutures and in the bosses; aperture a low arch at the base of the septal face on the inner periphery and extending a short distance toward the ventral umbo and dorsally for a distance of several chambers between the last two whorls, with slight upper lip. Diameter, 0.73 millimeter; thickness, 0.37 miUimeter. Plesiotype No. 4041, locality P254, Ponce formation. Rare at P251 and P254 of the Ponce formation, and at A93 of the Quebradillas formation. We include this form in Anomalina because of the nearly planispiral coiling and because the spire on neither side is completely covered by the last whorl. Other forms similar or identical with this species are Anomalina umbonata Cushman (in Howe 1939. La. Dept. Cons. Geol. Bull. 14) and Anomalina nokmi Hedberg (1937. Jour. Paleont. 11). GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 389 There is considerable variation in the amount of sutural Umbation and thickness of the umbos in this species (or these species), depending more on the ontogenetic age of the specimens than on the geologic age. They all occur between the middle Eocene and middle Miocene. Anomalina pompilioides NEW SPECIES Plate 22, figures 3a-c Anomalina grosserugosa Cole (not Gtinibel) (1928) Bull. Am. Paleont. 14 (53): 18. pi. 1, figs. 16, 17. Upper Eocene. Chapapote formation, Mexico. — Nuttall (not Giimbel) (1928) Quart. Jour. Geol. Soc. London 84: 99. pi. 7. fig. 18 (not 19). Upper Eocene and lower Miocene, Trinidad. Test small for the genus, nearly bilaterally symmetrical, subcircular in side view; periphery lobate; broadly elliptical in edge view, back broadly rounded; involute on ventral side, not completely involute on dorsal side; chambers inflated, rather large, regularly and gradually enlarging, 5 to 6 in the last whorl; sutures radial, nearly straight, depressed; wall coarsely perforate; aperture a low arch on the inner margin of the last septal face on the periphery; apertural face convex, lunate in shape, slightly more than twice as wide as its median height. Diameter, 0.32 by 0.39 miUimeter; thickness, 0.23 millimeter. Holotype No. 3867, locahty A86, Cibao formation. Rare at A86 of the Cibao formation. This species differs from A. grosserugosa (Gtimbel) (1870. Abh. k. bay. Akad. Wiss. Miinchen Math. Phys. CI. 10) in its greater thick- ness and fewer chambers. As Thalmann pointed out in 1932, Brady's specimens in the Challen- ger Report are not true A. grosserugosa (Gtimbel), but neither are they the same as this Porto Rican species. Brady's form has more cham- bers in the last whorl, i. e. eight. CYCLOLOCULINA Heron-Allen & Earland 1908 Cycloloculina cubensis Cushman & Bermudez Plate 21, figures 3a, b, and 4 Cycloloculina cubensis Cushman & Bermudez (1936) Contr. Cushman Lab. Foram. Res. 12: 61. pi. 11, figs. 15, 16. Eocene, Cuba. Test discoidal, compressed, the two sides much alike; edge narrowly rounded; about 5 rapidly enlarging chambers in the early part of the test, followed by 3 to 6 annular chambers; sutures indistinct except near the periphery where they are slightly depressed; surface finely granular and ornamented by scale-like additions of secondary tissue, most prominent on the last chamber, and radial depressions, the 390 SCIENTIFIC SURVEY OF PORTO RICO scales and radial depressions of each chamber independent of those on adjacent chambers, giving a creniilato appearance to the inner margins of the chambers; aperture not observed. Diameter, 0.57 millimeter; thickness, 0.12 millimeter. Plesiotypes No. 3888, locality L3C, San Sebastian formation, and No. 3886, locality A79, Los Puertos formation. Rare at L3C; common at L4C of the San Sebastian formation. Rare at P254, P255, and P431 of the Ponce formation. Common at A79, Los Puertos formation. CIBICIDES Mont fort 1808 Key to the Porto Rican Species A. Ventral side umbilicate, without umbo of secondary material B. Involute nearly to spire on dorsal side C. Test oval; later chambers gradually lengthening C. chnctau'cnsis, page 391. CC. Test round; later chambers rapidly lengthening C. americanus, page 390. BB. Not involute on dorsal side C. Test round; later chambers gradually enlarging D. Without dorsal umbo C. inexicanus, page 394. DD. With dorsal umbo C io, page 392. CC. Test oval; later chambers rapidly enlaiging D. Periphery strongly lobate C. lobatus, page 393. DD. Periphery slightly lubate C. scalenus, page 39(1. AA. Ventral side strongly umbonate with secondary material B. Last whorl wide, tending to be involute dorsally C. Dorsal secondaiy ornamentation weak C. pseudoungerianus, page .39."). CC. Dorsal secondary ornamentation strong.. C. flcrridanm, page 392. BB. Last whorl narrow, not involute dorsally C. Spiral suture not raised; dorsal deposit thin D. Whorls less than 3; pores very coarse C. perhicida, page 394. DD. Whorls more than 3; pores coarse. C. sinistralis, page 39(1. CC. Spiral suture raised C. sptroliyvbatns, page 397. Cibicides americanus (Cushnian) Plate 24, figures 3a-c Truncatulina americana Cushman (1918) U. S. Geol. Sur\ . lUill. 676: 03. pi. 20, flgs. 2, 3, pi. 21, fig. 1. Miocene, Atlantic coa.stal plain, IT. S. Cibicides americanus Cole & Gillespie (1930) Bull. Am. Paleont. 15 (57B) : 14. pi. 4, fig. 4. Meson formation, Mexico, middle Oligocone. — Cushman & Caliill (VXV.i) U. S. Geol. Surv. Prof. Pap. 175A: 34. pi. 13, figs. 2a-c. Miocene and Oligocene of Atlantic coastal plain; late Tertiary of California, Panama, and northern South America. CALLOWAY A\D IIEMINWAY, TERTIARY FORAMINIFERA 391 Tost small for the genus, planoconvex, dorsal side nearly flat, ventral side moderately convex; edge subacute; periphery smooth except for the last few chambers which are slightly lobulate; 8 to 11 chambers in the last adult whorl, rather rapidly increasing in size; in most specimens the last few chambers do not extend to the umbilicus either dorsally or ventrally; sutures distinct, slightly limbate on both sides in the early chambers, slightly depressed between later chambers; wall smooth, coarsely perforate; aperture at the base of the septal face extending onto the dorsal side only one chamber width and ex- tending to the umbilicus on the ventral side; aperture with slight lip. Diameter of figured specimen, 0.29 by 0.38 miUimeter; thickness, 0.13 millimeter. Maximum diameter, 0.50 millimeter. Plesiotype No. 3952, locality LlC, San Sebastian formation. Common at LlC and L2C of the San Sebastian formation. Rare at A86 of the Cibao formation. Cibicides choctawensis Cushman & McGIamery Plate 23, figures la-c Cibicides choctawejisis Cushman & McGIamery (1938) U. S. Gool. Surv. Prof. Pap. 189D: 111. pi. 28, flg. 6. Oligocene, Alabama. Test small, subcircular or broadly oval in dorsal view, planoconvex, dorsal side flat or slightly concave and nearly or quite involute to the middle of the spire, ventral side very convex, with small umbilicus, with tiuncate base and nearly straight sides, increasing rapidly in thickness with successive chambers; edge sciuare to subacute, but not keeled; periphery smooth, not lobulate; chambers about 9 in the last whorl, increasing gradually in size, the last few increasing more rapidly in all dimensions; sutures on the dorsal side strongly curved, slightly limbate, flush with the surface but depressed near the center of the spire; ventral sutures very little curved, limbate, flush with the surface except the last few which are a little depressed; surface smooth; wall finely but conspicuously perforate; aperture a small arch at the base of the septal face on the inner periphery, with upper lip, con- tinuing on the dorsal side between the last four chambers and the previous whorl. Diameter, 0.37 miUimeter; thickness, 0.15 millimeter. Plesiotype No. 4117, locality A43a, Cibao formation. Common at A43a of the Cibao formation. This form is more involute on the dorsal side and is much more finely perforate than is true of typical Cibicides. 392 SCIENTIFIC SURVEY OF PORTO RICO Cibicides floridanus (Cushman) Plate 23, figures 2a-c Truncatulina floridana Cushman (1918) U. S. Geol. Surv. Bull. 676: 62. pi. 19, fig. 2. Mio- cene, Florida. — Nuttall (1928) Quart. Jour. Geol. Soc. London 84: 98. pi. 7, figs. 14, 16. Miocene, Trinidad. Truncatuhna lobatula ornata Cushman (1918) U. S. Geol. Surv. Bull. 676: 61. pi. 18, figs. 1, 2. Miocene, Florida, Virginia. Cibicides floridana Cushman (1931) Bull. U. S. Nat. Mus. 104 (8): 122. pi. 23, figs. 3-5. Recent, off Florida; (1930) Fla. Geol. Surv. Bull. 4: 61. pi. 12, fig. 3. Miocene, Florida. Cibicides cf. floridana Palmer & Bermudez (1936) Mem. Soc. Cubana Hist. Nat. 10: 315. pi. 20, figs. 1-3. Lower Oligocene, Cuba. Cibicides floridanus Coryell & Rivero (1940) Jour. Paleont. 14: 334. pi. 44, figs. lOa-c. Mid- die Miocene, Haiti. Test small, subcirciilar in side view, equally biconvex or rather flat dorsally and strongly convex ventrally; edge subacute; periphery smooth, not lobulate, with border of clear shell material; the last whorl broad dorsally, tending to become involute, the early whorls covered with a thick, secondary deposit; ventral side with strong umbo of hyaline secondary material; chambers 10 to 12 in the last whorl, increasing slowly in length and thickness but more rapidly in width; dorsal sutures gently curved, limbate and raised, except for the last few which may be depressed; ventral sutures gently curved, limbate and raised, merging with the umbo and the border; wall coarsely perforate, the pores enlarged by the secondary material on the dorsal side; aperture a small arch, with upper lip, at the base of the septal face on the inner periphery, extending dorsally for a distance of one or two chambers. Diameter, 0.45 millimeter; thickness, 0.2 millimeter. Plesiotype No. 4118, locality A86, Cibao formation. Common at P2, P4, and P254 of the Ponce formation. Abundant at A86 of the Cibao formation. Common at A20 and A21 of the Quebradillas formation. This species is characterized by the strong secondary deposits in the form of covered spire and raised spiral and radial sutures. It has the wide last whorl which is partially involute on the dorsal side, also seen in C. pseudodoungerianus and C. io. Cibicides io Cushman Plate 22, figures 4a-c Cibicides pseudoungeriana io Cushman (1931) Bull. U. S. Nat. Mus. 104 (8): 125. pi. 23, figs. 1, 2. Recent, off Florida. Cibicides io Coryell & Rivero (1940) Jour. Paleont. 14: 334. pi. 44, fig. 11. Miocene, Haiti. Test subcircular in side view, nearly flat on the dorsal side, convex on the ventral side; periphery not lobulate; edge sharply rounded to GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 393 broadly rounded; whorls few, about two, the last whorl on the dorsal side increasing rapidly in width, considerably involute, the early whorls covered by secondary material, separated by a groove from the limbate and raised spiral suture; ventral side with small umbilicus, but without umbo of secondary material; chambers about 10 in the last whorl, enlarging gradually in length but rapidly in width as added; sutures on the dorsal side slightly curved, limbate and raised, except the last ones which may be slightly depressed; ventral sutures very little curved, nearly radial, narrowly limbate and flush with the surface or slightly depressed; aperture a low, broad arch at the base of the septal face, on the inner periphery, extending sHghtly ventrally and dorsally, with narrow upper lip. Diameter, 0.56 millimeter; thickness, 0.22 milHmeter. Plesiotype No. 4123, locaHty A21, Quebradillas formation. Rare at A21 of the Quebradillas formation. This species lacks the ventral umbo and raised sutures of C. flori- daniis, but has a dorsal umbo. Cibicides lobatus (d'Orbigny) Plate 24, figures 4a-c TruncatuHna lobata d'Orbigny (1839) in Barker, Webb & Berthelot, Hist. Nat. lies Canaries 2 (2): Foraminiferes 134. pi. 2, figs. 22-24. Recent, Canaries. TruncatuHna lobatula d'Orbigny (1846) Foram. Foss. Vienne 168. pi. 9, figs. 18-23. Middle Miocene, Vienna. — Brady (1884) Rep. Voy. Challenger Zool. 9: 660. pi. 92, fig. 10, pi. 93. flg. 1 (not 4 and 5). Recent, all latitudes. — Cushman (1918) U. S. Geol. Surv. Bull. 676: 16. pi. 1. fig. 10. Pliocene, North Carolina. Cibicides lobatula Cushman (1931) Bull. U. S. Nat. Mus. 104 (8): 118. pi. 21, figs. 3a-c. Recent, north Atlantic. Cibicides lobatulus Cushman (1935) U. S. Geol. Surv. Prof. Pap. 181: 52. pi. 22, fig. 4 (not 5 and 6). Upper Eocene, southeastern U. S. — Cole & Gillespie (1930) Bull. Am. Paleont. 15 (57B) : 15. pi. 4, fig. 2. Middle OUgocene, Mexico. Test of average size for the genus, planoconvex, flat or concave on the dorsal side, convex on the ventral side; periphery only slightly lobate; edge acute; chambers somewhat inflated, increasing gradually in size, 4 to 8 in the last whorl; sutures flush with the surface, curved and marked by clear shell material on the dorsal side, on the ventral side depressed, nearly radial; wall coarsely perforate; aperture on the periphery with slight lip, and extending both dorsally and ventrally along the base of the last chamber. Diameter of figured specimen, 0.65 by 0.74 millimeter; thickness, 0.20 millimeter. Plesiotype No. 3884, locality L4C, San Sebatian formation. Rare at LlC, L4C, and L5C of the San Sebastian formation, and at A43a and A86 of the Cibao formation. Abundant at A79 of the Los Puertos formation. Rare at A21 and r64 of the Quebradillas forma- 394 SCIENTIFIC SURVEY OF PORTO RICO tioii. Karc at P3, P251, and P254; common at P2o8, P431, and P432 of the Ponce formation. C. lohata (d'Orbigny) is distinct from C. lobatulus (Walker and Jacob), the type figure of which is conspicuously lobate. Cibicides mexicanus Nuttfill Plate 22, figures 5a-c CiMcides mexicana Nuttall (1932) Jour. Paleont. 6: 33. pi. 9, fig.s. 7-9. Lower Oligocene, Mexico. Test plano-convex, dorsal side flattened except for a moderate con- vexity of the early whorls, the last whorl being flat or even slightly concave, ventral side strongly convex, hemispherical; edge subacute, shouldered; early whorls indistinct on the dorsal side due to a thin layer of secondary shell material; 9 to 11 chambers in the last whorl, enlarging gradually in size; chambers involute ventrally, with .flight umbilical depression; sutures gently curved, somewhat oblique on the dorsal side, gently curved radial on the ventral side, marked by clear shell material on both sides of the test, flush with the surface or slightly depressed on the dorsal side, flush or slightly raised on the ventral side; wall coarsely perforate; aperture an elongate slit at the base of the septal face running from the periphery l(>ss than half way to the umbilicus, and extending on the dorsal side between the last two whorls for a distance of about 4 chambers, with upper lip. Diameter, 0.95 millimeter; thickness, 0.6 millimeter. Plesiotype No. 3924, locality P3, Ponce formation. Abundant at P2 and P255; rare at P2ol and P3 of the Ponce formation. Cibicides perlucidus Nuttall Pl.\te 23, FiGUKES 4a-c Cibicides perlucido Nuttall (1932) Jour. Pali out. 6: 33. pi. S, ilns. 10-12. Lower Oligooone, Mexico. Test subcircular in side view, unequally biconvex, dorsal side slightly elevated, ventral side strongly convex; edge subacute to sharp- ly rounded; periphery smooth, not lobulate; chambers 9 to 11 in the last whorl, enlarging gradually as added; whorls fcnv, only two or a little more, the last not involut(> beyond the ix'riphery of the previous whorl, the early whorls slightly obscured by a thin, smooth secondary deposit; spiral suture flush, bordered by a thick band of clear shell material; dorsal sutures narrowly limbate and flush with the surface; ventral sutures narrowly limbate and slightly depressed, merging GALLOWAY AND HEM I NW AY, TERTIARY FORAMINIFERA 395 with the central boss of clear shell material; wall very coarsely and conspicuously perforate, the wall material being translucent; aperture a low arch at the base of the septal face, on the inner periphery, with slight lip, and extending a short distance ventrally and dorsally. Diameter, 0.77 millimeter; thickness, 0.4 millimeter. Plesiotype No. 4119, locahty P251, Ponce formation. Rare at P251 of the Ponce formation. This species differs from C. flondanus and C. 'pseudoungerianus in the narrower last whorl which does not tend to be involute, and in lacking the thick, secondary deposit on the dorsal side. It has fewer whorls and is deeper ventrally than C. 'pseudoungerianus sinistralis and C. 'pseudoungerianus evolutus. C. mexicanus is hemispherical ventrally, whereas C. perlucidus is conical. Cibicides pseudoungerianus (Ciishman) Plate 23, figures 5a-c Truncatulina pseudoungeriana Cushman (1922) U. S. Geol. Surv. Prof. Pap. 129E: 97. pi. 20, flg. 9. Middle Oligocene, Mississippi. Cibicides pseudoungeriana Cu.shman (1931) Bull. U. S. Nat. Mus. 104 (8): 123. pi. 22, figs. 3-7. Recent, Atlantic. Test small, biconvex, slightly more convex on the ventral side, subcircular in side view; periphery not lobulate, with border of clear shell material; edge angled to sharply rounded; whorls 2 to 3, usually coiled dextrally, the last one only slightly overlapping earlier whorls; early whorls on the dorsal side covered by a thin, smooth layer of shell material; ventral side with large umbo of clear shell material; chambers 9 to 11 in the last whoi'l, gradually enlarging in size as added; sutures on both sides gently curved, narrowly limbate, flush with the surface excepting on the ventral side between the last few chambers, where they are .shghtly depressed; aperture a small arch, with upper lip, at the base of the septal face on the inner periphery, continuing a short distance ventrally and for a distance of two or three chambers dorsally. Diameter, 0.43 millimeter; thickness, 0.23 milli- meter. Plesiotype No. 4120, locahty A79, Los Puertos formation. Rare at A79 of the Los Puertos formation. This species is not well defined and not definitel.y recognizable, since several different forms have been included in it by its author. It is scarcely distinguishable from C. floridanus. As we distinguish the species, C. pseudoungerianus has little limbation and thin secondary deposits; C. floridanus has strong limbation on both sides, thick 396 SCIENTIFIC SURVEY OF PORTO RICO secondary deposit on the dorsal side in which the mural pores are enlarged. The difference in number of chambers and relative convexity is not significant. Cibicides scalenus NEW SPECIES Plate 23, figures 3a-c Test very small, oval in side view, dorsal side flat or slightly convex, ventral side very convex, with small umbilicus; whorls about 2; cham- bers about 8 in the last whorl, increasing gradually up to the last few chambers when the chambers increase very rapidly in all dimensions; edge sharp, but not keeled; periphery smooth up to the last few chambers when it becomes lobulate, bordered by clear shell material ; sutures on the dorsal side gently curved, limbate but not raised, the spiral suture being both limbate and raised; ventral sutures strongly curved, the outer ends limbate but not raised; wall conspicuously perforate; aperture a small arch on the inner periphery at the base of the last septal face, with upper lip, extending on the dorsal side for a distance of about two chambers. Diameter, 0.32 miUimeter; thickness, 0.16 millimeter. Plesiotype No. 4121, locality A43a, Cibao formation. Rare at 43a of the Cibao formation. The Porto Rican form resembles C. westi Howe (1939. La. Dept. Cons. Bull. 14), but C. scalenus is less conical on the ventral side, and the ventral sutures are not strongly sigmoid. Cibicides sinistralis Coryell & Rivero Plate 24, figures 5a-c Cibicides sinistralis Coryell & Rivero (1940) Jour. Paleont. 14: 335. pi. 44. figs. 12a-c. Mid- dle Miocene, Haiti. Test subcircular in side view, nearly equally biconvex, coiled sinistrally; periphery not lobulate; edge subangular; whorls 33^, gradually enlarging, not overlapping the earlier whorls on the dorsal side; chambers 10 to 11, gradually enlarging; dorsal side covered by a thin, transparent secondary deposit; ventral side \x\th strong umbo of clear shell material; dorsal sutures curved, slightly limbate but not raised; ventral sutures slightly curved, the later one sometimes sig- moid, narrowly limbate but flush with the surface; wall coarsely but not conspicuously perforate; aperture a small arch at the base of the septal face on the inner periphery, with upper lip, extending a very short distance ventrally and extending dorsally for a distance of 2 or 3 chambers. Diameter, 0.43 millimeter; thickness, 0.23 millimeter. GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 397 Plesiotype No. 4101, locality A86, Cibao formation. Rare at A86 of the Cibao formation. This species differs from C. floridanus and C. pseudoungerianus in having a narrower last whorl which does not overlap earlier whorls on the dorsal side. It differs from C. perlucida in the fewer whorls and less conspicuous pores. It differs from C. pseudoungerianus evolutus (1935. Contr. Cushman Lab. Foram. Res. 11: 64. pi. 9, fig. 11) only in coiUng to the left. The sinistral coiling seems to be of Uttle signifi- cance, in view of the fact that many species of Foraminifera coil normally both sinistrally and dextrally. It must be admitted, how- ever, that there are races, varieties or even species, which coil in only one direction for a given locality and geologic age. Cibicides spirolimbatus NEW SPECIES Plate 25, figures la-c Test of medium size for the genus, subcircular in side view, biconvex and biumbonate, the ventral side shghtly deeper; periphery not lobu- late; edge sharply rounded, not keeled; whorls about 4, unusually narrow for the genus, not increasing much in width as growth proceeds, not overlapping on the dorsal side; about 12 chambers in the last whorl, gradually increasing in size as added; dorsal sutures slightly curved, moderately obhque, more and more hmbate toward the center of the test, but not much raised, the spiral suture limbate and raised with clear shell material; the growth of the secondary material on the sutures obscures the spire especially toward the center but leaving the mural perforations in the reduced rhomboidal areas corresponding to the chambers; sutures on the ventral side shghtly curved, narrowly limbate and flush with the surface, merging into the umbo of clear shell material; aperture a low arch just below the periphery with upper hp, extending on the dorsal side between the whorls for a dis- tance of one or two chambers. Diameter, 0.87 milhmeter; thickness, 0.40 millimeter. Holotype No. 4122, locality A93, Quebradillas formation. Rare at A93 of the Quebradillas formation. This species differs from C. mundula (Brady, Parker and Jones) (1890. Trans. Zool. Soc. London 12) in being ventrally umbonate, in the curved sutures and the aperture not on the inner periphery. The narrow whorls and spiral limbation are characteristic. 398 SCIENTIFIC SURVEY OF I'ORTO RICO PLANULINA d'Orbigny 1826 Planulina crassa new species Plate 25, figures 2a-c Test nearly circular in side view, compressed in edge view yet rather thick, dorsal side flat or slightly concave, ventral side flat or slightly convex; back flat or slightly concave, a Httle oblique to the two sides; whorls 2 to 3, scarcely at all involute on th(^ ventral side, slightly involute on the dorsal side; chambers 9 to 10 in the last whorl, increasing gradually in size as added; sutures on the dorsal side much curved and strongly raised, merging with the raised border; ventral sutures strongly curved and raised; wall coarsely perforate, the ventral surface strongly granulate between the sutures and on the sutures, and finely granulate on the dorsal surface over the early whorls; aperture at the base of the septal face on the inner periphery, with upper lip, not extending onto the ventral side but extending onto the dorsal side between the last two whorls for a distance of one or two chambers. Diameter, 0.45 millimeter; thickness, 0.13 millimeter. Holotype No. 4065, locahty A21, Quebradillas formation. Abundant at A21; rare at A15; common at A20 and A93 of the Quebradillas formation. This species differs from P. ariminensis d'Orbigny in that it is nuich thicker and the chambers are narrower in proportion to their length. Planulina depressa (d'Orbigny) Plate 25, figures 3a-c Tnmcatulimi depressa d'Orbigny (1839) Voy. Amer. Morid. 5: Foianuniferes 39. pi. G, flgs. 4-G. Recent, oCf South America. Planulina depressa Cushnian (1930) Fla. Gool. Surv. Bull. 4: CO. pi. 12. fig. 2. Miocene. Florida. Test large, subcircular in side view, much compn^sscd, concavo- convex to i)lano-convex, ventral side moderately convex but without an umbilical boss of secondary shell material; edge sharply angular; periphciy lobulate, more strongly so in th(> last half whorl, with narrow, limbatc bordci-; some of the earlier whoi'ls show on both sides of the test, the; early dorsal spire covered with a thin, granular, .secondary depo.sit; chambers short and wide, a little inflated on the ventral .side, enlarging mor(> rai)idly in width than in length or thickness, about 9 in the last whorl; ventral sutures most distinct between the chambers of the last half whorl where they are conspicuously depressed, nearly radial near the center of the test, sharply curved near the piM-iphery; dor.sal sutures strongly curvc^d, flush with the surface or slightly de- GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 399 pressed, slightly limbate on the outer end where the Knibation merges into the bolder; wall moderately coarsely perforate; aperture a small, high arch, Avith strong upper lip, at the base of the septal face on the inner periphery, extending onto the dorsal side for a distance of 3 or 4 chambers. Diameter, 1.38 millimeters; thickness, 0.26 millimeter. Plesiotype No. 4124, locality P255, Ponce formation. Rare at P255 of the Ponce formation. Planulina marialana Hadley Plate 25, figuers 4a-c Planulina wucllcrslorfi Nuttall (not Schwager) (1932) Jour. Paleont. 6: 31. pi. 4, figs. 14, 15. Lower Oligocene, Mexico. Planulina marialana Hadley (1934) Bull. Am. Paleont. 20 (70A): 27. pi. 4. figs. 4-6. Oligo- cene, probably upper, Cuba. — Palmer & Bermudez (1936) Mom. Soc. Cubana Hist. Nat. 10: 313. pi. 20, figs. 10-12. Lower Oligocene, Cuba. Test large, nearly circular, biconvex, much compressed; periphery smooth, except for the last few chambers; edge angled, with narrow peripheral keel of clear shell material, more distinct on the earlier chambers of the last whorl; dorsal side of test showing 2>^ to 3 rapidly widening whorls, ventral side involute nearly to umbilicus; chambers short and wide, 10 to 12 in the final whorl of the adult; dorsal sutures broadly curved, limbate, usually raised with the raised limbation more conspicuous on the early whorls; ventral sutures broadly curved, the later ones sigmoid, limbate, usually raised; the largest specimens umbilicate; wall coarsely perforate; aperture at the base of the last chamber on the inner periphery, extending a short distance onto the dorsal side between the last two whorls. Diameter, 1.52 millimeter; thickness, 0.25 millimeter. Plesiotype No. 4125, locality P2, Ponce formation. Rare at P251; common at P3 and P255; abundant at P2 and P254 of the Ponce formation. It is quite possible that the forms figured by Hadley (umbonate, 9 chambers) and that figured by Palmer and Bermudez (umbilicate, 12 to 14 chambers) represent two different species, both of which are represented in the Porto Rican material. Planulina mexicana Cushman Plate 26, figures 3a-c Planulina mexicana Cushman (1927) Contr. Cushman Lab. Foram. Res. 3: 113. pi. 23, figs. 5a, b. Alazan, upper Eocene and lower Oligocene. Mexico. — Cole & Ponton (1930) Fla. Cieol. Surv. Bull. 5: 47. pi. 6, flg. 2. Lower Oligocene, Florida. — Nuttall (1932) Jour. Paleont. 6: 31. pi. 7, flg. 7. Lower Oligocene, Alazan, Mexico. — Hadley (1934) Bull. Am. Paleont. 20 (70A) : 28. pi. 4, flg. 12. Oligocene, Cuba. 400 SCIENTIFIC SURVEY OF PORTO RICO Planulina cocoaensis cooperensis Cushman (1933) Contr. Cushman Lab. Foram. Res. 9: 20. pi. 2, flg. 12; (1935) U. S. Geol. Surv. Prof. Pap. 181: 52. pi. 22, fig. 8. Upper Eocene, South Carolina. Test large, much compressed, complanate, sides flattened; edge narrowly rounded, not keeled; periphery slightly lobulate; earlj^ whorls visible on both sides of the test, becoming less involute with age, the final half whorl just touching the preceding whorl; chambers short and evenly curved, 7 to 8 in the last whorl; sutures distinct, sharply curved, limbate, flush with the surface or very slightly raised except between the last few chambers where they are depressed; wall coarsely perfo- rate ; aperture a low arch with slight lip at the base of the last chamber extending on the dorsal side about half a chamber length. Diameter, 0.82 by 1.04 millimeter; thickness, 0.11 millimeter. Plesiotype No. 4066, locality P254, Ponce formation. Rare at P254 of the Ponce formation. This is one of the least involute species of the genus, the complete spire being visible on both sides of the test. Planulina zigzag NEW SPECIES Plate 26, figures 4a-c Test oval in side view, much compressed, nearly planispiral, the dorsal side flat, or with slight spiral depression inside the last whorl, rarely slightly convex, as in the specimen figured; ventral side shghtly convex, with large, shallow umbilical depression; edge truncate or slightly concave, with sharp, continuous keel of clear shell material extending from the dorsal side, the ventral side with discontinuous keel formed by the outer edges of the sutural limbation; periphery smooth, formed by the dorsal keel; dorsal sutures nearly straight, radial, limbate and raised, the limbation continuing in a zigzag line from chamber to chamber, just inside the carina, the spiral suture limbate but not raised, sometimes depressed; whorls about 2, only slightly involute on each side; eight chambers in the last whorl, en- larging rapidly, not inflated; ventral sutures limbate and strongly raised, straight, radial, the raised limbation continuing in a con- spicuous zigzag line around the outer edge of the chambers, somewhat inside the dorsal carina, not joining to form a continuous carina; the ladial and zigzag ridges on both sides of the test have fine, longitudinal striae on them; wall very coarsely and conspicuously perforate where the preservation is good; surface smooth, except for the raised ridges; septal face finely granulate; aperture a short slit at the base of the septal face, not continuing onto either dorsal or ventral side, with GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 401 slight upper lip. Diameter, 0.53 by 0.72 millimeter; thickness, 0.11 millimeter. Holotype No. 4126, locality A86, Cibao formation. Common at A86 of the Cibao formation. It is probably that P. zigzag is related to P. alavensis Palmer (1938. Mem. Soc. Cubana Hist. Nat. 12: 345, 346, text fig.), but P. zigzag differs in the zigzag line of the peripheral limbation and the absence of peripheral secondary apertures. CIBICIDELLA Cushman 1927 Cibicidella variabilis (d'Orbigny) Plate 24, figures la-c Truncatulina variabilis d'Orbigny (1826) Ann. Sci. Nat. 7: 279. no. 8; refers to 22 plates of illustrations by Soldani, 1789-1798. Recent, Mediterranean. — Parker, Jones & Brady (1871) Ann. Mag. Nat. Hist. IV. 8: 177. pi. 12, fig. 138. (One of Soldani's figures, on which d'Orbigny named his species.) — Terquem (1878) Mem. Soc. Gecl. France III. 1: 20. pi. 1, figs. 18-25. Upper Pliocene, Isle of Rhodes. Test large, ovate in side view; periphery lobulate; edge round; dorsal side irregularly concave; ventral side strongly convex and um- bilicate; chambers 6 or 7 in the last formed whorl, inflated ventrally and sometimes dorsally, enlarging rapidly in all dimensions, especially the last few chambers, the last few chambers distorted but still retaining their normal position in a rotaloid coil, sometimes tending to become evolute; dorsal sutures flush and limbate in the early test, depressed in the later portion; ventral sutures strongly depressed, nearly hnear and radial; wall coarsely perforate, about 0.01 milUmeter in diameter, with httle if any secondary material; aperture a single, high arch, with upper lip, on the inner periphery at the base of the septal face. Length, 1.36 millimeter; Avidth, 0.85 millimeter; thickness, 0.2 milhmeter. Some specimens are much larger and thicker. Plesiotype No. 4127, locahty L4C, San Sebastian formation. Common at L4C; rare at L6C of the San Sebastian formation. Our specimens are all much alike and not much removed from the ancestral Cibicides, and are like the figures in the references above. They are not like the figure in d'Orbigny's Canaries paper, nor hke the specimens from the Miocene of Florida (1932. Fla. Geol. Surv. Bull. 9: pi. 15). 402 SCIENTIFIC SURVEY OF PORTO RICO SIPHONINA Reuss 1850 Siphonina advena Cushman Plate 26, figures la-c Siphonina advena Cushman (1922) U. S. Geol. Surv. Prof. Pap. 129E: 98. pi. 22, figs. 1, 2. Middle OUgocene. Mississippi; (1927) Proc. U. S. Nat. Mus. 72 (20) : 7. pi. 1, fig. 7. Oligo- cene, gulf coast and Mexico. — Cole & Ponton (1930) Fla. Geol. Surv. Bull. 5: 43. pi. 11, figs. 7, 8. Lower OUgocene, Florida. Test of moderate size, subcircular to oval in side view, biconvex, the ventral side slightly more convex; periphery not lobiilate; edge subangular, not keeled; chambers in about 2 whorls, 4 to 5 in the last whorl, enlarging gradually; dorsal sutures obscure, slightly curved and oblique, flush with the surface, somewhat broadened with clear shell material; ventral sutures distinct, narrow, slightly curved, radial, a little depressed; wall very coarsely and conspicuously perforate, ex- cept over the early dorsal spire where the pores are obscure; aperture peripheral, slightly ventral, elliptical with short neck and phialine lip. Diameter, 0.42 milUmeter; thickness, 0.2 millimeter. Plesiotype No. 3988, locality A20, Quebradillas formation. Rare at LlC and L5C of the San Sebastian formation, and at A6 of the Lares formation. Common at A43a and A86 of the Cibao forma- tion. Rare at A79 of the Los Puertos formation. Rare at A15, A93, and F358; common at A20; abundant at A21 of the Quebradillas for- mation. Rare at P251 and P432 of the Ponce formation. Siphonina tenuicarinata Cushman Pl.\te 26, FIGURES 2a-c Siphonina tenuicarinata Cushman (1927) Jour. Paleont. 1: 1C6. pi. 26, flgs. 11, 12. Lower OUgocene, Alazan, Mexico; (1929) Contr. Cushman Lab. Forani. Res. 5: 100. pi. 14, fig. 9. Middle Tertiary, Venezuela and Trinidad. — Hedl)erg (1937) Jour. Paleont. 11: 679. pi. 92, fig. 4. Oligocene, Venezuela. — Coryell & Rivero (1940) Jour. Paleont. 14: 337. pi. 43, flgs. 22, 29. Middle Miocene, Haiti. Test asymmetrically coiled, biconvex, the ventral side slightly more so; edge acute with a wide keel of clear shell material, in which there are pores extending out from the edge of the chamber; whorls about 3; chambers about 5 in the last whorl, increasing uniformly in size, the last ones somewhat intlated on the ventral side; dorsal sutures nearly straight, tang(^ntial to the previous whorl, flush with the sur- face and very obscure; ventral sutures slightly curved, radial, flush with the surface and obscure excepting the last one or two, which may be slightly d(>pressed; wall moderately coarsely but conspicuously perforate; apciture slightly below the periphery on the ventral side. GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 403 elliptical, with very short neck and broad lip. Diameter, 0.6 milli- meter; thickness, 0.26 millimeter. Plesiotype No. 4128, locality P254, Ponce formation. Rare at P2 and P255; common at P254, Ponce formation. PLANORBULINELLA Cushman 1927 Planorbulinella larvata (Parker & Jones) Plate 26, figures 5a-c Ptanorbulina vulgaris larvata Parker & Jones (1860) Ann. Mag. Nat. Hist. III. 5: 294. Recent. Planorbulina larvata Parker & Jones (1865) Phil. Trans. 155: 380. pi. 19, figs. 3a, b. Recent, Indian Ocean. — Brady (1884) Rep. Voy. Cliallenger Zool. 9: 658. pi. 92, flgs. 5, 6. Recent, south Paciflc — "exclusively tropical." — Hadley (1934) Bull. Am. Paleont. 20 (70A) : 30. pi. 6, fig. 9. Upper Oligocene, Cuba. Test subdiscoidal, more flattened on attached dorsal side; chambers in central part of test covered by laminae and fine, elongate knobs, only the chambers of the outermost whorl being iinobscured; edge rounded; apertures oval, two to each chamber, at the ends of the chamber in the median line. Diameter, 0.88 to 1.44 milUmeter; thickness, 0.15 millimeter Plesiotype No. 3947, locality A21, Quebradillas formation. Abundant at A86 of the Cibao formation. Rare at A20; abundant at A21 of the Quebradillas formation. Family ACERVULINIDAE Schultze 1854 RUPERTIA Wallich 1877 Rupertia verrucosa NEW SPECIES Plate 27, figures 9a-c Test large, attached in life by the concave dorsal side, irregularly tuberose, not columnar; early chambers close coiled like Cibicides for one whorl or more, followed by irregularly arranged chambers forming a tuberose mass of round or elongate, loosely or closely appressed chambers; wall calcareous, thick, coarsely perforate; sur- face in the young smooth, showing the closely spaced, large pores; in the adult the surface is usually covered with a rough, nodular deposit of secondary calcareous, shell material; aperture a sUt or large irregular arch at the base of the last septal face, with upper lip, never on the upper side of the test, nor tubular. Diameter, up to 4 milUmeters; height, up to 3 millimeters. Diameter of figured speci- men, 3.1 millimeters; height, 3 millimeters. 404 SCIENTIFIC SURVEY OF PORTO RICO Holotypc No. 4130, locality P4, Ponce formation. Abundant at P4 of the Ponce formation. This species differs from R. stabilis WaUich in the tuberose rather than columnar and bulbous shape, in the rough coating of secondary shell material in the adult test, in the lateral rather than terminal position of the aperture, and in the larger size. CARPENTERIA Gray 1858 Carpenteria bulloides NEW SPECIES Plate 27, figures 1 to 4 Test multilocular, irregularly cylindrical or columnar in the adult, globigerinoid in the young; chambers subglobular, increasing gradually in size from the apical, attached end, and arranged in an alternating series, 6 chambers in the adult test; sutures distinct, depressed; test attached in life, the place of attachment being on the first 2 or 3 chambers as shown by the cicatrix; aperture with a short neck of clear, imperforate shell material located on the side of the test opposite the point of attachment, in the adult terminal, not opening into the sutures or into an umbilical depression; aperture in the young oval in shape with a short flaring neck, in the adult round with a short conical neck; surface with large pores which are emphasized by deposits of secondary shell material, pores missing and the surface of the test smooth around the neck. Length of adult specimen, 1.61 millimeters; diameter, 1.21 millimeters. Holotypes No. 3881a, b, c, d, locality A21, Quebradillas formation. Common at A21 of the Quebradillas formation. C. bulloides lacks the broad basal attachment of C. lithothamnica Uhlig from the middle Eocene of West Galicia (1886. Jahrb. Geol. Reichsanstalt 36: 189. pi. 5, figs. 1,2). It resembles even more closely C. alternata Chapman and Crespin from the middle Tertiary of Vic- toria (1930. Proc. Roy. Soc. Victoria 43 (1): 99. pi. 5, figs. 9, 10) but differs in the coarsely perforate character of its surface and in the position and character of its aperture. We are not satisfied with the generic disposition of this species, but do not wish to redefine the genus at this time. Carpenteria proteiformis Goes Plate 27, figure 5 Carpenteria balaniformis proteiformis Goes (1882) Kon^l. Svensk. Vet. Akad. Handl. 19: 94. pi. 6, ngs. 208-214, pi. 7, flk's. 215-219. Recent, Caribbean Sea. Carpenteria proteiformis Cushnian (1931) liull. U. S. Nat. Mus. 104 (8): 140. pi. 26, fig. 1. Recent, West Indies. GALLOWAY AND HEM I NW AY, TERTIARY FORAMINIFERA 405 Test attached by a flat base, tending to be columnar, the basal portion more or less spreading; chambers inflated, the later chambers making an irregular subcylindrical column of several chambers which may be low or higher with staggered uniserial chambers; wall coarsely perforate, with a cellular, areolate surface suggestive of the mushroom Morchella esculenta; aperture circular, often with a tubular neck which is imperforate; position of aperture variable and several cham- bers may show apertures at once. Length of incomplete specimen, 2.0 milHmeters; diameter, 1.0 milHmeter. Plesiotype No. 4131, locality A21, Quebradillas formation. Common at A21 ; rare at A46 of the Quebradillas formation. Com- mon at P431 and P432 of the Ponce formation. Our specimens are not as elongate as specimens usually considered typical of this species, and the wall is more coarsely cellular or reticu- late. Perhaps they belong with C. utricularis. The species of Car- penteria vary so widely that it is difficult to identify any of them. GYPSINA Carter 1877 Gypsina discus (Goes) Plate 28, figure 4 Tinoporus vesicularis Goes (not P. and J.) (1882) Kongl. Svenska Vet. Akad. Hand]. My Foljd 19: 104. pi. 7, figs. 245-247. Recent, Caribbean Sea. Gypsina vesicularis discus Goes (1896) Bull. Mus. Comp. Zool. 21 : 74. pi. 7. figs. 4-6. Recent, west coast Central America. Test discoidal, biconvex; nearly circular in side view; edge sharply rounded; surface conspicuously reticulate, due to the chambers, the outer walls of which are usually broken away; the test consists of chambers added eccentrically around a small, coiled nucleoconch; the chambers are higher in the median region than on the flatter sides of the test; there is no median layer of chambers, as in the orbitoids, and the chambers are not in rays, as in Sphaerogypsina; the chambers are oval or subrhomboidal, as seen from the outside of the test and in parallel sections, averaging 0.13 millimeter in longer diameter; height of chambers, as seen in cross section, averaging 0.06 milUmeter; wall finely perforate; apertures, other than the perforations, absent. Di- ameter of test, up to 3 milUmeters; thickness, up to 1 millimeter. Plesiotype No. 4129, locaUty A21, Quebradillas formation. Common at A21; rare at F359, Quebradillas formation. Rare at P4, Ponce formation. The Tertiary Porto Rican specimens seem to be identical with the specimens figured by Goes, in the reference above. 406 SCIENTIFIC SURVEY OF PORTO RICO SPHAEROGYPSINA Galloway 1933 Sphaerogypsina globulus (Reuss) Plate 27, figure 7 Ceriopora globulus Reuss (1847) Haidinger's Naturwiss. Abh. 2: 33. pi. 5. fig. 7. Middle Miocene, Austria. Gypsina globulus Brady (1884) Rep. Voy. Challenger Zool. 9: 717. pi. 101, flg. 8. Recent, British Islands.— Cushman (1919) Carnegie Inst. Wash. Publ. 291 : 44. pi. 4, flg. 7. Upper Oligocene or early Miocene, Santo Domingo; (1935) U. S. Geol. Surv. Prof. Pap. 181: 54. pi. 23, figs. 4, 5. Upper Eocene, southeastern U. S. Sphaerogypsina globulus Galloway (1933) Manual of Foraminifera 309. pi. 28, figs. 13, 14. Test small, spheroidal; surface pustulose, or pitted, with a more or less regular polygonal network formed by the walls of the outside chambers; wall finely perforate, with no distinct aperture other than the perforations. Diameter, 0.27 to 1.11 millimeters. In cross section the chambers are seen to be rotaloid in the early portion with a microspheric nucleoconch (no megaspheric observed although several sections were made); the later chambers are con- centrically arranged in radial lines, enlarging mainly in width, very little in height toward the periphery; the later chambers are nearly equidimensional, circular or subcircular in shape near the center of the test, becoming more rectangular or oval near the periphery, the elongation being parallel to the periphery with maximum length up to 13/^ times the height; average diameter of chambers, 0.044 milli- meter; average height, 0.033 millimeter; layers of chambers relatively few, 8 to 14, outside of the nucleoconch. Plesiotype No. 3889, locality F358, Qucbradillas formation. Rare at LlC, L3C, and L4C of the San Sebastian formation. Common at A86 of the Cibao formation. Rare at P'358 of the Quc- bradillas formation. Rare at P2; common at P432 and P433 of the Ponce formation. Our specimens are identical in all respects with topotypes from the Miocene Leithakalk near Vienna, Austria. Sphaerogypsina pilaris (Brady) Plate 27, figure 8 Tinoporus pilaris Brady (1876) Ann. Soc. Mai. Belg. 11: 103. Miocene, Jamaica. Gypsina globulus Hill (1899) Bull. Mus. Comp. Zool. 34: 147. Miocene, Jamaica. Gypsina globulus (Reuss) var. pilaris Cuslunan (1919) Carnegie Inst. Wash. Publ. 291: 44. pi. 9, figs. 1, 2. Miocene, Cuba. Test spherical, large, the surface appearing almost smooth when observed megascopically, under the microscope ^vith a coarsely areolate or pustulose surface showing the outlines of the outer chambers; wall GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 407 finely but conspicuously perforate, with no aperture other than the perforations. Diameter, up to 3.2 or 4 millimeters. In cross section the microspheric nucleoconch is rotaloid, with later chambers added concentrically and arranged in radial lines; the radial tiers of chambers are conspicuous in natural cross sections of broken specimens as well as in laboratory sections; the later chambers are very numerous, in many concentric layers; chambers oval in shape, elongate parallel to the periphery with the length about twice the height, averaging 0.14 millimeter in diameter, height 0.067 millimeter. Plesiotype No. 3990, locality P258, Ponce formation. Common at A21, F359, and A64 of the Quebradillas formation. Abundant at P258 of the Ponce formation. S. ^pilaris occurs also in abundance in the calcareous strata 8.5 kilometers west of Isabel II on the north shore of the Island of Vieques, just east of Porto Rico. Meyerhoff correlates these beds with the upper part of the Ponce formation. S. 'pilaris differs from S. globulus in being generally larger in size, with much coarser areolation on the surface; in cross section the chambers are much more numerous, more closely appressed radially, and elongate rather than nearly equidimensional in shape. Family ASTERIGERINIDAE d'Orbigny 1839 AMPHISTEGINA d'Orbigny 1826 Amphistegina angulata (Cushman) Plate 28, figures 6a-c Asterigerina angulata Cushman (1919) Carnegie Inst. Wash. Publ. 291: 45. pi. 13, figs. la-c. Lower Miocene or upper Oligocene. Santo Domingo. Test rotaliform, nearly circular in side view; dorsal side moderately convex, convexity variable; ventral side strongly convex to conical; acutely angled at the edge but not carinate; about 14 chambers in the last whorl, of nearly uniform size; sutures flush with the surface, marked by clear shell material; sutures on dorsal side obliquely re- curved, on ventral side "with the sutures ending in an angle about midway between the center and the periphery, from which a secondary chamber is developed to the umbilical region, alternating with the main chambers; umbilical region solid, of clearer shell material" than the chamber walls; surface smooth except for fine papillae on the ventral side around the aperture and extending along the periphery; aperture rather long and narrow at the base of the last chamber on 408 SCIENTIFIC SURVEY OF PORTO RICO the ventral side of the test. Diameter, 0.67 (j^oiing specimens) to 1.38 millimeters; thickness, up to 0.83 millimeter. Plesiotype No. 3862, locality F359, Quebradillas formation. Common at F358; abundant at F359 of the Quebradillas formation. Common at P259 and 433; rare at P434; abundant at P258 of the Ponce formation. Only one whorl shows on the dorsal side of the test, not all the whorls of the spire as in Asterigerina. Amphistegina floridana Cushman & Ponton Plate 28, figures 5a-c Amphistegina floridana Cushman & Ponton (1932) Fla. Geol. Surv. Bull. 9: 96. pi. 14. figs. 6, 7. Lower Miocene, Florida. Amphistegina chipolensis Cushman & Ponton (1932) Fla. Geol. Surv. Bull. 9: 96. pi. 15, figs. la-c. Lower Miocene, Florida. Asterigerina choctawensis Cushman & McGlamery (1938) U. S. Geol. Surv. Prof. Pap. 189D: 111. pi. 28, flgs. 2a-c. Middle Oligocene, Alabama. Test planoconvex to biconvex, umbonal region more prominent on the ventral side; edge subacute with sUght keel; chambers numerous, 10-15 in the last whorl, increasing slowly in size, frequently with transparent alar prolongations over the umbonal region on the dorsal side of the test through which the earlier whorls are more or less clearly visible; sutures flush with the surface, marked by clear shell material, on dorsal side recurved about half way to the periphery, on ventral side nearly radial to junction with secondary sutures, then curving to the periphery which is met at an angle of about 45 degrees; secondary sutures appearing about midway between umbo and periph- ery; wall finely perforate; surface smooth except near the base of the aperture and along the periphery for a distance of about two chambers where the surface is strongly papillate; aperture ventral, at the base of the last septal face. Average diameter, 1 millimeter; thickness, 0.50 millimeter. Plesiotype No. 3863, locality L4C, San Sebastian formation. Rare at L2C; common at LlC, L3C, L5C; abundant at L4C of the San Sebastian formation. Common at A86 of the Cibao formation. Rare at F359; common at A 15, A21, and A93 of the Quebradillas formation. Rare at P2, P3, P4, and 434; common at P431; abundant at P258, P259, and P432 of the Ponce formation. There seems to be no appreciable difference between this species and A. chipolensis which was made at the same time as A. floridana and which comes from the same formation although from a slightly lower horizon. GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 409 Family CHILOSTOMELLIDAE Brady 1881 CHILOSTOMELLA Reiiss 1850 Chilostomella czizeki Reiiss Plate 28, figures 3a-c Chilostomella czizeki Reuss (1850) Denkschr. Akad. Wiss. Wien. 1 : 380. pi. 48, fig. 13. Middle Miocene Austria.— Schwager (1877) Boll. R. Com. Geol. Ital. 8: 26. pi., fig. 70. Tertiary. Italy. — Cushxaan (1926) Contr. Cusliman Lab. Foram. Res. 1: 74. pi. 11. fig. 2. Test small, smoothly elliptical in side view, the sides gently convex, the ends broadly rounded, not quite twice as long as thick, nearly circular in cross section but more sharply curved on the back; the last chamber covering two-thirds to three-fourths of the preceding chamber, the hne of contact being deeply concave on the back of the test, sometimes exposing a httle of several earlier chambers, including the allomorphinoid chambers and proloculum; wall very finely per- forate; surface smooth and sometimes mottled with yellow and white mosaic; aperture a narrow crescent comprising about one-fourth of the circumference of the test, with thick but not strongly everted lip. Length, 0.64 millimeter; diameter, 0.39 millimeter. Some specimens are larger. Plesiotype No. 4133, locality P254, Ponce formation. Rare at P254 and P255 of the Ponce formation. This species resembles C. cijlindroides Reuss but it is shorter and has more convex sides. Chilostomella globata NEW species Plate 28, figures 2a-c Test small, subglobular, the last chamber considerably larger than the previous one and constituting about two-thirds of the external test; subcircular in cross section; the suture line is only shghtly con- cave on the back of the test; wall very finely perforate; surface smooth; aperture a narrow crescentic slit, comprising about one-fourth of the circumference of the test, with large, strongly everted, shghtly undu- lating and fimbriate lip. Length, 0.38 millimeter; diameter, 0.33 millimeter. Other specimens up to 0.8 millimeter in diameter. Holotype No. 4134, locahty P254, Ponce formation. Rare at P254 and P255 of the Ponce formation. Chilostomella ovoidea Reuss Plate 28, figures la-c Chilostomella ovoidea Reuss (1860) Denkschr. Akad. Wiss. Wien 1: 380. pi. 48, figs. 12a-e. Middle Miocene. Austria.— Cushman (1924) Bull. U. S. Nat. Mus. 104 (5): 2. pi. 1. figs. 2. 3. Recent. Bermuda. 410 SCIENTIFIC SURVEY OF PORTO RICO Test ovoid, three-fourths as broad as long, nearly circular in cross section, tapering and more pointed at the apical than at the apertural end; greatest thickness above the middle of the test; the last chamber forms from two-thirds to three-fourths of the outside of the test; the suture line between the last two chambers nearly straight and trans- verse except for a slight apicad bend on the back of the test; wall finely perforate; surface smooth; aperture an arched opening com- prising about one-fourth of the circumference of the test, with thick, flaring lip. Length, 0.69 millimeter; diameter, 0.52 millimeter. Plesiotype No. 4135, locality P255, Ponce formation. Rare at P255 of the Ponce formation. Chilostomella urceolus NEW species Plate 27, figures 6a, b Test ovoid, circular in transverse section, broadly rounded at both apical and apertural ends, % as broad as long; greatest width more than twice that at the aperture; last chamber covering practically all of the preceding chamber, the line of contact between the two chambers being straight and at right angles to the axis of the test; wall smooth; aperture curved, at the base of the last chamber with a flaring lip. Length, 1 millimeter; diameter, 0.60 millimeter. Holotype No. 3883, locality P2, Ponce formation. Hare at P254, P255, and P4; common at P2 and P3 of the Ponce formation. This new species resembles C. ovoidea Reuss in many respects, but differs in that C. urceolus is rounded both apically and aperturally, is more elongate, and the line of contact between the last 2 chambers is not deflected posteriorly as in C. ovoidea. It is smaller than C. grandis and the suture is without posterior deflection. SPHAEROIDINA d'Orbigny 1S26 Sphaeroidina bulloides d'Orbigny Plate 30, figures la, b Sphaeroidina bulloides d'Orbigny (1826) Ann. Scl. Nat. 7: 267. no. 1; Modfeles, no. 65. Re- cent, Adriatic; Pliocene, Italy. ^Cushman (1914) Bull. U. S. Nat. Mus. 71 (4): 18. pi. 10, flg. 7, pi. 12. flg. 1. Itecent, north Pacific; (1924) Bull. U. S. Nat. Mus. 104 (5): 36. pi. 7. figs. 1-6. Recent, Caribbean, Gulf of Mexico, and western Atlantic from Brazil to Cape Cod. Test subspherical, exterior formed of a few chambers, usually 4 or 5; chambers slightly inflated, closely appressed; sutures moderately depressed; aperture a small arched opening with a flat tooth at the GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 411 inner margin of the last chamber. Diameter, 0.46 by 0.55 millimeter; thickness, 0.40 millimeter. Plesiotype No. 3991, locality P254, Ponce formation. Rare at F358 of the Quebradillas formation. Rare at P251, P254, P255, P431; common at P2 and P3 of the Ponce formation. Family ORBULINIDAE Schultze 1854 GLOBIGERINA d'Orbigny 1826 Globigerina bulloides d'Orbigny Plate 29, figures la, b Globigerina bulloides d'Orbigny (1826) Ann. Sci. Nat. 7: 277. no. 1; Modeles no. 76. Recent. Adriatic Sea; (1829-1844) Foraminiferes, Guerin-Meneville's Cuvier, IconograpWe. Mollusques 3: 9. pi. 2, fig. 12. Recent. Adriatic— Brady (1884) Rep. Voy. ChaUenger Zool. 9: 593. pi. 77, figs. 3-7. Recent, world wide: Cretaceous to Recent. — Galloway & Wissler (1927) Jour. Paleont. 1 : 40. pi. 7, fig. 4. Pleistocene. California.— Cole & Gillespie (1930) Bull. Am. Paleont. 15 (57B) : 12. pi. 2. fig. 16. Middle Oligocene. Meson for- mation. Me.Kico. Test of average size for the genus, longer than wide, consisting of about 23/^ whorls arranged in a trochoid spire; chambers about 12 in number, inflated, somewhat appressed, subglobular, rapidly enlarging, four constituting the last whorl ; sutures deeply depressed ; wall thick, finely reticulate ; aperture large, semicircular, opening into the umbili- cus. Length, 0.30 millimeter; width, 0.24 milUmeter; thickness, 0.19 millimeter, Plesiotype No. 3920, locality LlC, San Sebastian formation. Rare at LlC of the San Sebastian formation. Rare at A86; common at A43a of the Cibao formation. Rare at A79 of the Los Puertos formation. Rare at A20 and A21 of the Quebradillas formation. Rare at P2, P254, and P255; common at P3 of the Ponce formation. Globigerina dutertrei d'Orbigny Plate 29, figures 2a, b Globigerina dutertrei d'Orbigny (1839) in De la Sagra, Hist. Phys. Pol. Nat. Cuba Foramini- feres 84. pi. 4. figs. 19-21. Recent, oflf Cuba, Martinique. Guadeloupe. — Brady (1884) Rep. Voy. Challenger Zool. 9: 601. pi. 81. figs. la-c. Recent, Antarctic, south Pacific, West Indies.— Cushman (1921) Proc. U. S. Nat. Mus. 59: 55. pi. 12, figs. 7a-c. Recent, north coast Jamaica. Test rotahform, low spired, small for the genus; 5 chambers in last whorl; periphery lobulate; chambers inflated, slightly appressed; size of chambers rapidly increasing in early stages, the 5 chambers of the last whorl being more nearly equal in size; sutures depressed; wall finely reticulate; aperture a low arched opening into the umbilicus. Diameter, 0.28 by 0.36 millimeter; thickness, 0.20 millimeter. 412 SCIENTIFIC SURVEY OF PORTO RICO Plesiotype No. 3921, locality LlC, San Sebastian formation. Very rare at LlC of the San Sebastian formation. Rare at A43a; common at A86 of the Cibao formation. Rare at A21 of the Quebra- dillas formation. Rare at P432; common at P2 and P3 of the Ponce formation. This species differs from G. dubia Egger (1857. Neues Jahrb. Min. Geol. Pal.) in the lower spire. Globigerina inilata d'Orbigny Plate 29, figures 3a-c Globigerina inflata d'Orbigny (1839) in Barker, Webb & Berthelot, Hist. Nat. lies Canaries 2 (2): Foraminiferes 134. pi. 2, figs. 7-G. Recent, Canaries. — Cushman (1924) Bull. U. S. Nat. Mus. 104 (5): 12. pi. 3, figs. 1-3. Recent, mostly ofif New England. Test rotaloid, dorsal side nearly flat, ventral side very deep due to the rapid increase in height of the chambers; periphery moderately lobulate; edge flatly rounded with sharply rounded shoulder; whorls 2 to 3, the last usually having 4 chambers; sutures moderately de- pressed, slightly curved; wall finely perforate; surface smooth, the older tests covered with small, secondary knobs on the ventral side and on the dorsal early whorls; aperture a high arch on the ventral side extending from umbilicus to periphery, with thin upper lip. Diameter, 0.53 millimeter; height, 0.38 millimeter. Plesiotype No. 4137, locality P251, Ponce formation. Rare at P251 of the Ponce formation. Globigerina ouachitaensis Howe & Wallace Plate 29, figures 4a-c Globigerina ouachitaensis Howe & Wallace (1932) La. Dept. Cons. Geol. Bull. 2: 74. pi. 10, figs. 7a, b. Upper Eocene, Louisiana. — Howe (1939) La. Dept. Cons. Geol. Bull. 14: 83. pi. 12, figs. 18, 19. Middle Eocene, Louisiana. Test small, subglobular; periphery lobate; about 12 chambers visible on the dorsal side, 4 in the last whorl, differing very little in size in dorsal view, but increasing rapidly in thickness dorso-ventrally; chambers inflated; sutures distinct, depressed; aperture a fairl}^ high opening on the ventral side into the umbilical area, with previous apertures visible in the umbilical depression. Diameter, 0.40 by 0.42 millimeter; thickness, 0.30 millimeter. Plesiotype No. 4048, locality A21, Quebradillas formation. Common at A86 of the Cibao formation. Rare at A21 of the Que- bradillas formation. Rare at P2, P3, and P251; common at P255 of the Ponce formation. GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 413 Globigerina pachyderma (Ehrenberg) Plate 29, figures 5a-c Aristerospira pachyderma Ehrenberg (1861) Monatsber. k. preuss. Ak. Wiss. Berlin 303. Recent, Davis Strait; (1873) Abh. k. Akad. Wiss. Berlin pi. 1. fig. 4. Globigerina pachyderma Brady (1884) Rep. Voy. ChaUenger Zool. 9: 600. pi. 114, figs. 19, 20. Recent, northern oceans. — Heron-Allen & Earland (1909) Jour. Roy. Micr. Soc. 438. pi. 18, figs. 4, 5. Tertiary, Sussex. — Galloway & Wissler (1927) Jour. Paleont. 1: 43. pi. 7, flgs, 13a-c. Pleistocene, California. Test small, depressed subglobular; periphery slightly lobiilate; edge broadly rounded; whorls 2 or 3, the last consisting of 4 subequal chambers, the earlier whorls usually obscured by a secondary, granular deposit; sutures shghtly depressed, nearly straight and radial; dorsal side slightly convex, occasionally flat or slightly concave; ventral side very convex, shghtly umbilicate; wall coarsely and conspicuously perforate, thick; surface smooth or more usually covered with a secondary deposit of fine granules which obscure the pores; aperture a short, low arch at the base of the last chamber opening into the shal- low umbilicus, with thin upper lip. Length, 0.33 millimeter; width, 0.28 milUmeter; thickness, 0.26 millimeter. Plesiotype No. 4138, locaUty P254, Ponce formation. Rare at P254 of the Ponce formation. It is surprising to find this boreal species in subtropical deposits. The Porto Rican form seems to be typical in every way. Globigerina pseudotriloba White Plate 29, figures 6a, b Globigerina pseudotriloba White (1938) Jour. Paleont. 2: 194. pi. 27, figs. 17a, b. Cretaceous to Eocene, Mexico. Test longer than wide, composed of 2 whorls arranged in a low trochoid spire; chambers inflated, very httle appressed, rapidly and regularly enlarging, 3 comprising the last whorl; sutures deep; aper- ture a semi-circular opening at the base of the last septal face, opening into the umbihcal region. Length, 0.29 millimeter; width, 0.21 milli- meter; thickness, about 0.15 millimeter. Plesiotype No. 3922, locality LlC, San Sebastion formation. Rare at LlC of the San Sebastian formation. Common at A43a of the Cibao formation. Common at P3 and P254; abundant at P2 of the Ponce formation. This species is very similar to G. triloba Reuss (1850. Denkschr. k. Akad. Wiss. Wien 1) and has frequently appeared in the hterature of that species. Unhke G. triloba, G. pseudotriloba has only one aperture and that is located on the ventral side. 414 SCIENTIFIC SURVEY OF PORTO RICO Globigerina trilocularis d'Orbigny Plate 29, figures 7a, b, Globigerina trilocularis d'Orbigny (182G) Ann. Sci. Nat. 7: 277. no. 2. Miocene, near Bor- deaux, France. — Forna&ini (1898) Rendi. Sess. R. Accad. Sci. Istit. Bologna 2: 12. pi. 0, fig. 2. (from d'Orbigny's unpublished plates of 1826.), — Galloway & Morrey (1929) Bull. Am. Paleont. 15 (55): 10. pi. 3, fig. 9. Lower Oligocene, Ecuador. Test subglobular, consisting of 6 to 9 chambers arranged in a rota- loid coil of 2 whorls, with 3 chambers in the last whorl; periphery broadly rounded, slightl}^ lobulatc; chambers inflated, closely ap- pressed, enlarging rapidly; sutures slightly depressed; wall thick, reticulate; aperture an arched opening at the base of the last chamber opening into the umbilicus. Diameter of average specimen, 0.36 by 0.50 millimeter. Plesiotype No. 3923, locality L5C, San Sebastian formation. Rare at L5C of the San Sebastian formation. Abundant at A86 of the Cibao formation. Rare at A15 and A20; common at A21 and A93 of the Quebradillas formation. Rare at P2, P3, P254, and P255 of the Ponce formation. The Porto Rican forms are very similar to G. Iriloculinoides Plummer ((1926). Univ. Texas Bull. 2644), but lack the pecuHar aper- tural flap of that species. ORBULINA d'Orbigny 1839 Orbulina universe d'Orbigny Plate 30, figure 3 Orbulina universa d'Orbigny in Barker- Webb & Berthelot (1839) Hist. Nat. ties Canaries 2 (2) : Foraminifgres 123. pi. 1, flg. 1. Recent, Canaries. — Cusliman (1924) Bull. U. S. Nat. Mus. 104 (5) : 28. pi. 5, flgs. 2-9. Recent, Caribbean Sea, and widely distributed elsewhere. Test globular, consisting of only one exterior chamber; wall finely reticulate; aperture, when present, a simple round opening. Diameter of test, 0.26 millimeter. Plesiotype No. 3942, locality L6C, San Sebastian formation. Rare at LlC and L6C of the San Sebastian formation. Common at A93 of the Qu(>bradillas formation. Rare at A15, A20, and F359 of the Quebradillas formation. GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 415 Family PEGIDIIDAE Heron-Allen & Earland 1928 SPHAEROIDINELLA Cushman 1927 Sphaeroidinella seminulina (Schwager) Plate 30, figures 4a, b Globigerina seminulina Schwager (1866) Novara-Exped. Geol, Theil 2: 256. pi. 7, fig. 112. Pliocene, Kar Nikobar. Sphaeroidinella dehiscens Cusliman (not Parker & Jones) (1924) in part, Bull. U. S. Nat. Mus. 104 (5): 38. pi. 7, figs. 7, 8. Recent, Gulf of Mexico, and Atlantic Ocean. Test small for the genus, subglobiilar to broadly ovate, young specimens showing early globigerinoid chambers, adult forms with 3 chambers making up the entire visible portion of the test; sutures in the young test flush or very slightly depressed, the final suture of the adult fissure-like on the ventral side of the test, with jagged edges; wall thick with conspicuous, large pores which in some specimens make the surface rough ; aperture a small round opening at the bottom of the fissure-like suture. Length, 0.59 millimeter; diameter, 0.43 by 0.46 millimeter. Plesiotype No. 3992, locality A21, Quebradillas formation. Abundant at A21 of the Quebradillas formation. This species is distinctive in its slight development of a sutural fissure, and the complete absence of a sutural fissure on the dorsal side of the test. No specimens of the form commonly referred to *S. dehiscens (1865. Phil. Trans. 155) were found, hence it is improb- able that these are young or immature specimens of that species. Family HETEROHELICIDAE Cushman 1927 PAVONINA d'Orbigny 1826 Pavonina advena Cushman Plate 30, figures 2a, b Pavonina advena Cushman (1923) U. S. Geol. Prof. Pap. 133: 24. pi. 1, fig. 10. Middle Oligocene, Mississippi; (1926) Proc. U. S. Nat. Mus. 67 (25) : 22. (Summary of the genus.) Test very small, probably immature, fiabellate, compressed; chambers broadening very rapidly, tending to become annular, most of them arranged in an indefinite biserial manner, the later ones having radial depressions on the inner side (as if it were crimping due to bending of a tube, a feature seen in many bent, tubular forms) ; sutures slightly depressed, not Umbate; wall finely and evenly perforate; sur- face smooth but minutely granular; apertures absent excepting for the small mural pores on the septal face. Length, 0.24 millimeter; breadth, 0.22 millimeter; thickness, 0.05 millimeter. 416 SCIENTIFIC SURVEY OF PORTO RICO Plesiotype No. 4136, locality A6, Lares formation. Rare at A6 of the Lares formation. We have only a single specimen of this rare genus and species. It has the fine pores of P. advena, differing from most other species of the genus, and lacks the limbate sutures and lateral carina of other species. Our specimen is in shape very similar to the early stages of P. fiahelli- formis, as figured by Heron-Allen and Earland (1915. Trans. Zool. Soc. London 20: pi. 48, fig. 4). This form bears some resemblance to the early stages of Cyloloculina, but the chambers are not arranged in a uniformly spiral manner, and moreover this form does not occur with the numerous specimens we have from the Ponce and San Sebas- tian formations. BOLIVINA d'Orbigny 1839 Bolivina byramensis Cushman Plate 30, figures 11a, b Bolivina caclata byramensis Cusliman (1923) U. S. Geol. Surv. Prof. Pap. 133: 19. pi. 1, fig. 9. Middle Oligocene, Mississippi. Bolivina byramensis Cushman (1937) Cushman Lab. Foram. Res. Spec. Publ. 9: 69. pi. 8, figs. 18-20. Lower and middle Oligocene, southeastern U. S.; lower Oligocene, Texas: Oligocene or Miocene, Venezuela and Trinidad. — Coryell & Rivero (1940) Jour. Paleont. 14: 341. pi. 44, flgs. 17a-c. Middle Miocene, Haiti. Test small, tapering, about 13^ times as long as wide, rhomboidal in apertural view, oval in .side view; edge acute; chambers about 10 pairs, a little inflated near the center of the test becoming more com- pressed near the edge; sutures obhque, curved, all except the last few obscured by the ornamentation; .surface, except for the last 1 or 2 chambers, ornamented by a coarsely reticulate pattern of costae; aperture an elongate slit on the periphery at the base of the last cham- ber extending up onto the last septal face, bordered by a strong lip. Length, 0.57 millimeter; width, 0.31 millimeter; thickness, 0.13 millimeter. Plesiotype No. 3872, locality A86, Cibao formation. Common at A86 of the Cibao formation. Bolivina elongata Ilaiitkcii Plate 30, figures 5a, b Bolivina eUmoala IJantluu H87G) A magj . kir, foklt. int. fivkonyve 4: 55. pJ. 7. fig. 14; (1S81) Mitt. Jahrb. k. ungarn. geol. Anstalt 4: 65. pi. 7, fig. 14. Lower Oligocene, Hungary. — Cushman (1937) Cushman Lab. Foram. Res. Spec. Publ. 9: 51. pi. 6, flgs. 31. 32. Test elongate, very slightly tapering, a little mor(> than three times as long as wide, somewhat compressed; sides flat; edges rounded; GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 417 chambers enlarging gradually, about 8 pairs in the megaspheric form of the test; sutures oblique, near the edge of the test curving toward the apex, flush with the surface of the test and limbate except between the last few chambers where the sutures are slightly depressed and less conspicuously hmbate; wall finely perforate; aperture an elliptical opening in the last septal face, parallel to the compression of the test and extending upward from the base of the last chamber. Length, 0.73 millimeter; width, 0.22 millimeter; thickness, 0.15 milhmeter. Plesiotype No. 3875, locahty P255, Ponce formation. Very rare at P255 of the Ponce formation. This species resembles Cushman's figure (1937. pi. 6, fig. 32) of the topotype of B. elongata Hantken, although it does not have the stria- tions which are mentioned in the description and which are not shown in the figure of either the type or the topotype. Bolivina heineae NEW SPECIES Plate 30, figures 6a, b, and 7 Test small, regularly oval in side view, thick in the middle and thinning in all directions, widest and thickest part of the test midway between the apical and apertural ends; thickly elUptical in apertural view; chambers indistinct, obscured by the ornamentation, 7 to 9 pairs, low and broad, slightly overlapping, enlarging rather rapidly; sutures oblique, slightly curved, obscured by the ornamentation except for the last one near the apertural end; edge angular; surface ornamented by an anastomosing pattern of longitudinal costae which begin at the apical end, branch and are connected at irregular intervals by oblique cross costae; last chamber unornamented; aperture elon- gate, narrow, with lip. Length, 0.38 millimeter; width, 0.26 milli- meter; thickness, 0.15 miUimeter. Holotypes No. 3874a, b, locality P2, Ponce formation. Common at P2 of the Ponce formation. We take pleasure in naming this species for Professor Aida A. Heine who was one of the first paleontologists to take an interest in the foraminifera of Porto Rico. This species resembles B. hijramensis Cushman, but is a smaller, thicker, more oval form, not compressed, with reticulation that tends to be more longitudinal. Although B. heineae resembles B. reticulata Hantken (1875. Mitt. Jahrb. d. k. ung. geol. Anstalt 4) in general shape and size and fineness of reticulation, it differs in being thicker and having more definitely longitudinal costae which almost com- pletely obscure the sutures. 418 SCIENTIFIC SURVEY OF PORTO RICO Acid treatment was used to remove the surface ornamentation of one specimen and show the number of chambers and typical bolivine structure of this species. Bolivina jacksonensis Cushinan it Appliu Plate 30, figures 8a, b Bolivina jacksonensis Cushman & Applin (1926) Bull. Am. Assoc. Petr. Cieol. 10: 107. pi. 7. flgs. 3, 4. Upper Eocene, gulf coastal plain, U. S. and Mexico. — Cushman (1937J Cush- man Lab. Foram. Res. Spec. l^bl. 9: 57. pi. 7, flgs. 17, 18. Test small, compressed; ovate in side view, gradually tapering, broadest part of test about ^ distance from the apical end; narrowly ovate in apertural view; edge subacute but not carinate; chambers distinct, compressed, 8 or 9 pairs; sutures flush with the surface except between the last few chambers where they are a little depressed, marked by clear shell material, strongly recurved in earlier part of test, nearly straight and oblique to the edge in last part of the test; aperture narrow, at base of the last septal face, extending up toward a terminal position, with slight lip. Length, 0.33 miUimeter; \\4dth, 0.15 miUimeter; thickness, 0.05 millimeter. riesiotype No. 3876, locality LlC, San Sebastian formation. Rare at LlC of the San Sebastian formation. Bolivina matanzana convexa new variety Plate 30, figures 9a, b, and 10a, b Test small, subrhomboidal in side view, diamond shaped in end view, bluntly pointed at the apical end and rounded at the apertural (!nd, broadest part of test about % of the distance from apical end, near base of the last pair of chambers; edge acutely angled, slightly keeled; about 9 pairs of chambers in the test; sutures strongly curved, slightly raised, markful by clear shell material; wall conspicuously l)ei-forat(% some of the peiforations extending through the limbatc sutures; aperture extending from the base of the last septal face to a nearly terminal position, in shape elliptical with a raised rim. Length, 0.39 to 0.43 millimeter; width, 0.23 to 0.28 milHmetcr; thickness, 0.09 to 0.15 millimeter. Holotype No. 4038, locality A43a, Cibao formation. Paratype No. 4039. locality LlC, San Sebastian formation. Raw at LlC of the San Sebastian formation. Common at A43a of the Cibao formation. This variety is almost identical with B. matanzana Palmer and l^onnudez from th(> Oligocene of Cuba (1930. Mem. Soc. Cubana Hist. GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 419 Nat. 10), l)ut diftVrs in its smaller size and proportionally fj;reater thickness. Bolivina mexicana aliformis Cushnian Plate 31, figures la, b Bolivina mexicana aliformis Cushman (1926) Contr. Cushman Lab. Foram. Res. 1: 82. pi. 12, figs. 3, 4. Lower Oligocene, Ajazan formation, Mexico; (1937) Cuslunan Lab. Foram. Res. Spec. Pubi. 9: 66. pi. 8, figs. 9, 10. Oligocene, Mexico and Cuba. Test compressed, much thicker along the median line, elongate, tapering, 1.5 to 2.3 times as long as wide, greatest width near the apertural end; edge acutely angled with a broad, transparent keel, irregular in width, extending around the edges and apical end of the test; chambers numerous, about 12 pairs in the test, those of the early part low and broad, gradually increasing in height so that the last formed chambers are more nearly equidimensional; sutures distinct, limbate, those in the latter part of the test expanded toward the central part of the test; median thickness of the test due to a knobUke infla- tion of the inner ends of the chambers; wall finely but conspicuously perforate; aperture elongate, narrow, with a shght lip. Length, 0.23 to 1.00 millimeter; width, 0.14 to 0.50 millimeter; thickness, up to 0.20 millimeter. Plesiotype No. 4040, locality P3, Ponce formation. Rare at P254 and P255; common at P2 and P3 of the Ponce forma- tion. B. alazanensis venezuelana Hedberg (1937. Jour. Paleont. 11) from the middle Tertiary Carapita formation of Venezuela has a marked resemblance to this species, but differs in the absence of a broad keel. Bolivina tectiformis Cushnian Plate 31, figures 2a, b Bolivina tectiformis Cushman (1926) Contr. Cusliman Lab. Foram. Res. 1 : 83. figs. 6a, b. Oligocene, Mexico; (1937) Cushman Lab. Foram. Res. Spec. Publ. 9: 67. pi. 8, figs. 12-14. Oligocene, Mexico. Test small, elongate, oval in side view, nearly 2}/^ times as long as wide, rhomboid in apertural view; edge sharply rounded; periphery smooth; chambers closely appressed, about 8 pairs in the test, width greater than the length, enlarging gradually; sutures distinct, slightly curved, limbate, raised, fusing in the median line to form a rounded longitudinal costa or ridge; wall conspicuously but not coarsely perforate; surface of the early portion ornamented by obscure longi- tudinal grooves and costae; aperture nearly terminal, elliptical, with a thickened lip, extending from the base of the last chamber, up onto 420 SCIENTIFIC SURVEY OF PORTO RICO the septal face. Length, 0.40 minimctcr; width, 0.17 milHrneter; thickness, probably 0.08 or 0.10 millimeter. Plesiotype No. 3878, locality P2, Ponce formation. Ver}^ rare at P2 of the Ponce formation. Bolivina ventricosa NEW species Plate 31, figures 5a-c Test small with slight tendency to twist near the apical end, thickest in the middle of the test about midway of the length, tapering not only to the edges but also toward the apical and apertural ends; test widen- ing more rapidly after the first 3 or 4 pairs of chambers, apical end narrowly rounded, apertural end broadly rounded; edge angled, keeled; chambers closely appressed, about 9 pairs in the test, increas- ing markedly in width after the first 4 pairs, increasing very gradually in chamber length (i. e. adorally), greatest thickness at inner end of each chamber, the chamber thickness increasing rapidly for the first third of the test, decreasing equally rapidly for the last third of the test with maximum thickness of chaml^ers about midway between apical and apertural ends; sutures conspicuously curved, flush with the surface, continuous with the narrow keel of clear shell material that borders the test; aperture narrow, parallel to the compression of the test extending up onto the last septal face from the base of the last chamber on the periphery. Length, 0.35 millimeter; width, 0.30 millimeter; thickness, 0.17 millimeter. Holotype No. 3879, locality P254, Ponce formation. Rare at P254 of the Ponce formation. This species is quite different from all other species of Bolivina, particularly in its unusual thickness and the rapid widening of its test in side view. The shape of B. ventricosa suggests Ehrenhergina, but the planispirally coiled early portion is lacking and the aperture is straight, narrowly elliptical and nearly terminal, rather than lateral, in position. LOXOSTOMUM l']hrenbcrg 1S54 Loxostomum hiwanneense Howe Plate 31, figures 3a, b Loxostotna hiirannccnsc Howe (1030) Jour. I'aleont. 4: 32!). pi. 27, fig. 7. Lower Oligocene, Mississippi.— Ellisor (1933) Bull. Am. Assoc. Petr. Geol. 17: 1322, 1323. pi. 3. flg. 7. Lower Oligocene, Texas. Test small, elongalc, ovate in side and apcituial views, tapering lapidly near the apical end with slight apical spine; edge rounded; GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 421 chambers numerous, about 8 pairs; sutures straight, forming an oblique angle with the side of the test, flush with the surface, somewhat ob- scured by the ornamentation; all except the last chamber of the test ornamented by numerous fine, irregularly sinuous, longitudinal costae which are continuous across the suture lines; aperture small and elUp- tical, nearly terminal in position. Length, 0.39 millimeter; width, 0.18 millimeter; thickness, 0.11 millimeter. Plesiotype No. 3929, locality A43a, Cibao formation. Very rare at A43a of the Cibao formation. Loxostomum normale NEW SPECIES Plate 31, figures 4a, b Test small, elongate, about 23^ times as long as wide, tapering at both ends, thick and slightly twisted on the axis, widest a Uttle above the middle; periphery slightly lobulate, especially in the early part of the test; edge broadly rounded; chambers slightly inflated, about 5 pairs, increasing rapidly in size, the last chamber uniserial; sutures slightly depressed, oblique and slightly curved; wall very finely perforate; surface smooth; aperture terminal, elongate to ovate, parallel to the sides of the test, with slight lip. Length, 0.43 milli- meter; width, 0.16 milUmeter; thickness, 0.09 millimeter. Holotype No. 3930, locality A43a, Cibao formation. Rare at A43a of the Cibao formation. This species is close to the Recent species L. rostrum Cushman (1933. Contr. Cushman Lab. Foram. Res. 9: 82. pi. 8, fig. 13) from the tropi- cal Pacific, but differs in that L. normalis is smaller, is more tapering at the apical end, and has oblique sutures that become less oblique as chambers are added, rather than more oblique. PLECTOFRONDICULARIA Liebus 1902 I'lectofrondicularia trinitatensis Cushman & Jarvis Plate 31, figures 6a, b Plectofrondicularia trinitatensis Cushman & Jarvis (1929) Contr. Cushman Lab. Foram. Res. 5: 11. pi. 2, flg. 16. Upper Eocene, Trmidad. Test lanceolate, sides nearly parallel except toward the apical end where the test is gently tapering, initial end subacute; about 4 pairs of chambers in the early biserial part of the test, most of the test con- sisting of uniserial chambers which are slightly equitant; uniserial chambers increasing gradually in height; sutures obscured by the ornamentation which consists of several low costae, continuous from 422 SCIENTIFIC SURVEY OF PORTO RICO the initial to the apertiiral end, one on each edge and 4 on either side of the test; apertural end broken off. Length of broken specimen, 0.85 millimeter; width, 0.21 millimeter; thickness, 0.09 millimeter. Plesiotype No. 4068, locality P3, Ponce formation. Very rare, only one specimen being found at P3 of the Ponce formation. Plectofrondicularia vaughani Cushman Plate 31, figures 7a, b Plectofrondicularia vaughani Cushman (1927) Contr. Cushman Lab. Foram. Res. 3: 112. pi. 23, fig. 3. Lower Oligocene, Mexico; (1929) Contr. Cuslmian Lab. Foram. Res. 5: 92. pJ. 13, figs. 21, 22. Middle Tertiary, Ecuador, Venezuela. — Cushman & Jarvis (1930) Jour. Paleont. 4: 361. pi. 33. fig. 4. Miocene, Jamaica. — Hadley (1934) Bull. Am. Paleont. 20 (70) : 15. pi. 2. figs. 5, 6. Oligocene, probably upper, Cuba. — Coryell & Rivero (1940) Jour. Paleont. 14: 341. pi. 42, fig. 28. Middle Miocene, Haiti. Test elongate, compressed, oval in front view, initial end narrow, bluntly pointed ; proloculiim subspherical followed by several biseriallj'' arranged chambers, the major portion of the test consisting of about 5, very slightly inflated, equitant, chambers; sutures marked \\ith clear shell material, slightly depressed; aperture terminal, fissurine, not radiate. Length, L13 millimeters; width, 0.35 millimeter; thickness, 0.07 millimeter. Plesiotype No. 3949, locality P2, Ponce formation. Very rare at P2, P3, and P254 of the Ponce formation. The original description of this species states that, "this species is peculiar in the retention of the alternating character of the chambers as shown by the apertures pointing first to one side and then the other in succeeding chambers and the incurved portion of the wall even after the alar projections are developed", but it is evident from sub- sequent figures published by the originator of P. vaughani that this alternating character of the chambers throughout most of the test is not an essential differentiating characteristic. P. vaughani is essen- tially an elongate, oval form of moderate width with relatively few chambers, usually 5 to 7 equitant chambers following a short biserial portion of a few chambers. P. advena (Cushman) (1923. Bull. U. S. Nat. Mus. 104 (4)) is very similar, but is narrower and tends to be more irregular, or phylogerontic. GALLOWAY AND HEMINW'AY, TERTIARY FORAMINIFERA 423 Family BULIMINIDAE Jones 1876 REUSSELLA Galloway 1933 Reussella glabrata (Cushman) Plate 31, figures 8a, b Verneuilina spinulosa glabrata Cuslmian (1922) U. S. Geol. Surv. Prof. Pap. 129E: 92. Mid- dle Oligocene, Byrani, Mississippi. Reussia spinulosa Cushman & Ponton (not Reuss) (1932) in part, Fla. Geol. Surv. Bull. 9: 84. pi. 12, fig. 15 (not figs. 14, 16.) Area zone of Choctawhatchee, middle Miocene. Florida. Test small, inverted pyramidal in shape, 3-sided, greatest diameter near the apertural end; triangular in cross section, the sides somewhat concave; initial end and edges of the test angled, without spines or carinae; chambers closely appressed, regularly triserial in arrangement; sutures narrow, flush with the surface; surface smooth; aperture a small transverse slit at the inner edge of the last chamber. Length, 0.33 millimeter; diameter, 0.20 millimeter; breadth of one face, 0.24 millimeter. Plesiotype No. 4073, locality A79, Los Puertos formation. Common at L2C of the San Sebastian formation, and at A79 of the Los Puertos formation. Rare at A21 of the Quebradillas formation, and at P258 of the Ponce formation. BULIMINA d'Oibigny 1826 Bulimina socialis Bornemann Plate 31, ficiures 9a, b BuUmma socialis Bornemann (1855) Zeitsch. deutsch. geol. Ciesell. 7: .342. pi. 10, fig. 10. Middle Oligocene, German.v. — Cushman (1937) Contr. Cushman Lab. Foram. Res. 13: 36. pi. 4, figs. la-c. Middle OUgocene, Germany. Bulimina ovata Brady (1884) Rep. Voy. ChaUenger Zool. 9: 400. pi. 50. figs. 13a-b. Recent, north and south Atlantic and Pacific. — White (1929) Jour. Paleont. 3: 49. pi. 5, fig. 11. Mendez, upper Cretaceous, Mexico. — Nuttall (1932) Jour. Paleont. 6: 19. pi. 2, fig. 8. Lower Oligocene, Mexico. Bulimina sp. Cushman & Applin (1926) Bull. Am. Assoc. Petr. Geol. 10: 169. pi. 7, figs. 10-11. Upper Eocene, Texas. Test oval in front view, about 13^2 times as long as wide, widest about midway between apical and apertural ends, both ends of test rounded; test circular or nearly so in end view; 2 whorls of chambers visible; chambers slightly inflated, the 3 chambers of the last whorl constituting about ^ of the test; sutures distinct, somewhat depressed; wall finely perforate; aperture virguline, with a vertical, curved tooth. Length, 0.60 to 1.16 millimeters; diameter, 0.32 to 0.68 millimeter. Plesiotype No. 3880, locality P254, Ponce formation. 424 SCIENTIFIC SURVEY OF PORTO RICO Rare at P254 and P255; common at P3; abundant at P2 of the Ponce formation. Since 1846, when B. ovata was first named by d'Orbigny, workers on foraminifera have consistently used this name for all smooth speci- mens of Bulimina with generally oval shape in side view, circular cross section, slightly depressed sutures and relatively few chambers. Several authors, among them Cushman in 1922, Nuttall in 1932, and Palmer and Bermudez in 1936, have noted that two types of smooth Buliminas have been included in B. ovata, namely that of the original figure, and another which differs, "from the original figure of the species in being broader in proportion to the length, with the final whorl of chambers covering about three-fourths of the length of the test". (Palmer & Bermudez p. 287.) A comparison of the Porto Rican speci- mens with the original figure of B. socialis Bornemann and the figures of B. socialis pubhshed by Cushman in 1937 shows that the non-typical forms of B. ovata which have been erroneously included under B. ovata should be called B. socialis, a name which was proposed by Bornemann in 1855. It is improbable that the thicker type of Bulimina which we here call B. socialis, is the megaspheric form of B. ovata since only the B. socialis type occurs in the Porto Rican material, and not both types as would be expected if the two are the megaspheric and microspheric forms of the same species. B. socialis differs from B. ovata in the following respects: 1. the apical and apertural ends are narrowly to broadly rounded in shape, not sharply rounded or angular 2. the test is stouter; the form ratio (length divided by breadth) being 1.4 to 1.90, while B. ovnta ranges from 2 to 2.4. The type figuie of B. socialis was 1.57, of B. ovala 2.20. Family CASSIDULINIDAE d'Orbigny 1839 CASSIDULINA d'Orbigny 1S26 Cassidulina laevigata d'Orbigny Plate 32, figures la, b Cassidulina laevigata d'Orhifoiy (182C) Ann. Sci. Nat. 7: 282. pi. 1.5. figs. 4, 5; Model no. 14. Recent.— Brady (1884) Rep. Voy. Challenger Zool. 9: 428. pi. 54. figs. 1-3. Recent, widespread. — Cushman (1918) U. S. Geol. Surv. Bull. 676: 9. pi. 1, figs. 5a, b. Pliocene, North Carolina. Test lenticular, nearly equally biconvex, biumbonate; nearly circular in side view with slightly lobate outline; edge angled but not carinate; chambers short and wide with nearly parallel edges, about 5 pairs in the last whorl; sutures slightly curved, limbate, flush with the GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 425 surface or very slightly depressed near the edge of the test; wall smooth, finely perforate; aperture an elongate virguline slit on the face of the last chamber nearly parallel to the periphery. Diameter, 0.24 by 0.26 millimeter; thickness, 0.13 millimeter. Plesiotype No. 4142, locality A43a, Cibao formation. Rare at A43a of the Cibao formation, and at A79 of the Los Puertos formation. Cassidulina subglobosa Brady Plate 32, figures 2a, b Cassidulina subglobosa Brady (1881) Quart. Jour. Micr. Sec. 21: 60; (1884) Rep. Voy. Chal- lenger Zool. 9: 430. pi. 54, figs. 17a-c. Recent, Atlantic and Pacific Oceans. — Galloway & Morrey (1929) Bull. Am. Paleont. 15 (55) : 40. pi. 6, fig. 6. Lower Oligocene, Ecuador.— Hedberg (1937) Jour. Paleont. 11: 680. pi. 92, flg. 5. Upper Oligocene, Venezuela. Test subglobular, only slightly compressed on the sides; broadly oval in side and edge views; edge broadly rounded; chambers few, 3 pairs in the last whorl, inflated; sutures somewhat depressed; surface smooth, wall finely perforate; aperture a large virguline open- ing extending up onto the last septal face, obliquely to the plane of coiling. Diameter, 0.40 millimeter; thickness, 0.32 milhmeter. Plesiotype No. 4180, locality P2, Ponce formation. Rare at P2, P3, and P254 of the Ponce formation. Cassidulina tricamerata NEW SPECIES Plate 32, figures 3a-c Test subequally biconvex, biumbonate, not quite planispirally coiled, subcircular to broadly oval in side view; periphery very slightly lobulate; edge sharply rounded, not keeled; chambers large, closely ' appressed, very little inflated, 3 pairs in the last whorl, enlarging rapidly, the last chamber constituting from one-third to two-fifths of the circumference of the test; sutures narrowly limbate, a little de- pressed, slightly curved, nearly parallel on the two sides of each chamber; wall very finely perforate; surface smooth; aperture a narrow, curved slit in the plane of coiling, extending about one-sixth of the circumference of the test or about one-half the circumferential dimen- sion of the septal face, without upper lip, but with lower, undulating, plate-like tooth; septal face narrow, convex, with outer, inflated rim. Diameter, 0.53 by 0.60 millimeter; thickness, 0.30 millimeter. Holotype No. 4143, locality P255, Ponce formation. Rare at P2 and P3; common at P254 and P255 of the Ponce forma- tion. This species differs from C. delicata Cushman (1927. Scripps Inst. 426 SCIENTIFIC SURVEY OF PORTO RICO Oceanog. tech. scr. 1 : 168. pi. 6, fig. 5) in the fewer chambers and much shorter aperture. It differs from C. rariloculn Cushman (1933. Contr. Cushman Lab. Foram. Res. 9: 93. pi. 10, fig. 4), from the tropical Pacific, in the less inflated chambers, sharper edge, longer aperture, and larger size. Perhaps C. ''delicatula" Cushman (obviously an error for "delicata") (1929. Contr. Cushman Lab. Foram. Res. 5: 101. pi. 14, fig. 12, Tertiary, Venezuela) is the same as our species, but the chambers are rather too inflated and the sutures too much curved. EHRENBERGINA Reuss 1850 Ehrenbergina caribbea new species Plate 32, figures 4a-d EhTenbcrgina bradyi Coryell & Rivero (not Cushman) (1040) Jour. Paleont. 14: 342. pi. 44 fig. 22. Middle Miocene, Haiti. Test short subtriangular in side view, oval in edge view, pyramidal in apertural view; dorsal side convex, ventral side more strongly convex, with a broafl, flat ridge down the center, bounded by concave slopes; edge sharp, with a short, flat spine at the end and upper side of each chamber and a few, small spines on the apical, ventral end; chambers closely appressed, not inflated, about 7 pairs; dorsal sutures curved, overlapping about half their length, limbate, mostly flush with the surface, the middle part of the later ones raised; ventral sutures obscure, nearly flush with the surface, curved and overlapping on the ventral ridge; wall very finely perforate; surface smooth except foi- the raised sutures and spines on the edge and ai:)ical end; aperture a thin, curved slit just under the terminal part of the ])eriphery, nearer the sutural side of the chamber, with thin lower lip. Length, 0.61 millimeter; width, 0.56 millimeter; thickness, 0.30 millimeter. Holotypc No. 4140, locality P2, Ponce formation. Rare at P2 of the Ponce formation. This species is similar to E. hradyi Cushman (1927. Proc. U. S. Nat. Mus. 70 (16) : 5. pi. 2, fig. 1.) from deep water of the Pacific, but it lacks the median ventral groove and spines, the fine peripheral spines, and it is shorter. The subtropical middle Tertiary forms from Haiti and Porto Rico seem to be identical, and are readily distinguishable from the cold water Recent forms from the Pacific. E. navali^ Hadley (1934. Bull. Am. Paleont. 20 (70A)) seems very close to ?J. caribbea, but it is moic slender, has more delicate spines, and lacks the small spines on the apical, ventral end of the test. GALLOWAY AND IIEMINWAY, TERTIARY FORAMINIFERA 427 Ehrenbergina serrata gibbera new variety Plate 32, figures 5a-d Test subtriaiigiilar in dorsal viow, oval in odge view, tho early part as thick as the later part; biconvex, oval in cross section with the back humped (hence the name gibbera), the later part uncoiled and perhaps also the earlier part, the exact nature of which is not determinable because of the addition of spines and other secondary material; edge sharply rounded in the earlier part, angled and slightly carinate in the later part; earlier part with a round spine extending out from the edge of each chamber and other spines on the apical end and on the early, ventral side; chambers 6 to 7 pairs, on the dorsal side short and wide, overlapping each other by about half their width; sutures on the back of the test limbate but flush with the surface, curved in three parts, usually orad, sometimes apicad; ventral sutures depressed, obscured by secondary material; surface smooth except for the spines; aperture elongate, narrow and curved, near the middle of the test, nearly parallel to the margin, with fimbriate lower lip and welt-like upper lip. Length, 0.60 millimeter; width, 0.55 millimeter; thickness, 0.30 millimeter. Holotype No. 3902, locality P2, Ponce formation. Rare at P2 of the Ponce formation. This species differs from E. serrata Reuss (1850. Denkschr. k. Akad. Wiss. Wien 1: 377. pi. 48, fig. 7), from the Miocene of Austria, in the less definitely coiled and more spinose early stage, the humped back, and fewer chambers. Family UVIGERINIDAE Galloway & Wissler 1927 UVIGERINA d'Orbigny 1826 Uvigerina bulbacea new species Plate 33, figures 1, 2 Test small, elongate bulbous, more than twice as long as wide, enlarging rapidly from the rounded apical end to the greatest width, about one-third of the length from the apical end, thence gradually tapering to the aperture; chambers in about 3 whorls, closely appressed, little inflated, regularly triserial in the first third, becoming more inflated and fewer to a whorl up to the terminal chamber which is uniserial; sutures flush and obscure in the first third, depressed and plain in the later two-thirds; wall finely perforate; apical end of the test provided wiih very small spines, not in lines, the later part of the 428 SCIENTIFIC SURVEY OF PORTO RICO last chamber and apertural neck finely granulate; most of the test with fine, low, round costae, not continuous across the sutures, about 20 on one side of the test or 16 to 20 in a width of 0.2 millimeter; last chamber without shoulders, the last chamber tapering into a short neck, without phialine lip. Length, 0.51 millimeter; diameter, 0.21 millimeter. Variation in length from 0.32 to 0.54 millimeter; diameter from 0.18 to 0.24 millimeter; form ratio from 1.8 to 2.7 millimeters, averaging 2.2 millimeters. Holotype No. 4144, paratype No. 4144a, locality A86, Cibao forma- tion. Common at A86 of the Cibao formation. This form is like U. postica, with which it occurs, but differs consist- ently in the following ways: it is smaller, the greatest width is nearer the apical end, the tapering to the aperture is more marked, the spines are smaller on the apical end and absent on the apertural end, and the costae are smaller. We have considered the possibility that one form is megaspheric and the other microspheric, but find no proof. Note the greater magnification of U. hulhacea. Uvigerina elongata Cole Plate 33, figure 5 Uvigerina elongata Cole (1927) Bull. Am. Paleont. 14 (51): 26. pi. 4, flgs. 3. 4. Middle Eo- cene, Mexico. — Cusliman (1937) Contr. Cushman Lab. Foram. Res. 13: 78. pi. 11, flgs. 15, 16. Middle Eocene. Mexico. Test small, elongate, greatest width below the middle of the test; early chambers closely appressed, enlarging rather rapidly in size, tending to become uniserial in the later part of the test; sutures obscure in the early part of the test, depre.s.sed in the later part; surface hirsute, i. e. with small fine spines irregularly arranged; apertural end sloping with a neck; aperture terminal, round. T^ength, 0.52 milli- meter; diameter, 0.18 millimeter. Plesiotype No. 4017, locality P254, Ponce formation. Rare at P254 and P255 of the Ponce formation. Uvigerina fusiformis NEW SPECIES Plate 33, figure 6 Test small, fusiform, tajieriiig toward both ends, apertural end tapering to the slender, cylindrical neck; chambers about six on a side, rather loosely appressed; sutures distinct, d(5pressed; surface of the last few chambers ornamented by fine, irregularly arranged spines; earlier part of the test with fine spines in vertical lines, apical end of GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 429 the test finely spinose; aperture round at the end of a slender, tapering neck, with lip. Length, 0.44 millimeter; diameter, 0.18 millimeter. Holotype No. 4018, locality P2, Ponce formation. Rare at P2, P3, P254, and P255 of the Ponce formation. This species differs from U. -pygmaea d'Orbigny (1826. Ann. Sci. Nat. 7) in the finer striae. It seems close to U. elongata, but is shorter, more regular in shape, and has the spines in lines on the early part of the test. The specific name fusiformis is used because of the fusiform shape of the test. Uvigerina gallowayi Cushman Plate 33, figure 8 Uvigerina alaia Galloway & Morrey (not Cushman & Applin) (1929) Bull. Am. Paleont. 15 (55): 38, pi. 6, fig. 1, Oligocene, Ecuador. Uvigerina gallowayi Cushman (1929) Contr. Cushman Lab. Foram. Res. 5: 94. pi. 13. flgs. 33, 34. Oligocene, Ecuador. Test of medium size, stout fusiform, with pointed apical end and shouldered apertural end; length twice the thickness, greatest thick- ness above the middle; chambers regularly triserial, closely appressed, little inflated; sutures slightly depressed, marked by clear shell material; surface ornamented with 14 to 16 strong costae which begin below the proloculum, cross the sutures and fade out on the last chamber or two; aperture round with small lip, at the end of a short neck, situated in an encircling depression. Length, 0.75 millimeter; diameter, 0.40 millimeter. Plesiotype No. 4145, locaUty A86, Cibao formation. Rare at A86 of the Cibao formation. Uvigerina gardnerae Cushman Plate 33, figures 13a, b Uvigerina gardnerae Cushman (1926) in Cushman & Applin, Bull. Am. Assoc. Petr. Geol. 10: 175. pi. 8, figs. 16, 17. Upper Eocene, Texas. — Cushman & Edwards (1937) Contr. Cushman Lab. Foram. Res. 13: 79. pi. 11, figs. 19, 20. Upper Eocene, Atlantic and gulf coastal plain. Test large, stout, widest at or above the middle; ends rounded; chambers regularly triserial, inflated; sutures depressed but indistinct; surface ornamented by about 20 plate-like costae, 5 or 6 in a width of 0.2 miUimeter, not continuous with those on adjacent chambers, partly breaking up into spines or disappearing on the last chamber; aperture round, at the end of a short neck, with small lip. Length, 0.83 millimeter; diameter, 0.45 millimeter; form ratio 1.8. Plesiotype No. 4019, locaUty P254, Ponce formation. 430 SCIENTIFIC SURVEY OF PORTO RICO Rare at P254 of the Ponce formation. Thi.s form is not, "much elongated, slender", as stated in the descriptions of U. gardnerae, and the measurements in the descriptions and in the plates do not agree. U. gardnerae belongs to the U. pere- grina group, but differs in being thicker and shorter, with fewer costae, and absence of a granular surface. It is much like U. gardnerae ciibaua Hadley (1934. Bull. Am. Paleont. 20 (70A)) which lacks spines on the last chamber. It is also much like U. capayana Hedberg (1937. Jour. Paleont. 11 : 677. pi. 91), but is much larger and has much coarser costae. The various groups of species of Uvigerina have been insufficiently differentiated for definite recognition. Uvigerina mantaensis Cusliiiian & Edwards Plate 33, figure 7 Uvigerina proboscidea Galloway & Morrey (not Schwager) (1929) Bull. Am. Paleont. 15 (55): 39. pi. 6, fig. 4. Lower Oligocene, Ecuador. Uvigerina mantaensis Cushman & Edwards (1938) Contr. Cushman Lab. Foram. Res. 14: 84. pi. 14, flg. 8. Oligocene, Ecuador. Test stout, about IJ^ times as long as wide, broadly fusiform in shape; chambers rather few, inflated; sutures distinct, depressed; wall hispid throughout; shoulders broad; aperture terminal, with a short, cylindrical neck and phialine lip. Length, 0.59 millimeter; diameter, 0.32 milHmeter. Plesiotype No. 4076, locality P3, Ponce formation. liare at P3 and P254 of the Ponce formation. U. ma7}taensis differs from U. proboscidea Schwager (1866. Novara- Exped. C.eol. Theil 2) in that it has a less tapering test and broad, not sloping, shoulders. Uvigerina mexicana Nnttall PlATK 33, FIGURE 9 Uvigerina mexicana Nuttall (19:12) Jour. Paleont. 6: 22, pi. r>, ligs. 12. 13. I^wer Oligocene. Mexico. Test short, stout fusiform in shai)e; apical end .>^ubacut(> or pointed; thickest part of the test near or below the middle; lower % of the test ornamented by numerous low, round, longitudinal costae, which are continuous from chamber to chamber, and increase by division, less frequciilly l)y implantation; last chamber unornamented; early chambers rather obscure, little inflated, last chamber slightly inflated; sutures flush with the surface, obscure in the early part of the test, last few marked by dear shell material; ai)erturc loimd, at the end of GALLOWAY AND IIEMINWAY, TERTIARY FORAMJNIFERA 431 a short cylindrical neck which is set in a depression or on a flattened area of the last septal face. Length, 0.70 millimeter; maximum diameter, 0.44 millimeter. Plesiotype No. 4021, locality P2, Ponce formation. Rare at P2, P3, and P255 of the Ponce formation. Uvigerina mexicana bulbosa new variety Plate 33, figure 10 This variety differs from typical U. mexicana in being subglobular in shape, and broadly rounded, not tapering at both apical and apertural ends. Length, 0.54 millimeter; chameter, 0.48 millimeter. Holotype No. 4077, locahty P3, Ponce formation. Rare at P3 of the Ponce formation. Uvigerina mexicana ranunculus NEW VARIETY Plate 33, figure 11 Test stout, circular in cross section, more tapering at both ends than typical U. mexicana; maximum diameter about midway the length of the test; apical end with a flat, spine-like process formed by the union of several costae; other characteristics like typical U. mexicana. Length, 0.92 miUimeter; diameter, 0.50 milhmeter. Holotype No. 4022, locality P2, Ponce formation. Rare at P2 of the Ponce formation. Uvigerina postica new species Plate 33, figures 3, 4 Test elongate, about twice as long as AAide, bulbous, the later two- thirds converging to the apical end, enlarging rapidly from the apical end to the greatest width below the middle; apical end pointed, some- times with short apical spine; chambers in about 4 whorls, closely appressed and regularly triserial in the first third of the test, slightly inflated and tending to become biserial in the second third, and con- siderably inflated and tending to become uniserial in the final third of the test; sutures flush with the surface in the first third of the test, marked by clear shell material, somewhat depressed in the later two- thirds; wall finely perforate; surface of the apical and apertural ends provided with small spines which are not in definite lines; most of the test ornamented by fine, low, round longitudinal costae, not continu- ous across the sutures, about 20 on one side of the test or 13 in a width of 0.2 millimeter; shoulders of last chamber round and sloping; aper- 432 SCIENTIFIC SURVEY OF PORTO RICO ture with a short, tapering neck, not set in a depression, without Up or with a very narrow phiaUne Hp. Length, up to 0.66 milUmeter; diameter, 0.30 millimeter. Form ratio (length over diameter) 1.9 to 2.28, averaging 2.1. Holotype No. 4146a, paratype No. 4146b, locality A86, Cibao formation. Common at A86 of the Cibao formation. This species is similar to U. striata d'Orbigny in the fine costae, but scarcely deserves comparison. The comparison with U. bulbacea is given under that species. The specific name postica refers to the enlarged posterior part of the test, above the pointed apical end. Uvigerina vicksburgensis Cushman & Ellisor Plate 33, figures 12a, b Uvigerina vicksburgensis Cushman & Ellisor (1931) Contr. Cushman Lab. Foram. Res. 7: 54. pi. 7, figs. 7a, b. Lower Oligocene, Texas. — Ellisor (1933) Bull. Am. Assoc. Petr. Geol. 17: pi. 3, fig. 10. — Cushman & Edwards (1938) Contr. Cushman Lab. Foram. Res. 14: 76. pi. 13, figs. 10, 11. Lower Oligocene, gulf coastal plain of U. S. and Mexico. Test regularly fusiform, over twice as long as wide, greatest width near the middle; chambers in about three whorls, closely appressed, regularly triserial to near the apertural end, where the chambers be- come biserial and tend to be uniserial; sutures slightly depressed, obscure; surface ornamented, from apex to apertural neck, with about 18 or 20 strong, sharp costae, about 5 in a width of 0.2 millimeter, which curve considerably; some costae are confined to one chamber, others cross the sutures; aperture terminal, round, at the end of a short neck, which is not set in a depression, with small lip. Length, 0.73 millimeter; width, 0.35 millimeter. Plesiotype No. 4020, locality P251, Ponce formation. Rare at P251 of the Ponce formation. This is one of the better characterized species of the genus, with its fusiform shape and part of the costae crossing the sutures. SIPHOGENERINA SchlumbLM-ger 1883 Siphogenerina costostriata N'EW SPECIES Pl,,\TK 34. FIGURES 5a, b Test of medium size, slender, the early part conical, the later part somewhat compressed; the early part sharply rouiuUnl, gradually expanding to the broadly rounded apertural end; chamb(>rs triserial and closely appress(-(l in the first third of th(> t(>st, becoming biserial in GALLOWAY AND HEMINWAY, TERTIARY FORAMINJFERA 433 the initklle third and iiniserial in the final third, progressively more inflated from the middle to the oral end; sutures obscure in the first third, d(!pressed in the later two-thirds; surface ornamented with fine striae in the first one-third of the test, about 20 in a width of 0.2 millimeter; the later two-thirds with thin, low costae, 5 to 6 in a width of 0.2 millimeter, between each pair of which there are 2 or 3 much smaller, raised striae; neither the costae nor striae cross the sutures; aperture oval, without phiahne lip, at the end of a short neck, which is not set in a depression. Length, 0.78 milUmeter; width, 0.26 miUi- meter; thickness, 0.21 miUimeter. Holotype No. 4147, locaUty P254, Ponce formation. Rare at P254 of the Ponce formation. This species has just passed from the Uvigerina to the Siphogenerina stage. It differs from described species of both genera in the two sets of costae, and in other ways. Siphogenerina cumingsi NEW species Plate 34, figures 6 and 7 Test elongate, subcylindrical ; apical end pointed in the more elon- gate, microspheric form, bluntly rounded in the stouter megaspheric form; triserial stage very short, uniserial stage with about 7 chambers which are about % as long as wide; sutures flush with the surface, curved adorally between the costae; wall ornamented by 7 to 8 longitudinal costae, each continuous the length of the test except for the last chamber; apical end spinose; triserial portion of megaspheric test with short costae intercalated between the primary costae; apertural end rounded with a short neck; aperture terminal, round, with slight phialine lip. Dimensions of average megaspheric form: length, 0.94 miUimeter; diameter, 0.26 miUimeter. Dimensions of figured microspheric form: length, 0.86 miUimeter; diameter, 0.26 mUlimeter. Maximum size of microspheric form: length, 1.26 milli- meters; diameter, 0.30 millimeter. Holotype No. 3983, locaUty P3, paratype No. 3984, locality P2, Ponce formation. Holotype is megaspheric form; paratype micro- spheric. Rare at P3 and P255; common at P2 of the Ponce formation. Both megaspheric and microspheric forms occurred at all three localities. S. cumingsi difl^ers from S. spinosa Bagg (1904. Md. Geol. Surv. Mio.; refigured by Cushman & Cahill, 1933. U. S. Geol. Surv. Prof. Pap. 175A) in the lower costae and more slender test in both mega- spheric and microspheric forms, and in the absence of spines on the 434 SCIENTIFIC SURVEY OF I'ORTO RICO fostao. The lip is also loss flaring. >S'. latncUala ( 'iishmaii (1918. U. S. Gcol. Surv. Bull. 676) has lamellae rather than costae. This species is named in honor of Professor E. R. Ciimings, who, as head of the Department of Geology at Indiana University, has fostered scientific research foi- many years. Siphogenerina hubbardi ^E^\ speciks Plate 34, figure 2 Test elongate, cylindrical, nearly circular in cross section; triserial portion about 3^-^ the length of the test, 4 chambers in uniserial part of the test; apical end either blunt in what is evidently the megaspheric form, or pointed in the microspheric form; most specimens with small spinosities near the apical ends of the costae; chambers slightly in- flated, uniserial chambers broader than long; sutures depressed in carl}^ part of the test, only slightly depressed in later part ; wall ornamented by about 20 longitudinal costae most of which are con- tinuous from chamber to chamber; aperture terminal, circular with rounded lip at the end of a definite neck. Total length, 1.00 milli- meter; diameter, 0.30 millimeter; length of neck, O.OG millimeter. Holotype No. 3985, locality P2, Ponce formation. Conmion at P2, P3, and P255 of the Ponce formation. aS'. hubbardi differs from *S. multicodata Gushman and Jar vis (1929. Contr. Cushman Lab. Foram. Res. 5) in that the costae are continuous from chamber to chamber and the apical spinosity is less pronounced. This species is named in honor of Dr. Bela Hubbard who has made important contributions to the stratigraphy and paleontology of Porto l^ico. Siphogenerina mexicana Cushman Plate 34, figures la, b Siphogenerina mexicana ('ushnian (1H2G) Proc. U. S. Nat. Mus. 67 (2.S): 15. pi. r>. figs. 4a. b. Lower Oligoceno, Alazan, Mexico. Test small, slender, subcylindrical, the first third of the test tri- serial, tapering from a subacute initial end, the rest of the t(>st uni- serial with about 5 rectilinear chambers; chambers distinct, very slightly inflated; sutures distinct, nearly flush with the surface, marked by clear shell material; wall with conspicuous pores, surface orna- mented with numerous fine, longitudinal striae; apertural end without a neck, truncate; aperture small, nearly circular, surrounded by a raised rim or lip. Length, 0.75 millimeter; diameter, 0.18 millimeter. Plesiotype No. 3986, locality P2, Ponce formation. Rare at P2 and P3 of the Ponce formation. GALLOWAY AND HEMINWAY, TERTLARY FORAMINIFKRA 435 Siphogenerina multicostata Ciishman & Jarvis Plate 34, figures 3 and 4 Siphouinerina multicoslata Cushman & Jarvis (1929) Contr. Cusliman Lab. Foram. Res. 5: 14. pi. 3, fig. 6. Miocene, Trinidad. — Cushman (1929) Contr. Cusliman Lab. Foram. Res. 5: 95, pi. 13, flg. 38. Middle Tertiary, Venezuela. — Hadley (1934) Bull Am. Paleont. 20 (70A) : 17. pi. 2. flg. 8. Oligocene, probably upper, Cuba. Test small, elongate, subcylindrical to fusiform, 3 to 4 times as long as wide, greatest ^vidth in the later half of the test; uniserial chambers occupying nearly half the length of the test; chambers distinct, slightly inflated, uniserial chambers shorter than Avide, except for the last chamber which is approximately equidimensional ; sutures de- pressed; surface ornamented by 18 to 20 narrow, low, longitudinal costae which extend up to the base of the neck; costae on each chamber usually independent of those on adjacent chambers; costae somewhat spinose on triserial part of the test; apertural end tapering to a short neck; aperture terminal, circular. Length, 0.74 millimeter; diameter, 0.20 millimeter. Plesiotypes Nos. 3987a and 3987b, locality P254, Ponce formation. Rare at P2 and P254 of the Ponce formation. The Porto Rican specimens have the phialine lip broken off. ANGULOGERINA Cushman 1927 Angulogerina cibaoensis new species Plate 34, figures 12a, b Test small, short, thick, bulbous at the lower end, apical end pointed and rapidly expanding to the greatest width at about two- fifths of the length of the test from the apical end, thence tapering to the apertural end; test triangular throughout, with angled edges; chambers regularly triserial and closely appressed in the earlj^ part, becoming biserial and tending to become uniserial, not inflated; sutures indistinct, little depressed; surface smooth at apical and aper- tural ends, provided with 3 or 4 small, low costae on each face in the middle of the test, about 4 in a width of 0.2 millimeter; the costae do not cross the sutures; aperture oval, at the end of a short neck, with small lip. Length, 0.62 millimeter; width, 0.36 millimeter; form ratio 1.7. Holotype No. 3864, locality A86, Cibao formation. Common at A86 of the Cibao formation. This species is shorter than A. cooperensis, with which it occurs, and lacks the apical costae. 43G SCIENTIFIC SURVEY OF POIiTO RICO Angulogerina cooperensis Cuslmiaii Plate 34, figures 13a, b Angulugerina cooperensis Cushnian (1935) U. S. Geol. Surv. Prof. Pap. 181: 42. pi. 16, fig. 9. Upper Eocene, South Carolina. Test of average size, slender, apical end pointed and rapidly ex- panding to the greatest width about one-third of the length of the test from the apioal end, thence tapering to the apertural end, tri- angular throughout, with angled edges; chambers regularly triserial in the first third, l)ecoming biserial and the final third uniserial, closely appressed but irregularly inflated; sutures indistinct, little depressed; wall finely perforate; surface ornamented with low, indis- tinct costae on the first two-thirds, disappearing on the final third, not crossing the sutures, 3 to 6 on each face of the test, 6 in a width of 0.2 millimeter; aperture at the end of a long neck, subtriangular, with slight lip. Length, 0.74 millimeter; width, 0.31 millimeter; form ratio 2.4. Plesiotype No. 4148, locality A86, Cibao formation. Rare at A86 of the Cibao formation. The Porto Rican form differs from the type figure of .1. cooperensis ill having the greatest width nearer the apertural end, and in having a longer neck. Such differences may be of varietal or even specific rank, particularly since the Porto Rican form is nearly twice as large as the tj'pe specimens from the Eocene. Angulogerina decorissima NEW species Plate 34, figures 8 to 10 Test small, short fusiform, tending to hv bulbous, greatest width a little below the middle of the test; triangular in cross section, sharply angled at the edges; chambers relatively few, closely appressed, flat on the faces of the test; sutures slightly depressed; surface ornamented by numerous striae which cover the entire test up to the base of the neck and arc discontinuous from chamber to chamber; neck sub- triangular of moderate l(>ngth; apertun^ terminal, round, with a slight l)hialine lip. Length of holotype, 0.53 millimeter; diameter, 0.28 millimeter. Holotype No. 38()5a, paratypes Nos. 3865b and 38C5c, locality A21, Quebradillas formation. Rare at A21 of the Quebradillas formation. This species has a slight resemblance to A. albalrossi ornata Cushman (1932. Contrib. Cushnian Lab. Forani. Res. 8), but is more regular GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 437 ill shape, wdth striae on all the chambers. It differs from A. occi- (Jentalis (Ciishmaii) (1933. U. S. Geol. Surv. Prof. Pap. 175A) iu having finer striae and less lobulate chambers with triangular cross section throughout the test. Angulogerina ponceana new species Plate 34, figures lla-c Test small, stout, 1^ times as long as wide; apical end broadly rounded; apertural end truncate; in end view subtriangular with rounded angles; chambers few, 4 to 6, regularly triserial, somewhat inflated, overhanging previous chambers; sutures depressed deeply in some places and shallow in others, undulating, with the uneven chamber inflation producing a knotty appearance to the test; wall finely perforate; surface ornamented with 9 to 12 low, obscure costae, mostly confined to the early and middle part of the test, all fading out on the last chamber; aperture terminal, with short neck set in an en- circling depression in the last septal face, with slight lip. Length, 0.45 millimeter; thickness, 0.33 millimeter. Holotype No. 4023, locahty P254, Ponce formation. Rare at P254 of the Ponce formation. This species differs from A. naranjoensis Cushman and Bermudez (1937. Contrib. Cushman Lab. Foram. Res. 13: 16, pi. 1, fig. 56) in the shorter form, broadly rounded apical end, less inflated chambers, and in the presence of low costae. It is much like the paratype of that species (figs. 57, 58), which seems to be costate. The authors say in the original description (p. 17), "wall generally smooth", and in the discussion again, "smooth wall". The costae in the Porto Rican form are so small and inconspicuous that they might ])e overlooked. TRIFARINA Cushman 1923 Trifarina bradyi Cushman Plate 35, figures 4a, b Bhabdogonium tricarinalum Brady (1884) Rep. Voy. ChaUenger Zool. 9: 525. pi. 67, figs. 1-3. Recent, Atlantic Ocean. Trifarina bradyi Cushman (1923) Bull. U. S. Nat. Mus. 104 (4) : 99. pi. 22, flgs. 3-9. Recent, CJulf of Me.\ico, Caribbean Sea, and off southeastern U. S.; (1929) Contr. Cushman Lab. Foram. Res. 5: 96. pi. 13, fig. 39. Middle Tertiary, Venezuela. Test small, elongate, tapering toward both apical and apertural ends; triangular in cross section, concave on all three sides, with carinae at the three angles running from the initial end to the aper- ture; chambers triserial for about H the length of the test; sutures 438 SCIENTIFIC SURVEY OF PORTO RICO not depressed; aperture terminal, round, at the end of a short neck. Length, 0.52 millimeter; diameter, 0.24 millimeter. Plesiotype No. 4005, locality P3, Ponce formation. Rare at P3 of the Ponce formation. Family PLEUROSTOMELLIDAE Reuss 1860 PLEUROSTOMELLA Reuss 1860 Pleurostomella bierigi Palmer & Bermudez Plate 35, figures 2a-c Pleurostomella bierigi Palmer & Bermudez (1936) Mem. Soc. Cubana HLst. Nat. 10: 294. pi. 17, figs. 7, 8. Lower Oligocene, Cuba. Test stout in appearance, oval in side view, nearly circular in end view, apical end obtusely angled or rounded, maximum width about midway of the test; chambers inflated, last one constituting more than half the length of the test; aperture in a depression of the final chamber, an arcuate slit, with subsidiary vertical notch. Length, 0.58 milli- meter; width, 0.41 millimeter; thickness, 0.43 millimeter. Plesiotype No. 4069, locality P3, Ponce formation. Rare at P3 and P254; common at P2 of the Ponce formation. Th{> Porto Rican forms are slightly different from the type figures of P. bierigi in that they are more rounded at the apical end. Pleurostomella elliptica new species Plate 35, figures 3a, b Pleurostomella alternans C^uslmian & Harris (not Schwager) (1927) Contr. Cusliman Lab. Forain. Kes. 3: 129. pi. 2,'>. (If^. 2H. Kocene, Trinidad. Test small, elongate, elliptical to fusiform in shape; 6 to 8 chambers in the test, regularly alternating, increasing gradually in size, Uttle inflated; sutures slightly depressed; surface unornamented ; aperture in an oval depression on the side of the last septal face, consisting of a lunate opening with two small teeth on the lower side. Length, 0.66 millimeter; diameter, 0.16 millimeter. Holotype No. 4070, locality P2, Ponce formation. Common at P2 of the Ponce formation. P. elliptica differs from P. alternans Sch wager (1866. Novara- p]xpcd. Geol. 2: 238. pi. 6, fig. 79, which we here designate as the type of P. alternans; not fig. 80) in that the test has fewer chambers, is less tapering and is more rounded at the apical end. It differs from P. jacksonensis Cushman and Applin (1926. Bull. Am. Assoc. Petr. GALLOWAY AND II EM I NW AY, TERTIARY FORAMINIFERA 439 Geol. 10) in the less pointed apical end and absence of any, "series of depressions in longitudinal lines". Pleurostomella gerontica NEW species Plate 35, figures la-d Test stout, regularly oval in front view, both apical and apcrtural ends narrowly rounded; subcircular in end view; 5 to 6 chambers in the test, biserially arranged, enlarging rapidly with the exception of the final chamber which is unusually small, the second and third chambers from the last constituting about % of the test; aperture subterminal, an arcuate slit in a slight depression on the septal face. Length, 0.67 miUimeter; width, 0.50 millimeter; thickness, 0.46 millimeter. Holotype No. 4071, locality P2, Ponce formation. Rare at P2 and P3 of the Ponce formation. We have several specimens, all of which have the small final cham- ber and distinctive aperture. P. gerontica is distinctive in the shape of its stout test, with small final chamber and subterminal, crescentic aperture without a tooth. In general appearance it is very similar to P. hierigi, but differs in the small final chamber and in the character of the apertiu'c. NODOSARELLA Rzehak 1885 Nodosarella constricta granulifera new vahietv Plate 35, figukes 5a, h , This variety dift'ers from typical A^. constricta Cushman and Ber- mudez (1937. Contr. Cushman Lab. Foram. Res. 13) in having a finely granular surface. Length, 0.83 millimeter; diameter, 0.33 millimeter. Holotype No. 4059, locality P2, Ponce formation. Rare at P2 of the Ponce formation. Nodosarella crassielegans (Nuttall) Plate 35, figures 6a, b Nodosaria crassielegans Nuttall (1928) Quart. Jour. Geol. Soc. London 84: 80. pi. 4, figs. 6, 7. Upper Eocene and lower Miocene, Trinidad. Test elongate, arcuate, round in section, consisting of 6 to 10 chambers which are longer than thick, and more closely appressed in the early part of the test; sutures transverse, depressed, marked by 440 SCIENTIFIC SURVEY OF PORTO RICO clear shell material; apical end of the test with one, rarely two, short, eccentric spines; surface smooth; aperture produced, crescentic. Length, 1.77 millimeters; diameter, 0.26 millimeter. Plesiotype No. 4078, locality P3, Ponce formation. Common at P3 of the Ponce formation. This species differs from N. paucistriata Galloway and Morrey in the absence of the septal bridges, and from N. cocoaensis (Cushman) in the inflated chambers. Nodosarella paucistriata Galloway & Morrey Plate 35, figures 7 to 9 ? Nodosaria catenulata Cushman (1923) U. S. Geol. Surv. Prof. Pap. 133: 28. pi. 3, fig. 14. Lower Oligocene, Alabama. Nodosaria intermittens Nuttall (1928) Quart. Jour. CJeol. Soc. London 84: 82. pi. 4, fig. 17. Upper Eocene and lower Miocene, Tiinidad. Nodosarella paucistriata Galloway & Morrey (1929) Bull. Am. Paleont. 15 (55): 42. pi. 6, figs. 12a, b. Lower Oligocene, Ecuador. — Coryell & Rivero (1940) Jour. Paleont. 14: 343. pi. 42, figs. 21-23. Middle Miocene, Haiti. Ellipsonodosaria verneuili paucistriata Cushman (1929) Contr. Cushman Lab. Foram. Res. 5: 97. pi. 14. figs. 4, 5. Middle Tertiary, Ecuador and Trinidad. Test elongate, slender, slightly curved; about 11 chambers in the adult test, most of the chambers inflated; sutures depressed, or flush with the surface between the early uninflated chambers; apical end blunt, with a short, stout, eccentic spine; surface unornamcnted except for about 6 septal bridges on the depressed sutures, flush sutures unornamented; aperture terminal, crescentic in shap(\ at the end of a neck-like constriction of the last chamber. TiCngth, up to 2.58 millimeters; diameter, up to 0.40 millimeter. l^lcsiolypcs No. 4058a and 4058b, locality P3; and No. 3891, locality P2, Ponce formation. Common at A86 of the Cibao formation. Rare at P3; common at P2, P254, and P255 of the Ponce formation. The septal bridges of this species arc very small and easil}-^ over- looked. Nodosarella verneuili (d'Orbigny) Plate 35, figures 10a, b Dentalina verneuili d'Orbigny (184C) Foram. Foss. Vlenne 48. pi. 2, flgs. 7, 8. Middle Miocene, Vienna. Ellipsonodosaria verneuili Cushman (1929) Contr. Cushman Lab. Foram. Res. 5: 96. pi. 14, flg.s. 1-3. Middle Tertiary, Ecuador, Venezuela, Trinidad. — Cushman & Jarvis (1930) Jour. Paleont. 4: 364. pi. 33, flg. 12. Middle Miocene, Jamaica. — Palmer & Bermudez (1936) Mem. Soc. Cubana Hist. Nat. 10: 295. pi. 18, flgs. 1, 2. 14-16. Lower Oligocene, Cuba. Test slender, elongate, slightly curved, gradually tapering from the initial end which terminates in a spine; early part of the test with GALLOWAY AND HEM I NW AY, TERTIARY FORAMINIFERA 441 parallel, straight sides, later part with the chambers slightly inflated; chambers increasing rather slowly in size, being usually slightly shorter than wide; sutures distinct, of clear shell material, flush with the sur- face in the early part of the test and slightly depressed in the later part; apertural end produced; aperture terminal, crescentic, with small, pointed tooth. Length of average specimen, 1.33 millimeters; diameter, 0.25 millimeter. Plesiotype No. 4034, locaUty A86, Cibao formation. Common at A86 of the Cibao formation. ELLIPSOGLANDULINA Silvestri 1900 Ellipsoglandulina exponens (Brady) Plate 36, figures la, b EUipsoidina exponens Brady (1892) in Jukes-Brown & Hanison, Quait. Jour. Geol. Soc. 48: 198. Upper Tertiary, Barbados. — Guppy (1894) Proc. Zool. Soc. London, 650. pi. 41, fig. 13. Miocene, Trinidad. Ellipsoglandulina laevigata Silvestri (1900) Att. Rend. R. Acad. Sci. Lett. Arti. Zelanti, Acireale, Me. CI. Sci. 10 : 9. pi. 1, figs. 3-10, 12, 13. Upper Miocene or basal Pliocene, Sicily. Ellipsoglandulina erponens Nuttall (1928) Quart. Jour. Geol. Soc. 84: 95. pi. 6. flg. 17. Oligocene and Miocene, Trinidad. — Cushman (1928) Contr. Cusliman Lab. Foram. Res. 4: 103. pi. 14, flg. 17. Upper Cretaceous, Trinidad. Ellipsoglandulina principiensis Cushman & Bermudez (1937) Contr. Cushman Lab. Foram. Res. 13: 18. pi. 2, figs. 1-3. Eocene, Cuba. Test oval in side view, narrowly rounded at the apical end, slightly tapering at the apertural end; chambers 3 to 4, overlapping so that the final chamber constitutes nearly one-half of the test; sutures distinct, .slightly depressed; aperture terminal, crescentic. Length, 1.20 milli- meters; diameter, 0.70 millimeter. Plesiotype No. 3903, locality P2, Ponce formation. Rare at P2 of the Ponce formation. ELLIPSOIDINA Silvestri 1923 EUipsoidina ellipsoides abbreviata Seguenza Plate 36, figures 2 and 3 EUipsoidina abbreviata Seguenza (1859) Eco. Peloritano II. 5: 14. pi., figs. 5a, b. Miocene, Italy. EUipsoidina ellipsoides abbreviata Brady (1868) Ann. Mag. Nat. Hist. IV. 1: pi. 13, flg. 3. Miocene, Italy.— Silvestri (1925) Atti Soc. Ital. Sci. Nat. 54: 51. text flg. 1. Molasse, Tertiary. Italy. EUipsoidina e/hpsoides Hadley (not Seguenza) (1934) Bull. Am. Paleont. 20 (70A) : 19. pi. 2. figs. 18, 19. Oligocene, probably upper, Cuba. Test smoothly ovoid, apical end slightly more pointed than apertural end; chambers uniserial, enlarging rapidly, last chamber completely embracing earlier chambers; aperture terminal, an arcuate slit with 442 SCIENTIFIC SURVEY OF PORTO RICO raised lip; shape of slit varies from a short arc to a short U or V shape. Length, 1.35 iiiillinieters; diameter, 1.17 milhmeters. Plesiotypes No. 3904a and 3904b, locahty P2, Ponce formation. Eare at P3 and P255; abundant at P2 of the Ponce formation. K. abbreriata is, as the name indicates, much shorter in proportion than E. eUipsoides Seguenza (1859. Eco. Peloritano II. 5:). Family CAMERINIDAE ^k-ek it Hayden 1865 OPERCULINELLA Yabe 191S Operculinella sinuata new SPECIES Plate 36, figures 4a-c Test very small fur the genus, compressed, biconvex, biumbonate, nearly bilaterally symmetrical; edge narrowly rounded but not keeled; periphery smooth; whorls about two, involute to umbos, only the last whorl exposed, part of the earlier whorl visible through the clear umbonal material; chambers 10 to 12 in the last whorl; sutures distinct, marked by clear shell material, flush with the surface, strongly curved, meeting the periphery with a small angle, recurved onto the umbo, making the sutures sigmoid in the last half whorl; the two sides of the test nearly identical; wall very finely perforate; aperture a small sUt, ^\^th small upper lip, at the base of the septal face on the inner periphery, not extending beyond the edge of the test on either side. Diameter, 0.55 millimeter; thickne.>^s, 0.16 millimet(M-. Holotype No. 4067. locahty A21, Quebradillas formation. Rare at A2() and A21 of the Quebradillas formation. There are few described species with which to compare this Porto Eican form. It is smaller and has fewer chambers than O. -v I b L ft 2 c 2 b 9 c 9 b GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 451 Plate 2 PAGE Figure 1. Quinqueloculina akneriana d'Orbigny. X 40; a, side view; b, opposite side; c, apertural view; Plesiotype No. 3961, locality L6C, San Sebas- tian formation. ''"^ P'iGURE 2. Quinqueloculina apiculata n. sp. X 56; a. side view; b, opposite side; c, apertural view; Holotype No. 3962, locality L6C, San Sebastian forma- tion. 302 Figure 3. Quinqueloculina maculata n. sp. X 28; a, side view; b, opposite side: c, apertm-al view; Holotype No. 3965, locality L6C, San Sebastian forma- tion. . 303 Figure 4. Qumqmlondina philippi Reuss. X 40; a and c, side views; b, edge view; d, apertural view; Plesiotype No. 3966, locality A79, Los Puertos formation. ^^"^ Figure 5. Quinqueloculina ponceana n. sp. X 76; a, side view; b, opposite side; c, apertural view; Holotype No. 3967, locality P2, Ponce formation. 304 Figure 6. Quinqueloculina vulgaris d'Orbigny. X 76; a, side view; b, opposite side; c, apertural view; Plesiotype No. 3970, locality L6C, San Sebas- tian formation. ^^^ Figure 7. Quinqueloculina pijgmma Reuss. X 56; a, side view; b, opposite side; c, apertural view; Plesiotype No. 3968, locality L6C, San Sebastian for- mation. 304 Figure 8. Quinqueloculina seminulum (Linne). X 40; a, side view; b, oppo- site side; c, apertural view; Plesiotype No. 3969, locality L6C, San Sebastian formation. 305 Figure 9. Triloculina elliptica n. sp. X 56; a and b, side view; c, edge view; d, apertural view; Holotype No. 4009, locality LlC, San Sebastian formation. 307 452 SCIENTIFIC SURVEY OF PORTO RICO Plate 3 PAGE Figure 1. Trilocuhna avslriaca d'Orbigny. X 40; a, side view; b, opposite side; c, apertural view; Plesiotype No. 4006, locality L6C, San Sebastian for- mation. 30G Figure 2. Trilocuhna brongniaTtiana d'Orbigny. X 27; a, side view; b, op- posite side; c, apertural view; Plesiotype No. 4007, locality P258, Ponce for- mation. 306 Figure 3. Triloculina cervicula, n. sp. X 56; a, side view; b, opposite side; c, apertural view; Holotype No. 4008, locality L6C, San Sebastian formation. 306 Figure 4. Triloculina gibba d'Orbigny. X 40; a, edge view; b, side view; c, apertural view; Plesiotype No. 4010, locality L4C, San Sebastian formation. 307 Figxtre 5. Triloculina injlala d'Orbigny. X 56; a, side view; b, opposite side; c, apertural view; Plesiotype No. 4011, localitj' L3C, San Sebastian formation. 307 Figure 6. Triloculina laevigata d'Ovhi^ny . X 40; a, side view; b, opposite side; c, apertural view; Plesiotype No. 4013, locality L4C, San Sebastian for- mation. 308 Figure 7. Triloculina longis.sima, n. sp. X 56; a, side view; b, opposite side; c, apertural view; Holotj'pe No. 4012, locality L6C, San Sebastian formation, 308 FiGi'RE 8. Triloculina oblonga (Montagu). X 56; a, side view; b, opposite side; c, apertural view; Plesiotype No. 4014, locality L6C, San Sebastian for- mation. 309 Figure 9. Triloculina qnadnlateralis n. sp. X 40; a, side view; b, opposite side; c, apertural view; Plesiotype No. 4015, localitj- L6C, San Sebastion for- mation. 309 Figure 10. Tnloculina tricar inala d'Orbigny. X 53; a, edge view; b, aper- tural view; Plesiotype No. 4016, locality P253, Ponce formation. 309 Scientific Survey of Porto Rico and the Virgin Islands Volume III, Part 4, Plate 3 / 2 2 b 3 Q a 2 c 3 b 1^ 5 Q "Ky 7 a ■'V-^X 7 \^ 8 b 5 c 5 b x\ 9 b 9 GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA Scientific Survey of Porto Rico and the Virgin Islands Volume III, Part 4, Plate 4 I C N 2 c I b i I 2 Q ^■' 2 b -' 3 c V I ! ) 5 Q 5 c 4 a ( y 7c V 6 b ^ -fQ \ J V 6 c 9 b ■■" — »J>'*^9 c 8 c /\ >v ■^1 8 8 b 9 Q GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 453 Plate 4 PAGE Figure 1. Pyrgo hougainvillei (d'Orbigny). X 56; a, front view; b, side view; c, apertural view; Plesiotype No. 3953, locality L6C, San Sebastian for- mation. 310 Figure 2. Pyrgo dypeata (d'Orbigny). X 56; a, front view; b, side view; c, apertural view; Plesiotype No. 3954, locality A79, Los Puertos formation. 311 Figure 3. Pyrgo inomata (d'Orbigny). X 53; a, front view; b, side view; c, apertm-al view; Plesiotype No. 3955, locality A79, Los Puertos formation. 311 Figure 4. Pyrgo lunula (d'Orbigny). X 40; a, front view; b, side view; c, apertural view; Plesiotype No. 3956, locality P258, Ponce formation. 311 Figure 5. Pyrgo oblonga (d'Orbigny). X 76; a, front view; b, side view; c, apertural view; Plesiotype No. 3957, locality P251, Ponce formation. 312 Figure 6. Pyrgo peruviana (d'Orbigny). X 40; a, front view; b, side view; c, apertural view; Plesiotype No. 3958, locality L4C, San Sebastian formation. 312 Figure 7. Pyrgo sutspherica (d'Orbigny). X 56; a, front view; b, side view; c, apertural view; Plesiotype No. 3959, locality L2C, San Sebastian for- mation. 313 Figure 8. Miliolinella circularis (Bornemann). X 40; a, side view; b, oppo- site side; c, apertural view; Plesiotype No. 3997, locality P2, Ponce formation. 310 Figure 9. Pyrgoella globulus (Bornemann). X 56; a, front view; b, side view; c, apertural view; Plesiotype No. 3960, locality P255, Ponce formation. 313 454 SCIENTIFIC SURVEY OF PORTO RICO Plate 5 PAGE Figure 1 . Maasilina ikcorala. Cushman. X 40; Plesiotype No. 3932, locality A79, Los Puertos formation. 314 Figure 2. Massilina inaequalis d'Orbigny. X 25; a, side view; b, opposite side; c, apertural view; Plesiotype No. 3933, locality A79, Los Puertos formation. 314 Figure 3. Haumna mnsebasliannif;if< n. sp. X 25; Immature specimen: a, side view; b, apertural view; c, opposite side; Paratype No. 3925b, locality L6C, San Sebastian formation. 315 Figure 4. Hauerina sansebastianensin n. sp. X 25; Adult specimen: a, side view; b, apertural view; c, opposite side; Holotype No. 3925a, locality L6C, San Sebastian formation. 315 Figure 5. Dtndritma ])rcelegan.s n. sp. X 20; a, side view; b, apertui-al view; Holotype No. 3889, locality A79, Los Puertos formation. 315 Figure 6. PenewpHis rarinatus d'Orbigny. X 28; a, side view; b, apertural view; Plesiotype No. 3945, locality LlC, San Sebastian formation. 316 Figure 7. Peneroplis jrrotevs d'Orbigny. X 28; a, side view; b, apertui-al view; Plesiotype No. 3946, locality L6C, San Sebastian formation. 316 Figure 8. ArcJiaias adunaia (Fichtel & Moll). X 20; a, side view; b, aper- tural view; c, portion of surface showing the chamberlets; Plesiotype No. 3868, locality P258, Ponce formation. 317 Figure 9. Archaias anyidalus (Fichtel & Moll). X 40; a, side view; b, aper- tural view; Plesiotype No. 3869, locality LlC, San Sebastian formation. 318 Figure 10. Archmas comprc'^tius (d'Orbigny). X 20; a, side view; b, edge view; Plesiotype No. 3870, locality LlC, San Sebastian formation. 318 Scientific Survey of Porto Rico and the Virgin Islands Volume III, Part 4, Plate S \ >v. \v y J 2 c ; \ f 4 Pi 3 Q 3 b Vr~^ ; 3 c •*» - 9 a 9 b 10 b 10 a GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA Scientific Survey of Porto Rico and the Virgin Islands Volume III, Part 4, Plate 6 k^ .^\ ^ ' GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 455 Plate 6 PAGE Figure 1. Trochamminmdes approximatus n. sp. X 53; a, side view; b, aper- tural view; Holotype No. 4064, locality A91a, Los Puertos formation. 319 Figure 2. Cydammina acutidorsata (Hantken). X 40; a, side view; b, aper- tural view; Plesiotype No. 3887, locality P3, Ponce formation. 319 Figure 3. Valvulina oviedoiana d'Orbigny. X 25; a, side view; b, apertural view; Plesiotype No. 4026, locality L6C, San Sebastian formation. 320 Figure 4. V alvtdammina cornucopia n. sp. X 25; a, dorsal view; b, side view; c, ventral view; Holotype No. 4025, locality L6C, San Sebastian formation. 320 Figure 5. Verneuilina mexicana Nuttall. X 43; Plesiotype No. 4031, locality P2, Ponce formation. 321 Figure 6. Dorothia caribaea Cushman. X 27; a, side view; b, apertural view; Plesiotype No. 3901, locality P255, Ponce formation. 322 Figure 7. Dorothia cylindrica (Nuttall). X 27; a, side view; b, edge view; c, apertural view; Plesiotype No. 3916, locality P2, Ponce formation. 322 Figure 8. Dorothia pra^longa (Karrer). X 40; a, side view; b, apertural view; Plesiotype No. 4046, locality P2, Ponce formation. 322 Figure 9. Liebusella byramensis (Cushman). X 14; a, side view; b, aper- tural view; Plesiotype No, 4132, locality P255, Ponce formation. 325 Figure 10. Listerella cf. communis (d'Orbigny). X 27; a, side view; b, apertural view; Plesiotype No. 4056, locality P255, Ponce formation. 326 456 SCIENTIFIC SURVEY OF PORTO RICO Plate 7 PAGE Figure 1. Gaudryina asiphonia Andreae. X 27; a, side view; b, apertural view; Plesiotype No. 4045, locality P2, Ponce formation. 323 Figure 2. Gmidryina glabrata maxima n. var. X 27; a, side view; b, aper- tural view; Holotype No. 4050, locality A43a, Cibao formation. 323 Figure 3. Gaudryina {P neudogaudr^jina) atlantica (Bailey). X 25; a, side view; b, apertural view; Plesiotype No. 4085, locality P3, Ponce formation. 325 Figure 4. Clavulinoides polygonahs n. sp. X 40; a, side view; b, apertural view; Holotype No. 4091, locality F64, Quebradillas formation. 327 Figure 5. Clavulinoides triangularis (Nuttall). X 27; a, side view; b, aper- tural view; Plesiotype No. 4086, locality P3, Ponce formation 327 Figure 6. Gaudryina karreriana Cushman. X 20; a, side view; b, apertural view; Plesiotype No. 3917, locality P2, Ponce formation. 324 Figure 7. Gaudryina puertoricana n. sp. X 77; a, side view showing aper- ture; b, opposite side; Holotype No. 3918, locality L6C, San Sebastian formation. 324 Figure 8. Clavulina tricarinata d'Orbigny. X 40; a, side view; b, apertural view; Plesiot3^pe No. 3885, locality L3C, San Sebastian formation. 326 Figure 9. Tcxhdariella barrettii Jones & Parker. X 27; a, side view; b, aper- tural view; Plesiotype No. 4004, locality A86, Cibao formation. 332 Figure 10. Vulvidina pachyheilus Hadley. X 40; a, side view; b, apertural view; Plesiotype No. 4032, locality A93, Quebradillas formation. 332 Figure 11. Vulvuhna pcnnatula (Batsch). X 20; a, side view; b, apertural view; Plesiotype No. 4033, locality P2, Ponce formation. 333 Scientific Survey of Porto Rico and the Virgin Islands Volume III, Part 4, Plate 7 % ^^4b 6i ;^^iiy A a m \. v^ 5 b 5 Q / a r 7 b 8 b ^■«F*>- 9 b 8 Q 9 a lO a II b GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA Scientific Survey of Porto Rico and the Virgin Islands Volume III, Part 4, Plate 8 ^:# 2 a r f^' 2c y^v \ •^. 3 b 4 a /^rd:^' V / 4 c "^ 4 b 2 b 5 5 c / 5 b 30 i i ^'^' ^*^" ea 8c -^ 8 b i^ 9 b \,, 10 b Mb 9a 10 a II GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA GALLOWAY AND HEMINWAY, TERllARY FORAMINIFERA 457 Plate 8 PAGE Figure 1. Textularia articulata d'Orbigny. X 29; a, side view; b, edge view; Plesiotype No. 3998, locality LlC, San Sebastian formation. 329 Figure 2. Textularia agghitinans d'Orbigny. X 45; a, side view; b, edge view; c, apertural view; Plesiotype No. 3996, locality LlC, San Sebastian forma- tion. 328 Figure 3. Tcvtularia broussardi Howe & Wallace. X 27; a, side view; b, apertural view; Plesiotype No. 4092, locality P2, Ponce formation. 329 Figure 4. Textularia gramen d'Orbigny. X 45; a, side view; b, edge view; c, apertui-al view; Plesiotype No. 4003, locality LlC, San Sebastian formation. 330 Figure 5. Textularia candeana d'Orbigny. X 45; a, side view; b, edge view; c, apertural view; Plesiotype No. 3999, locality L3C, San Sebastian formation. 329 Figure 6. Textularia grenadava Hedberg. X 27; a, side view; b, apertm-al view; Plesiotype No. 4002a, locality P3, Ponce formation. 330 Figure 7. Textularia grenadana Hedberg. X 27; apertural view of another specimen; Plesiotype No. 4002b, locality P3, Ponce formation. 330 Figure 8. Textularia indenta n. sp. X 20; a, side view; b, edge view; c, aper- tural view; Holotype No. 4051, locality A43a, Cibao formation. 331 Figure 9. Astacolus insolitus (Schwager). X 40; a, side view; b, apertural view; Plesiotype No. 4037, locality P254, Ponce formation. 334 Figure 10. Astacolus ovatus n. sp. X 53; a, side view; b, apertural view; Holotype No. 3871, locality A86, Cibao formation. 334 Figure 11. Astacolus sublituus (Nuttall). X 53; a, side view; b, apertural view; Plesiotype No. 3872, locality A86, Cibao formation. 335 > 4U89; 458 SCIENTIFIC SURVEY OF PORTO RICO Plate 9 PAGE Figure 1. Margirmlina insnlensis n. sp. X 53; a, side view; b, apertural view; c, apical view; Holotype No. 4062, locality P2, Ponce formation. 337 Figure 2. Marginulina subcrassa Schwager. X 40; a, side view; b, aper- tural view; Plesiotype No. 4057, locality P2, Ponce formation. 337 Figure 3. Vaginulina faha n. sp. X 27; a, side view; b, apertural view; c, apical view; Holotype No. 4024, locality P3, Ponce formation. 335 Figure 4. Vaginulina mexicana Nuttall. X 20; a, side view; b, apertural view; Plesiotype No. 4090, locality P3, Ponce formation. 336 Figure 5. Vagmidina siliqiwidm n. sp. X 53; a, side view; b, apertural view; Holotype No. 4052, locality P2, Ponce formation. 336 Figure 6. Saraccnaria arcuata (d'Orbigny). X 27; a, side view; b, front view; c, apertural view; Plesiotype No. 3982, locality P3, Ponce formation. 333 Figure 7. H cmicristellaria fragana (Giimbel). X 27; a, side view; b, aper- tural view; Plesiotype No. 4053, locality A21, Quebradillas formation. 335 Figure 8. Amphicoryne obhqua n. sp. X 53 ;a, side view; b, apertural view; Holotype No. 3861, locality P2, Ponce formation. 338 Figure 9. Denlali'na advcna Cushman. X 27; a, side view; b, apertural view; Plesiotype No. 3895. locality P254, Ponce formation. 339 P'iGURE 10. Dentalina halkyardi Cushman. X 40; a, side view; b, apertural view; Plesiotype No. 3892, locality P2, Ponce formation. 340 Figure 11. Dnilahna halkyardi ponccana n. var. X 20; a, side view; b, apertural view; Holotype No. 3893, locality P3, Ponce formation. 340 Figure 12. Dmtalma hinaeformm n. sp. X 53; Holotype No. 3890, locality P2, Ponce formation. 341 Figure 13. Dentalina nnilhlincala Rornomann. X 40; Plesiotype No. 4035, locality P254, Ponce formation. 341 Figure 14. Dentalina semilaevis Hantken. X 20; Plesiotype No. 3894, lo- cality P3, Ponce format inn. 342 Scientific Survey of Porto Rico and the Virgin Islands Volume III, Part 4, Plate 9 I b /'■'"l 6 b 9b 10 b 9 a •""> 3b II b 10 a -^ 7 a II a p iiin 14 GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA Scientific Survey of Porto Rico and the Virgin Islands Volume III. Part 4, Plate 10 \. 5 a 5 c / / >^a- 5 b Wa -i lOa 6 b /rv\ 6 6 c l-^ -^ o %.. t« 9 b A ^ \ r;f '^* 12 b 2 a JHT II 3 a ^«K.»*' 13 b '-i .. 3 c GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 459 Plate 10 PAGE Figure 1. N onion jvicoharense Cushman. X 82; a, apertural view; b, side view; Plesiotype No. 3938, locality P254, Ponce formation. 357 Figure 2. Nodosaria halkyardi antiUana Palmer & Bermudez. X 45; Plesiotype No. 4060, locality A86, Cibao formation. 342 Figure 3. Nodosaria simplex Silvestri. X 45; Plesiotype No. 4061, locality P2, Ponce formation. 344 Figure 4. Nodosaria scalaris (Batsch). X 40; Plesiotype No. 3936, locality P255, Ponce formation. 344 Figure 5. PolelUna corrugata Williamson. X 82; a, dorsal view; b, ventral view; c, edge view; Plesiotype No. 3944, locality P432, Ponce formation. 299 Figure 6. Glandulina comatvla (Cushman). X 40; immatiu-e specimen: a, side view; b, apertural view; c, apical view; Plesiotype No. 3919b, locality P2, Ponce formation. 338 Figure 7. Glandulina mauricensis (Howe & Roberts). X 57; Plesiotype No. 4047, locality P254, Ponce formation. 339 Figure 8. Lagena nuttalli n. sp. X 40; a, side view; b, apertural view; Holo- type No. 4054, locality P254, Ponce formation. 346 Figure 9. Lagena strumosa Reuss. X 40; a, side view; b, apertural view; Plesiotype No. 4055, locality P255, Ponce formation. 346 Figure 10. Lagena inipi^essa n. sp. X 40; a, side view; b, opposite side; c, apertural view; Holotype No. 4094, locality P2, Ponce formation. 346 ■ Figure 11. Lrngulina semicostata n. sp. X 82; a, side view; b, apertural view; Holotype No. 3928, locality P2, Ponce formation. 353 Figure 12. Antillesina marielensis (Palmer). X 82; adult form: a, aper- tural view; b, side view; Plesiotype No. 3943, locality P254, Ponce formation. 367 Figure 13. Antillesina marielensis (Palmer). X 82; young specimen: a, apertural view; b, back view; c, side view; Plesiotype No. 4063, locality LlC, San Sebastian formation. 367 460 SCIENTIFIC SURVEY OF PORTO RICO Plate 11 PAGE Figure 1. Glatuhdinn comaiula (Cushman). X 27; a, side viow; h, aper- tural view; Plesiotype No. 3919a, locality P2, Ponce formation. 338 FiGUBE 2. GlanduHna gcdlowayi (Cushman). X 52; a, side view; h, ai)er- tural view; Plesiotype No. 3951, locality A86, Cibao formation. 338 Figure 3. Fisfiiirinn lacvis Seguenza. X 76; a, side view; b, apcrturai view; Plesiotype No. 4043, locality P2, Ponce formation. 353 Figure 4. Fissurina marginata (Montagu). X 53; a, side view; h, apertural view; Plesiotype No. 4044, locality P254, Ponce formation. 353 Figure 5. Nodomria nhhquntn (Batsch). X 40; Plesiotype No. 3935, locali- ty P3, Ponce formation. 343 Figure 6. Nodosaria raphanintnim (Linne). X 15; Plesiotype No. 4097, locality P3, Ponce formation. 344 Figure 7. Nodosaria parinna Hedberg. X 40; Plesiotype No. 4099, locality P254, Ponce formation. 343 Figure 8. Nodosaria Umgiscata d'Orbigny. X 27; Plesiotype No. 3934, locality P3, Ponce formation. 342 Figure 9. Lngoia amjmlla n. sp. X 56; a, side view; b, apical view; Holo- type No. 3926, locality L6C, San Sel)astian formation. 345 Figure 10. Lagcna bullosa n. sp. a, side view, X 40; b, apertural \iew, X 80; Holotype No. 4079, locality P2, Ponce formation. 345 Figure 11. Lingvlina ponrcann n. sp. X 20; a, side view; b, apertui-al view; Holotype No. 3927, locality P3. Ponce formation. 352 Figure 12. Robulm calliferus (Stache). X 27; a, side view; b, apertural view; Plesiotype No. 4096, locality P2, Ponce formation. 347 Figure 13. Rnhulus prntubvrnns (Cushman). X 14; a, side view; 1), apertural view; Plesiotype No. 3980, locality P2, Ponce formation. 351 Figure 14. Robulus planulus n. sp. X 53; a, side view; b, apertural view; Holotype No. 3979, locality A86, Cibao formation. 350 Scientific Survey of Porto Rico and the Virgin Islands Volume III, Part 4, Plate 11 .Mk. 'Si*^ I a ■^im^-*-^' 2 b 4 b 3 b 4 a 10 b -N, 9 b I Q e*" 13a 13 b A \^ . V ■^-^- ,2 a GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA (4 b Scientific Survey of Porto Rico and the Virgin Islands Volume III, Part 4, Plate 12 -i lb 2a ^ 2b ~-^ 30 A K \ 4a ▼ 4b *\ ■* V / 7a \ 8a "^ 8b i:! I 9a GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 461 Plate 12 PAGE Figure 1. Rolulus chambersi Garrett. X 43; a, side view; b, apertural view; Plesiotype No. 3973, locality P254, Ponce formation. 347 Figure 2. Robuhis subpaptllosus (Nuttall). X 30; a, side view; b, apertural view; Plesiotype No. 4095, locality P3. Ponce formation. 351 Figure 3. Robulus convergens (Bornemann). X 28; a, side view; b, aper- tural view; Plesiotype No. 3974, locality P3, Ponce formation. 348 Figure 4. Robulus occidentalis torridus (Cushman). X 43; a, side view; b, apertural view; Plesiotype No. 4096, locality P3, Ponce formation. 349 Figure 5. Robulus fakifer (Stache). X 53; Plesiotype No. 4074a, locality A21, Quebradillas formation. 349 Figure 6. Robulus fakifer (Stache). X 53; apertural view of another speci- men; Plesiotype No. 4074b, locality A21, Quebradillas formation. 349 Figure 7. Robuhis iota (Cushman). X 20; a, side view; b, apertural view; Plesiotype No. 3976, locality P2, Ponce formation. 349 Figure 8. Robulus cibaocnsis n. sp. X 53; a, side view; b, apertural view; Holotype No. 4036, locality A86, Cibao formation. 348 Figure 9. Robuhis ■plummerae Cole. X 53; a, side view; b, apertural view; Plesiotype No. 3977, locality P254, Ponce formation. 350 462 SCIENTIFIC SURVEY OF PORTO RICO Plate 13 PAGE Figure 1. Ainopterina cylindroides (Roemer). X 76; a, side view; b, oppo- site side; c, apertural view; Plesiotype Xo. 4089, locality P259, Ponce formation. 355 Figure 2. Raphnnulina gibba glohosa (Von Miinster). X 53; a, side view; b, oppot^ite side; c, apertural view; Plesiotype No. 3971, locality L2C, San Sebas- tian formation. 355 Figure 3. Pobjinorpbina terquemiana Fornasini. X 76; a, side view; b, op- posite side; c, apertural view; Plesiotype No. 4088, locality A79, Los Puertos formation. 543 Figure 4. GuUulina basalis n. sp. X 40; a, side view; b, opposite side; c, apertural view; Holotype No. 4049, locality A21, Quebradillas formation. 354 Figure 5. Noniondla niodcsta n. sp. X 77; a, side view; b, opposite side; c, apertural view; Holotype No. 3941, locality LlC, San Sebastian formation. 359 Figure 0. Nonion dilutatum n. sp. X 76; a, side view; b, apertm-al view; c, opposite side; Holotype No. 4081, locality A43a, Cibao formation. 356 FiGX"RE 7. Nonion nitdHporatunt n. sp. X 76; a, side view; b, apertural view; c, opiK)site side; Holotype No. 4083, locality A43a, Cibao formation. 356 Figure 8. Nonion chapnpotense Cole. X HO; a, side view; b, apertural view; Plesiotype No. 3937, locality L6C, San Sebsatian formation. 355 Scientific Survey of Porto Rico and the Virgin Islands Volume III, Part 4, Plate 13 GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA Scientific Survey of Porto Rico and the Virgin Islands Volume III, Part 4, Plate 14 4 Q i \ ?y 2 4 b 2 b 3 Q ,<»-A •"..; '»... ■•". ^ I"*'"' 9 b 9 Q 10 b 10 GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA GALLOWAY AND HEMfNWAY, TERTIARY FORAMINIFERA 463 Plate 14 PAGE Figure 1. Nonion povipilioidcs (Fichtel & Mf)ll). X 58; a, side view; b, apertuial view; Plesiotype Xo. 3939, locality P2, Ponce formation. 357 Figure 2. Nonion subgrateloupi n. sp. X 56; a, side view; b, apertural view; Holotype No. 3940a, locality L5C, San Sebastian formation. 358 Figure 3. Nonion mhgraidoupi n. sp. X 5G; a, side view; b, apertvual view; Paratype No. 3940b, locality L5C, San Sebastian formation. 358 Figure 4. Pseudononion papillatum n. sp. X 53; a, side view; b, opposite side; c, apertural view; Holotype No. 4072, locality A43a, Cibao formation. 358 Figure 5. Elphidiimi, naiUiloideum n. sp. X 55; a, side view; b, apertm-al view; Holotype No. 3907, locality L5C, San Sebastian formation. 362 Figure 6. Elphidiwn poeyanvm. (d'Orbigny). X 55; a, side view; b, aper- tural view; Plesiotype No. 3909, locality LlC, San Sebastian formation. 3(j3 Figure 7. Elphidium lameri (d'Orbigny). X 30; a, side view; b, apertural view; Plesiotype No. 4102, locality P258, Ponce formation. 360 Figure 8. Elphidium lobatum n. sp. X 76; a, side view; b, apertural view; Holotype No. 3906, locality A6, Lares formation. 362 Figure 9. Elphidium owcmianmn (d'Orbigny). X 40; a, side view; b, aper- tural view; Plesiotype No. 3908, locality L5C, San Sebastian formation. 363 Figure 10. Elphidium lens n. sp. X 40; a, side view; b, apertural view; Holotype No. 3905, locality L4C, San Sebastian formation. 361 464 SCIENTIFIC SURVEY OF PORTO RICO Plate 15 PAGE Figure 1. Elphidium sagrni crassum n. var. X 56; a, side view; b, apertural view; Holotype No. 3910, locality LlC, San Sebastian formation. 366 Figure 2. Elphidium puertoricaisc n. sp. X 56; a, side view; b, apertural view; Holotype No. 3911, locality L6C, San Sebastian formation. 364 Figure 3. Elphidium. sagrai (d'Orbigny). X 56; a, side view; b, apertural view; Plesiotype No. 4139, locality LlC, San Sebastian formation. 365 Figure 4. Pullcnia hidloides (d'Orbigny). X 80; a, side view; b, apertural view; Plesiotype No. 4087, locality P255, Ponce formation. 360 Figure 5. Lamarckina echinata n. sp. X 90; a, dorsal view; b, ventral view; c, apertural view; Holotype No. 4111, locality P4, Ponce formation. 368 Figure 6. Gyroidma stellifern n. sp. X 48; a, dorsal view; b, ventral view; c, apertural view; Holotype No. 4105, locality A86, Cibao formation. 378 Figure 7. Gyroidma cf. soldami d'Orbigny. X 53; a, dorsal view; b, ventral view; c, apertural view; Plesiotype No. 4114, locality P3, Ponce formation. 377 Scientific Survey of Porto Rico and the Virgin Islands Volume III, Part 4, Plate IS ■"-JC5^^* 2 a VSI 3 b ,K^ v>Ov 5 a 5 c 3 a / 6 fi.^ y'' ,,^"^ 2c 2q 3 b FyS >»>'' 3a 4 b / V. 4 a GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA Scientific Survey of Porto Rico and the Virgin Islands Volume III, Part 4, Plate 20 I I Q ^^ u 2 b 2 c 2 Q 3 a 3 c 3 b 4 b r \ 4 c 40 GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 469 Plate 20 PAGE Figure 1. Rotalia meyerhoffi n. sp. X 53; a, dorsal view; b, ventral view; c, apertural view; Holotype No. 4108, locality P431, Ponce formation. 381 Figure 2. Rotalia tholtis n. sp. X 60; a, dorsal view; b, ventral view; c, apertural view; Holotype No. 4110, locality P431, Ponce formation. 382 Figure 3. Discorbis havanensis Cushman & Bermudez. X 53; a, dorsal view; b, ventral view; c, apertural view; Plesiotype No. 4112, locality P258, Ponce formation. 383 Figure 4. Discorbis inultisedus n. sp. X 93; a, dorsal view; b, ventral view; c, apertural view; Holotype No. 3898, locality L5C, San Sebastian for- mation. 384 470 SCIENTIFIC SURVEY OF PORTO RICO Plate 21 PAGE Figure 1, DiscorUs ohgospiratus n. sp. X 37; a, dorsal view; 1), ventral view; c, apertural view; Holotype No. 3899, locality A86, Cibao formation. 384 Figure 2. Dtscorbis svbaraucan.us Cushman. X 53; a, dorsal view; b, ven- tral view; c, apertural view; Plesiotype No. 3900, locality L5C, San Sebastian formation. 386 Figure 3. Cydoloculina cubensis Cushman & Bermudez. X 53; adult specimen: a, side view; b, edge view; Plesiotype No. 3888, locality L3C, San Sebastian formation. 389 Figure 4. Cydoloculina cubensis Cushman & Bermudez. X 53; side view of young specimen; Plesiotype No. 3886, locality A79, Los Puertos formation. 389 Figure 5. Discorbis flondanus Cushman. X 53; a, dorsal view; b, ventral view; c, apertural view; Plesiotype No. 3897, locality L5C, San Sebastian for- mation. 383 Figure 6. Discorbis pelliculatus n. sp. X 53; a, dorsal view; b, ventral view; c, apertural view; Holotype No. 4141, locality A21, Quei)radillas formation. 385 Scientific Survey of Porto Rico and the Virgin Islands Volume III, Past 4, Plate 21 2 a i A-x /'' y ■ \ ■J: \ \ I C J 3 a 3 b 5 b ■■ r 6c 6a GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA Scientific Survey of Porto Rico and the Virgin Islands Volume III, Part 4, Plate 22 /'-'■^ V I Ic t: ■ I V 2 b 2 c 2 \ 3 c i \ / 4 b GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 471 Plate 22 PAGE Figure 1. Anomalina alazanensis Nuttall. X 40; a, dorsal view; b, ventral view; c, apertui-al view; Plesiotype No. 3866, locality P3, Ponce formation. 387 Figure 2. Anomalina nudeata (Seguenza). X 40; a, dorsal view; b, ven- tral view; c, apertural view; Plesiotype No. 4041, locality P254, Ponce formation. 388 Figure 3. Anomalina pompihoides n. sp. X 76; a, dorsal view; b, ventral view; c, apertural view; Holotype No. 3867, locality A86, Cibao formation. 389 Figure 4. Cibicides to (Cushman). X 53; a, dorsal view; b, ventral view; c, apertural view; Plesiotype No. 4123, locality A21, Quebradillas formation. 392 Figure 5. Cibicides mexicanus Nuttall. X 27; a, dorsal view; b, ventral view; c, apertural view; Plesiotype No. 3924, locality P3, Ponce formation. 394 472 SCIENTIFIC SURVEY OF PORTO RICO Plate 23 PAGE Figure 1, Cibicides chodawensis Cushman & McGIamery. X 76; a, dorsal view; b, ventral view; c, apertural view; Plesiotype No. 4117, locality A43a, Cibao formation 391 Figure 2. CiUcides fioridanus (Cushman). X 76; a, dorsal view; b, ventral view; c, apertural view; Plesiotype No. 4118, locality A86, Cibao formation. 392 Figure 3. Cibicides scalenus n. sp. X 76; a, dorsal view; b, ventral view; c, apertural view; Holotype No. 4121, locality A43a, Cibao formation. 396 Figure 4. Cibicides perlucidus Nuttall. X 40; a, dorsal view; b, ventral view; c, apertural view; Plesiotype No. 4119, locality P251, Ponce formation. 394 Figure 5. Cibicides pseud(mngcrianus (Cushman). X 76; a, dorsal view; b, ventral view; c, apertural view; Plesiotype No. 4120, locality A79, Los Puertos formation. 395 Scientific Survey of Porto Rico and the Virgin Islands Volume III, Part 4, Plate 23 \' 2 Q ■■■/ 7 2 c i4:: 'I ^ 2 b 4 b 3 a 3 c '^' 3 b 5 a ^ 5 c GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA Scientific Survey of Porto Rico and the Virgin Islands Volume III, Part 4, Plate 24 I ,4 '*""'*'7\ I b /?• "^' ''r^' '^ > 2 I c ^^ \ >. y\ — ; 30 3 c 2 b 4 b ^. -^..^"^ 3 b 5 a v5 \. 4 c 4 Q vV,'«* 5 b V 5 c GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 473 Pl.ATE 24 PAGE Figure 1. CihicideUa variabilis (d'Orbigny). X 25; a, dorsal view; b, ven- tral view; c, apertural view; Plesiotype No. 4127, locality L4C, San Sebastian formation. 401 Figure 2. E pistomaria pontifera n. sp. X 40; a, dorsal view; b, ventral view; c, apertural view; Holotype No. 4115, locality P258, Ponce formation. 386 Figure 3. Cibicides amcricanun (Cushman). X 76; a, dorsal view; b, ven- tral view; c, apertural view; Plesiotype No. 3952, locality LlC, San Sebastian formation. 390 Figure 4. Cilindes lobaius (d'Orbigny). X 53; a, dorsal view; b, ventral view; c, apertural view; Plesiotype No. 3884, locality L4C, San Sebastian for- mation. 393 Figure 5. Cibicides sinistralis Coryell & Rivero. X 76; a, dorsal view; b, ventral view; c, apertural view; Plesiotype No. 4101, locality A86, Cibao for- mation. 396 474 SCIENTIFIC SURVEY OF PORTO RICO Plate 25 PAGE Figure 1. Cibicides s]nrolimbaius n. sp. X 40; a, dorsal view; b, ventral view; c, apertural view; Holotype No. 4122, locality A93, Quebradillas formation. 397 Figure 2. Planulina crassa n. sp. X 76; a, dorsal view; b, ventral view; c, apertural view; Holotype No. 4065, locality A21, Quebradillas formation. 398 Figure 3. Planulina depressa (d'Orbigny). X 27; a, dorsal view; b, ven- tral view; c, apertural view; Plesiotype No. 4124, locality P255, Ponce formation. 398 Figure 4. Planulina ■manalana Hadley. X 27; a, dorsal view; b, ventral view; c, apertui-al view; Plesiotype No. 4125, locality P2, Ponce formation. 399 Scientific Survey of Porto Rico and the Virgin Islands Volume III, Part 4, Plate 25 ■J' f ""^ I C I b y 3 b r 4 b ^ \ 4 c X 4 GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA Scientific Survey of Porto Rico and the Virgin Islands ./IJK I Volume III, Part 4, Plate 26 X" \ ^ "^ # 4a f\ i W.5- ..'■.;■- •^ ,>— 2 Q s. 'i ^w 3 b 3 c ) i>. / .{A*, U'^A*- ^ A 5 \., __: 5c -A 6a 9^ 6 b V ^ 6c GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA '^11 Plate 28 PAGE Figure 1. ChiloatomeUa ovoidea Reuss. X 53; a, side view; b, back view; c, apertui-al view; Plesiotype No. 4135, locality P 255, Ponce formation. 409 Figure 2. ChilostomeUa ghbata n. sp. X 76; a, side view; b, back view; c, apertural view; Holotype No. 4134, locality P254, Ponce formation. 409 Figure 3. Chilostomella czizeki Reuss. X 53; a, side view; b, back view; c, apertuial view; Plesiotype No. 4133, locality P254, Ponce formation. 409 Figure 4. Grj/mna discus (Goes). X 15; Plesiotype No. 4129, locality A21, Quebradillas formation. 405 Figure 5. Amphistegma floridana Cushman. X 27; a, dorsal view; b, ven- tral view; c, apertural view; Plesiotype No. 3863, locality L4C, San Sebastian formation. 408 Figure 6. Amphistegina angulata (Cushman). X 40; a, dorsal view; b, ven- tral view; c, apertural view; Plesiotype No. 3862, locality F359, Quebradillas formation. 407 478 SCIENTIFIC SURVEY OF PORTO RICO Plate 29 PACK FiGVRE 1. Globigcrhia huUoidcs d'Orbignj'. X 77; a, dorsal view; b, ventral view; Plesiotype No. 3920, locality LlC, San Sebastian formation. 411 Figure 2. Ghbigerhia dutertrei d'Orbigny. X 77; a, dorsal view; b, ven- tral view; Plesiotype No. 3921, locality LlC, San Sebastian formation. 411 Figure 3. Ghbigcrlna inflata d'Orbigny. X 53; a, dorsal view; b, ventral view; c, apertural view; Plesiotype No. 4137, locality P251, Ponce formation. 412 Figure 4. Globigmna ouachitainsis Howe & ^^'allace. X 76; a, dorsal view; b, ventral view; c, apertiual view; Plesiotype No. 4048, locality A21, Quebra- dillas formation. ^^^ Figure 5. Globigcrinn pachydcrma (Elu-enberg) . X 90; a, dorsal view; b, ventral view; c, apertural view; Plesiotype No. 4138, locality P254, Ponce for- mation. ^'-'^ Figure 6. Globigcrina pseudotriloba White. X 77; a, dorsal view; b, ven- tral view; Plesiotype No. 3922, locality LlC, San Sebastian formation. 413 Figure 7. Globigmna IrtlocularU d'Orbigny. X 77; young individual: a, dorsal view; b, ventral view; Plesiotype No. 3923, locality L5C, San Sebastian formation. ^^'^ Scientific Survey of Porto Rico and the Virgin Islands Volume III, Part 4, Plate 29 ■K. 2a 2 b '■■■'d 3 b 3 Q ■\ 7^' 4 b 4 C 4 i 5 b ^'''■"x 6 b 6a 5 c ' 27 5 a 7 b 70 GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA Scientific Survey of Porto Rico and the Virgin Islands Volume III, Part 4, Plate 30 / i I b 2 a 2 b 3 I 4 a 6 Q 8 b 9 b 4 b /^^. 8 Q 9 Q b 5 b i 1 5 a II b X \ \--v^' iOa — II a GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 479 Plate 30 PAGE Figure 1. Sphaeroidina bulloides d'Orbigny. X 53; a, dorsal view; b, ven- tral view; Plesiotype No. 3991, locality P254, Ponce formation. 410 Figure 2. Pavonina advma Cushman. X 110; a, side view; b, edge view; Plesiotype No. 4136, locality A6, Lares formation. 415 Figure 3. OrluUna universa d'Orbigny. X 110; Plesiotype No. 3942, lo- cality L6C, San Sebastian formation. 414 Figure 4. SphaeroidineUa seviinuUna (Schwager). X 53; a, ventral view; b, dorsal view; Plesiotype No. 3992, locality A21, Quebradillas formation. 415 Figure 5. Bolivina elongaia Hantken. X 57; a, side view; b, apertural view; Plesiotype No. 3875, locality P255, Ponce formation. 416 Figure 6. Bolivina heineae n. sp. X 90; a, side view; b, apertural view; Holotype No. 3874a, locality P2, Ponce formation. 417 Figure 7. Bolivina heineae, n. sp. X 90; Specimen with surface removed with acid to show chambers and sutures; Paratype No. 3874b, locality P2, Ponce formation. 417 Figure 8. Bohvina jacksonensis Cushman & Applin. X 110; a, side view; b, apertural view; Plesiotype No. 3876, locality LlC, San Sebastian formation. 418 Figure 9. Bohvina mantanzana co7we.ra n. sp. X 76; a, side view; b, aper- tural view; Holotype No. 4038, locality A43a, Cibao formation. 418 Figure 10. Bohvina 7nantanzana convexa n. sp. X 76; a, side view; b, aper- tm"al view; Paratype No. 4039, locality LlC, San Sebastian formation. 418 Figure 11. Bolivina byramcnsis Cushman. X 76; a, side view; b, apertural view; Plesiotype No. 3872, locality A86, Cibao formation. 416 480 SCIENTIFIC SURVEY OF PORTO RICO Plate 31 PAGE Figure 1. Bolivina mexicana nliformis Cushman. X 37; a, side view; b, apertural view; Plesiotype No. 4040, locality P3, Ponce formation. 419 Figure 2. Bolivina tediformis Cushman. X 93; a, side view; b, apertural view; Plesiotype No. 3878, locality P2, Ponce formation. 419 Figure 3. Loxostomum hiwanneense Howe. X 76; a, side view; b, apertural view; Plesiotype No. 3929, locality A43a, Cibao formation. 420 Figure 4. Loxosiomum normale n. sp. X 76; a, side view; b, apertural view; Holotype No. 3930, locality A43a, Cibao formation. 421 Figure 5. Bolivina ventricosa n. sp. X 93; a, side view; b, apertural view; c, edge view; Holotype No. 3879, locality P254, Ponce formation. 420 Figure 6. Plcctofrondicularia trinitaten.-ii..s Cushman & Jarvis. X 53; a, side view; b, end view; Plesiotype No. 4068, locality P3, Ponce formation. 421 Figure 7. Plectojrondicularia vaughani Cushman. X 53; a, side view; b, apertural view; Plesiotype No. 3949, locality P2, Ponce formation. 422 Figure 8. Reusaella glabrata (Cushman). X 100; a, edge view; b, apertural view; Plesiotype No. 4073, locality A79, Los Puertos formation. 423 Figure 9. Buliminn socialis Bornemann. X 76; a, side view; b, apertural view; Plesiotype No. 3880, locality P254, Ponce formation. 423 Scientific Survey of Porto Rico and the Virgin Islands 2 b Volume III, Part 4, Plate 31 3 b 4 b 3 a \/ 4 a 5 b ■ I 5 a \. .^ 5 c 6 b ti "|l; I ^ 6q . /. 8 b y 9 a GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA Scientific Survey of Porto Rico and the Virgin Islands / i / Volume III, Part 4, Plate 32 2 b 3 b ■V^ 3 c 3 a 4 a 5 a /^~ ■-■ 4 d 4 c 4 b ri C^^\ <£>-' 5 d / yr 5 c '*r' 5 b GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 481 Plate 32 PAGE Figure 1. Cassidulma laevigata d'Orbigny. X 110; a, side view; b, aper- tm-al view; Plesiotype No. 4142, locality A43a, Cibao formation. 424 Figure 2. Cassidulma subglobosa Brady. X 76; a, apertiiral view; b, back view; Plesiotype No. 4180, locality P2, Ponce formation. 425 Figure 3. Cassididma tricamerala n. sp. X 53; a, side view; b, opposite side; c, apertuial view; Holotype No. 4143, locality P255, Ponce formation. 425 Figure 4. Ehrtnhergina caribbea n. sp. X 53; a, dorsal view; b, ventral view; c, edge view; d, apertural view; Holotype No. 4140, locality P2, Ponce for- mation. ^^" Figure 5. Ehrcnbirgina serrata gibbera n. var. X 53; a, dorsal view; b, ven- tral view; c, edge view; d, apertural view; Holotype No. 3902, locality P2, Ponce formation. *^' 482 SCIENTIFIC SURVEY OF PORTO RICO Plate 33 PAGE Figure 1. Uvigerina bulbacea n. sp. X 76; Holotype No. 4144, locality A86, Cibao formation. "^27 Figure 2. Uvigerina bulbacea n. sp. X 76; Paratype No. 4144a, locality A86, Cibao formation. ^^27 Figure 3. Uvigerina postica n. sp. X 53; Holotype No. 4146a, locality A86, Cibao formation. "^'^^ Figure 4. Uvigerina -jiostica n. sp. X 53; Paratype No. 4146b, locality A86, Cibao formation. "^"^^ Figure 5. Uvigerina elongata Cole. X 76; Plesiotype No. 4017, locality P254, Ponce formation. "*28 Figure 6. Uvigerina f us if ormis n. sp. X 76; Holotype No. 4018, locality P2, Ponce formation. "^^^ Figure 7. Uvigerina ^nantaensis Cushman & Edwards. X 53; Plesiotype No. 4076, locality P3, Ponce formation. ^ -^SO Figure 8. Uvigerina gallowayi Cushman. X 53; Plesiotype No. 4145, lo- cality A86, Cibao formation, 429 Figure 9. Uvigerina mexicana Nuttall. X 53; Plesiotype No. 4021, locali- ty P2, Ponce formation. ^ "^30 Figure 10. Uvigerina mexicana biilbosa n. var. X 53; Holotype No. 4077, locality P3, Ponce formation. '*31 Figure 11. Uvigerina mexicana ranunculus n. var. X 40; Holotype No. 4022, locality P2, Ponce formation. -^31 Figure 12. Uvigerina vicksburgensis Cushman & Ellisor. X 53; a, side view; b, apertural view; Plesiotype No. 4020, locality P251, Ponce formation. 432 Figure 13. Uvigerina gardnerae Cushman. X 40; a, side view; b, apertnial view; Plesiotype No. 4019, locality P254, Ponce formation. 429 Scientific Survey of Porto Rico and the Virgin Islands Volume III, Part 4, Plate 33 8 r\ \^ \ - ( ■ \ . f • 10 A I I y-T» 12 a 12 b / ,VH^. 13 b 'Kfo^ua^ 11 13 Q GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA Scientific Survey of Porto Rico and the Virgin Islands Volume III, Part 4, Plate 34 I b iV 1-^ ^ J f 5 b \s^ 5a ■ » ' v> \ ^ II C II b \ .1 8 FN*r -f ^yr 12 b \Z a If 13 b 1 13a GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 483 Plate 34 PAGE Figure 1. Siphogenerina mexicana Cushman. X 53; a, side view; b, aper- tural view; Plesiotype No. 3986, locality P2, Ponce formation. 434 Figure 2. Siphogenerina hubbardi n. sp. X 40; Holotype No. 3985, locality P2, Ponce formation. '*34 Figure 3. Siphogenerina multicostaia Cushman & Jarvis. X 53; Plesiotype No. 3987a, locality P254, Ponce formation. 435 Figure 4. Siphogenerina multicostata Cushman & Jarvis. X 53; Plesiotype No. 3987b, locality P254, Ponce formation. 435 Figure 5. Siphogenerina eostostriata n. sp. X 53; a, side view; b, apertm-al view; Holotype No. 4147, locality P254, Ponce formation. 432 Figure 6. Siphogenerina cumingsi n. sp. X 53; Holotype No. 3983, locali- ty P3, Ponce formation. ^^^ Figure 7. Siphogenerina cumingsi n. sp. X 53; Paratype No. 3984, locali- ty P2, Ponce formation. "^33 Figure 8. Angulogerina decorissima n. sp. X 53; Holotype No. 3865a, lo- cality A21, Quebradillas formation. -136 Figure 9. Angidogerina decorissima n. sp. X 53; Paratype No. 3865b, lo- cality A21, Quebradillas formation. _ -136 Figure 10. Angidogerina decorissima n. sp. X 53; Paratype No. 3865c, locality A21, Quebradillas formation. ^36 Figure 11. Angulogerina ponceana n. sp. X 53; a, side view; b, back view; c, apertmal view; Holotype No. 4023, locality P254, Ponce formation. 437 Figure 12. Angulogerina cibaoensis n. sp. X 53; a, side view; b, apertural view; Holotype No. 3864, locality A86, Cibao formation. 435 Figure 13. Angulogerina cooperensis Cushman. X 53; a, side view; b, apertural view; Plesiotype No. 4148, locality A86, Cibao formation. 436 484 SCIENTIFIC SURVEY OF PORTO RICO Plate 35 PAGE Figure 1. PJeuroHtomeUa gerontica n. sp. X 48; a, front view; b, side view; c, apertural view; d, back view; Holotype No. 4071, locality P2, Ponce formation. 439 Figure 2. PleurodomcUa bierigi Palmer & Bermudez. X 53; a, front view; b, side view; c, apertural view; Plesiotype No. 4069, locality P3, Ponce formation. 438 Figure 3. Pleurodomella elHpiica n. sp. X 53; a, front view; b, side view; Holotype No. 4070, locality P2, Ponce formation. 438 Figure 4. Trifarina bradyi Cushman. X 76; a, edge view; b, side view; Plesiotype No. 4005, locality P3, Ponce formation. 437 Figure 5. NodosareUa consirida granuHfera n. var. X 40; a, side view; b, apertural view; Holotype No. 4059, locality P2, Ponce formation. 439 Figure 6. NodosareUa crassielegans (Nuttall). X 20; a, side view; b, aper- tural view; Plesiotype No. 4078, locality P3, Ponce formation. 439 Figure 7. Nodomrella paucistriala Galloway & Morrey. X 20; a, side view; b, apertural view; Plesiotype No. 4058a, locality P3, Ponce formation. 440 Figure 8. NodosareUa paucislriata Galloway & Morrey. X 20; a, side view; b, apertural view; Plesiotype No. 4058b, locality P3, Ponce formation. 440 Figure 9. Nodosareda paucistriala Galloway & Morrey. X 27; a, side view; b. apertural view; Plesiotype No. 3891, locality P2, Ponce formation. 440 Figure 10. NodosareUa verneuili (d'Orbigny). X 27; a, side view; b, aper- tural view; Plesiotype No. 4034, locality A86, Cibao formation. 440 Scientific Survey of Porto Rico and the Virgin Islands Volume III, Part 4, Plate 35 /m>^ :^ I 'A 10 3 a 2 b % 6 b :frs i I 2 c 7b 2 -^4 8 b 9 b r .V'*^ 4 b 10 b V~f 8 a 9 a 10 GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA Scientific Survey of Porto Rico and the Virgin Islands Volume III, Part 4, Plate 36 I ^ro 4 Q 4 C 4 b GALLOWAY AND HEMINWAY: TERTIARY FORAMINIFERA GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 485 Plate 36 PAGE Figure 1. Elhj)sogla7iduKna exponens (Brady). X 27; a, side view; b, aper- tural view; Plesiotype No. 3903, locality P2, Ponce formation. 441 Figure 2. Ellipsoidina ellipsoides abbreviata Seguenza. X 20; a, side view; b, apertural view; Plesiotype No. 3904a, locality P2, Ponce formation. 441 Figure 3. EUipsoidina ellipsoides abbreviata Seguenza. X 20; apertural view of another specimen; Plesiotype No. 3904b, locality P2, Ponce formation. 441 Figure 4. Operculinella sinuata n. sp. X 53; a, side view; b, opposite side; c, edge view; Holotype No. 4067, locality A21, Quebradillas formation. 442 Figure 5. Heferostegina antillea Cushman. X 13; Plesiotype No. 4149, lo- cality L3C, San Sebastian formation. 442 Figures 6 and 7. Miogypsinotdes complanata (Schlumberger). X 20. 444 Figure 6. Side view of a small specimen, 1 millimeter long, Plesiotype No. 4150a, locality L4C, San Sebastian formation. 444 Figure 7. Side view of a medium sized specimen: breadth 1.83 millimeters, length 1.5 miUimeters; Plesiotype No. 4150b, locality L4C, San Sebastian for- mation. ■*■*■* Figures 8 to 10. Miogypsinotdes complanata (Schlumberger). X 30. 444 Figure 8. Longitudinal vertical section through proloculum (P) showing median chamber, fibrous lateral laminae and chamber walls and pillars; Plesio- type No. 4150c, locality L4C, San Sebastian formation. 444 Figure 9. Median section of a small specimen (0.9 millimeter diameter) showing the proloculum, coiled nucleoconch and later spatulate chambers; Plesiotype No. 4150d, locality L4C, San Sebastian formation. 444 Figure 10. Median section of a medium sized specimen (1.33 millimeters) showing the proloculum, coiled nucleoconch, and later spatulate chambers; Plesiotype No. 4151, locality L6C, San Sebastian formation. 444 GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 487 INDEX=' Page Amphicoryne falx 338 obliqua 278, 294, 338 Amphistegina angulata. .281, 292, 296, 407 chipolensis 408 floridana 281, 288, 289, 292, 296, 408 Angulogerina albatrossi ornata 436 cibaoensis 282, 289, 435 cooperensis 282, 289, 435, 436 decorissima 282, 292, 436 naranjoensis 437 ponceana 282, 297, 437 Anomalina alazanensis 280, 295, 387 grosserugosa 389 nolani 388 nucleata 280, 292, 295, 388 pompilioides 280, 289, 389 subtenuissima 377 umbonata 388 Antillesina 366 marielensis 279, 286, 287, 295, 367 Apiopterina cylindroides 279, 295, 355 Archaias aduncus, 277, 287, 290, 291, 293, 317 angulatus 277, 287, 290, 291, 293, 317, 318 compressus 277, 287, 318 Aristerospira pachy derma 413 Astacolus crepidula 334, 335 insolitus 278, 294, 334 ovatus 278, 289, 334 sublituus 278, 289, 294, 335 Asterigerina angulata 407 choctawensis 408 Bigenerina capreolus 333 pennatula 333 Biloculina bougainvillei 310 clypeata 311 ' globulus 313 inornata 311 lunula 311 oblonga 312 peruviana 312 subspherica 313 Bolivina byramensis 281, 289, 416, 417 caelata byramensis 416 elongata 281,296,416 heineae 281, 296, 417 jacksonensis 281, 288, 418 matanzana 418 matanzana convexa. . .281, 288, 289, 418 me.\icana aliformis 281, 296, 419 reticulata 417 tectiformis 281, 296, 419 ventricosa 281, 296, 420 Page Bulimina ovata 423, 424 sociaUs 281, 296, 423 Cancris sagra 370 Carpenteria alternata 404 balaniformis proteiformis 404 bulloides 281, 292, 404 lithothamnica 404 proteiformis 281, 292, 296, 404 utricularis 405 Cassidulina delicata 425 delicatula 426 laevigata 281,289,290,424 rarilocula 426 subglobosa 281, 296, 425 tricamerata 281, 296, 425 Ceriopora globulus 406 Cliilostomella cylindroides 409 czizeki 281, 296, 409 globata 281, 296, 409 grandis 410 ovoidea 281,296,409,410 urceolus 281, 296, 410 Cibicidella variabilis 280, 287, 401 Cibicides americanus .... 280, 287, 289, 390 choctawensis 280, 289, 391 floridanus 280, 289, 292, 295, 392, 393, 395, 397 io 280, 292, 392 lobatula 393 lobatulus 393, 394 lobatus 280, 287, 289, 290, 292, 295, 393 mexicanus 280, 295, 394 niundula 397 nucleata 388 perlucida 394 perlucidus 280, 295, 394, 397 pseudoungeriana io 392 pseudoungerianus . 280, 290, 392, 395, 397 pseudoungerianus evolutus 397 scalenus 280, 289, 396 sinistralis 280, 289, 396 spirolimbatus 280, 292, 397 trinitatensis 388 westi 396 Clavulina Ijyramensis 325 communis 326 cylindrica 325 triangularis 327 tricarinata 278, 284, 287, 290, 291, 293, 326 Clavulinoides guayabalensis 327 jarvisi 327 polygonalis 278, 291, 327 triangularis 278, 293, 327 * New genera, species, and varieties, and pages where treated, are in bold faced type. 488 SCIENTIFIC SURVEY OF PORTO RICO Page Cristellaria acutauricularis 333, 334 arcuata 333 articulata 348 calHfera 347 convergens 348 crassa 351 crepidula 334 falcifer 349 insolita 334 iota 349 karreri 351 occidentalis torridus 349 protuberans 351 subalata 349 sublituus 335 subpapillosa 351 Cubanina alavensis 326 Cyclamniina acutidorsata . . . .278, 293, 319 Cycloloculina cubensis 280, 287, 290, 295, 389 Dendritina antillarum 316 preelegans 277, 290, 315 Dentalina advena 278, 294, 339 communis 339, 340 debilis 340 lialkyardi 278, 294, 340, 341 halkyardi ponceana 278, 294, 340 hillaeformis 278, 294, 341 multilincata 278,294, 341 semilaevis 279, 294, 342 verneuili 440 Discorbina globularis 384 polystomelloides 386 tuberculata 369 Discorbis floridanus 280, 287, 290, 383 havancnsis 280, 290, 292, 295, 383 mirus 384 multisectus 280, 287, 384 oligospiratus 280, 289, 384 orbicularis 384, 3S5 pelliculatus 280, 292. 385 subaraucanus 280, 287, 290, 295, 386 Dorothia caribaea 278, 293, 322 cylindrica 278, 286, 293. 322 praelonga 278, 293, 322 E;iH-enl)ergina bradyi 426 caribbea 281, 296, 426 navalis 426 serrata 427 serrata gibbera 281, 296. 427 Ellipsoglandulina exponens. . .282, 297, 441 lae\'lgata 441 principiensLs 441 Ellipsoidina al)brev-iata 441 ellipsoides 441 ellipsoides ahbreviata 282, 297, 441 exponens 44 1 Ellipsonodosaria verneuili 440 verneuili paucistriata 440 Elpliidium australe 363 Page chapmani 361 chipolense 366 craticulatum 361 crispum 363 incertum 363 lanicri 279, 295, 360, 361 lens 279, 287, 361 lobatum 279, 288. 291, 362 nautiloideum 279, 287, 291, 362 owenianum. . .279, 287, 289. 290. 295. 363 pocyanum 279. 287, 288. 290. 291, 295, 362, 363 puertoricense 279, 287, 290. 291. 295, 364. 366 rugosum 366 sagrai 279, 287, 289, 290, 291, 295, 361. 365, 366 sagrai crassum 279, 287, 290, 292, 295, 365, 366 Epistomaria pontifera 280, 295,386 rimosa 387 Eponides advena 280, 289, 371. 375 antillarum 374 byramensis cubensis . 374 ellisorae 280. 295. 372 exiguus 280. 287. 373 guayabalensis 374 jacksonensis 375 me.xicana 374 nana 372 ornatissimus . 280, 287, 289, 290, 295, 373 parantillarum 280, 292, 295. 374 pulvinus 280. 289. 292. 375 repandus 280. 292. 375 ventricosus 280, 289. 292. 295, 376 vortex 280, 289, 376 I'Mssurina laevis 279. 295. 353 marginata 279, 295, 353 (iaudryina asiphonia 278, 293, 323 cylindrica 322 glabrata 324 glabrata maxima 278, 289, 323 jacksonensis 325 karreriana 278, 291, 293, 324 praelonga 322 (Pseudogaudryina) atlantic^ . . . 278, 293, 324,325 puertoricana 278, 287, 324 rugulo.sa 331 siphoncUa asiphonia 323 Glandulina comata 338 comatula 278, 294, 338 gallovvayi 278, 289, 294, 338 mauriconsis 278, 294, 339 Globigerina bulloides. .281, 288, 289, 290. 292, 296, 411 dubia 412 dutertrei 281, 288,289.292, 296,411 inflata 281.296,412 ouachitaensis 281, 289, 292, 296, 412 GALLOWAY AND HEMINWAY, TERTIARY FORAMINIFERA 489 Page pachydernia 281, 296, 413 pseudotriloba 281, 288, 280, 296, 413 seniinulina 415 triloba 413 trilocularis . . . 281, 288, 289, 292, 296, 414 triloculinoides 414 Globorotalia menarclii 280, 292, 368 palniarealensis 370 Globulina gibba slobosa 355 Guttulina basalis 279, 291, 354 kisMnouyi 354 Gypsina discus 281, 292, 296, 405 globulus 406 globulus pilaris 406 vesicularis discus 405 Gvroidina c/. soldanii 280, 295. .377 steUifera 280, 289, 378 Haplophragmium acutidorsatuni 319 Hauerina bradyi 315 compressa 315 sansebastianensis 277, 287. 315 Hemicristellaria fragaria 278, 291, 335 Heterostegina antillea . . . 282, 285, 288, 442 ocalana 443 Karreriella mexicana 321 Lagena amphora 345 ampulla 279, 287, 345 buUosa 279, 294. 345 curvilineata 346 impressa 279, 294, 346 nuttalli 279, 294, 346 raricosta 345 striata strumosa 346 strumosa 279, 294, 339, 346 sulcata 346 tenuis 345 Lamarckina echinata 280, 295, 368 Lenticulina convergens 348 Lepidocyclina cf. parvula . . . 282, 285, 288, 297, 443 (Nephrolepidina) tournoueri. . .282, 288, 297, 443 Liebusella byramensis 278, 293, 325 Lingullna cubensis 352 ponceana 279, 295. 352 semicostata 279. 295. 353 Listerella cf. communis 278, 293, 326 Loxostomum liiwanneense . . . .281, 289, 420 normale 281, 289, 421 rostrum 421 ]Marginulina eximJa 337 insulensis 278, 294, 337 subcrassa 278, 294. 337 Massilina decorata 277, 290, 314 inaequalis 277, 290, 314 jacksonensis 303 IMiliolina pygmaea 304 Miliolinella circularis . . . .277, 290, 293, 310 Miogypsina complanata 444 Miogypsinoides complanata 282, 285, 286, 288, 289. 290, 297, 444 Page Nautilus aduncus 317 angulatus 318 pompilioides 357 raplianistrum 344 repandus 375 (Orthocuras) obliquatus 343 pennatula 333 scalaris 344 Nodosarella constricta 439 constricta granulifera 282, 297, 439 crassielegans 282, 297, 439 paucistriata 282, 289. 297. 440 verneuili 282. 289, 440 Nodosaria ad vena 339 catenulata 440 comatula 338 crassielegans 439 halkyarcU antillana 279, 289, 342 intermittens 440 longiscata 279, 294, 342, 343 obliquata 279, 294, 343 pariana 279, 294, 343 pauperata 340 raphanistrum 279, 294, 344 scalaris 279, 294, 344 simplex 279, 294, 344 Xonion chapapotense 279, 287, 355 dUatatum 279. 289, 356 elongatum 358 grateloupi 358 marielensis 366. 367 multiporatum 279, 289, 356 nicobarense 279, 289, 291, 295, 357 pompilioides 279, 289, 295, 357 subgrateloupi 279, 287, 290, 295. 358 umbilicatulus 357 Nonionella auris 359 dauvillon.sis 360 modesta 279. 287, 359 pauciloba 359 pseudoauris 359 Nonionina bulloides 260 umbilicatula 357 Operculinella dia 442 sinuata 282,292.442 Orbiculina adunca 316, 317, 318 compressa 318 Orbulina universa 281, 288, 292, 414 Ovulina tenuis 345 Patellina advena 300 corrugata 277,293,299 Pavonina advena 281, 288, 415 flabelliformis 416 Peneroplis carinatus.277, 287, 291, 293, 316 dubia ; • ■ 316 elegans 316 pertusus carinatus 316 prote; 316 proteus 277,287,290,291,316 Planorbulina vulgaris larvata 403 Planorbulinella larvata. .280, 289, 292, 403 490 SCIENTIFIC SURVEY OF PORTO RICO Page Planulina alavensis 401 arirninensis 398 camagueyana 374 cofoaonsis cooperensis 400 crassa 280, 292, 398 dcpressa 280, 295, 398 marialana 280, 295, 399 niexicana 280, 295, 399 wut'llt'istorfi 399 zigzag 280, 289, 400 Plecaniuni ru^osum 331 Plectofrondicularia advena 422 trinitatensis 281, 296, 421 vaughani 281, 296, 422 Pleurostomella alternans 438 hU'vi'Ai 282, 297, 438, 439 elliptica 282, 297, 438 gerontica 282, 297, 439 jacksoin'nsis 438 Polyniorpliina amygdaloides terquemiana 354 cylindroides 355 globosa 356 torquciriana 279, 290, 354 Polystonu'lla craticulata 361 lanieri 360 oliscura 367 oweniana 363 poeyana 363 sagra 365 Pseudarcplla arenata 298 patella 277, 288, 298 Pseudoglandulina gallowayi 338 niauriccnsis 339 Pseudononion papillatum 279, 289, 358 Pullcnia Imlloides 277, 295, 3(50 sphaeroides 360 Pulvinulina l>yrainensis 372 exigua 373 monardii 368 repanda 375 Pyrgo l)ougainvilIci 277, 287, 310, 312 bulloidcs 313 clypcata 277, 287, 288, 289, 290, 291, 311 dt'pri'ssa 312 clongata 312 inornata 277, 287, 290, 291, 31 1 liimila 277, 287. 290, 291, 293, 31 1 oblonga 277, 287, 288, 289, 290, 291, 293. 312 peruviana. . . .277, 287, 289, 290, 293, 312 sphaora 314 subsphacrica 277, 287, 288. 290, 291, 293, 313 Pyrgoella globulus 277, 293, 313 Pyrulina cyliiKb-oidcs 355 Quinriuoloculina aknoriana. .277, 287, 289, 301 anguina 304 angusta 304 Page august is.sima 304 apiculata 277, 287, 302 audacula 277, 290, 302 badcnsis 302 jamaicensis 303 lat'vigata 277, 287. 302 maculata 277, 287, 303 phiiippi 277, 290. 303 ponceana 277. 293. 304 pyginaca 277, 287, 290, 291, 304 seminuluiii . . .277, 287, 289, 301. 304. 305 vulgaris 277. 287. 290. 291. 293. 305 Raphanulina gibba globosa. .279. 287. 290. 291,355 Reussi'lla glabrata . 281, 288. 290, 292, 296. 423 Reu.ssia spinulosa 423 Rhabdammina abyssorum 299 irregularis 277, 293, 299 Rhabdogonium tricarinatum 437 Robulus arcuatostriatus carolinianus . .347 brevispinosus 352 calliferus 279. 294, 347 chanibersi 279, 289, 291. 294. 347 cibaoensis 279. 289, 348 clericii 347. 348 convergens 279, 289, 295. 348 falcifer 279, 291, 349 gutticostatus 352 iota 279, 295, 349 occidentalis torridus 279, 295, 34!» planulus 279, 289, 350 plunuiicrae 279. 295. 3.")0 protuberans 279. 295. 351 sul)pai)illosu.s 279, 295. 351 Rotalia aculeata 382 advena 371. 372 alabaniensis 382 byraniensis 280. 295. 378 ohocfawensis 280, 285, 287, 379, 381 mcnar^iii 368 mexicana 280, 295. 379. 380. 381 niexicana mecatepecensis. 280. 28(). 295, 3 SO meyerhoffi 280, 295. 381 tholus 280, 295. 382 Rupertia stabilis 10 I verrucosa 281, 295. 403 Saraeenaria areuata 278. 294, 333 Serpula (Lagena) marginata 353 seniinulum 305 Sigmomorpliina seniitecta terquemiana 354 Siphogenerina costostriata . . . 2S2. 297. 432 cumingsi 2S2, 297. 433 hubbardi 282, 297, 434 lamellata 434 niexicana 282, 286, 297, 434 inulticostata 282, 297, 434, 435 Sii)hoiiina advena. .280, 288, 289, 290, 292, 295, 402 GALLOWAY AND HEMINWAV, TERTIARY FORAMINIFERA 491 Page tenuicarinata 280, 295, 402 Sphaerogypsina globulus 281, 288, 289, 292, 29G, 406 pUaris 281, 292, 296. 406 Sphaeroidina bulloides. . .281, 292, 296, 410 Sphaeroidinella deliiscens 415 .semlnulina 281, 292, 415 Spirillina \i\-ipara 277, 289, 293, 299 Spiroloculina antillarum 277, 290. 291, 293, 300 elongata 277, 290, 300 (ixcavata 301 oculina 277, 290, 301 Textularia agglutinans. . .278, 287, 289, 328 articulata 278, 287, 329 atlantica 325 liarrettii 332 broussardi 278, 294, 329 candeana 278, 287. 294. 329 candeiana 329 gramen 278, 287, 330 firenadana 278, 294, 330 indenta 278, 289, 331 pseudotrochu.s 332 rugosa 331 Textulariella barrettii 278, 289, 291. 293, 332 Tinoporus pilaris 406 vesicularis 405 Trifarina bradyi 282, 297, 437 Triloculina austriaca 277, 287, 288, 291, 293. 306 brongniartiana 277, 293. 306 cervicula 277. 287. 306 circularis 310 elliptica 277, 287. 307 gibba 277, 287, 290, 291. 293, 307 gracilis 307 inflata 277,287,290,291, 293,307 lae^^gata 277, 287, 290, 293, 308 , longissima 277, 287, 308 Oblonga.277, 287, 288. 290. 291, 293. 307. 308. 309 quadrilateralis 277. 287. 290, 309 tricarinata 277. 293, 309 Trochamminoides approximatus . 277. 290. 319 proteus 319 Truncatulina araericana 390 Page depressa 398 fioridana 392 lobata 393 lobatula 393 lobatula ornata 392 nucleata 388 trinitatcmsis 388 variabiles 401 Uvigerina bulbacea 282, 289, 427. 432 capayana 430 elongata 282, 296, 428 fusiformis 282. 296, 428 jiallowayi 282. 289. 429 gardnerae 282, 296, 429 gardnerae cubana 430 mantaensis 282, 296, 430 mexicana 282, 296. 430 mexicana bulbosa 282. 297. 431 mexicana ranunculus 282, 297. 431 peri'grina 430 postica 282, 289, 428, 431 proboscidea 430 pygniaea 429 striata 432 vicksburgensis 282, 286, 297. 432 Vaginulina elegans 337 elegans mexicana 336 faba 278, 294. 335 legumen elegans 336 mexicana 278, 286, 294, 336 siliquoidea 278, 294, 336 Valvulaniinina afflnis 321 cornucopia 278, 287, 293. 320 deforiuLs 321 Valvulina oviedoiana. . . .278, 287. 293, 320 Valvulineria cubana 371 inaecjualis 370 maclureaformis 280, 289. 369 nuttalli 280, 295. 370. 371 palraarealensis 280. 286. 295. 370 paucilocula 280. 289, 290, 295. 371 Yermiculum marginatum 353 oblongum 309 Verneuilina mexicana. .278. 286, 291, 293, 321 spinulosa glabrata 423 Vulvulina pachyheilus . .278, 289. 291. 294, 332 pennatula 278, 294. 333 THE NEW YORK ACADEMY OF SCIENCES SCIENTIFIC SURVEY OF PORTO RICO and the VIRGIN ISLANDS VOLUME III NEW YORK Published by the Academy 1941 CONTENTS OF VOLUME III Page Title-page i Contents "i Dates of Publication of Parts and Editors iii Part 1. Tertiary Mollusca from Porto Rico. By C. J. Maury 1 Part 2. Tertiary Mollusca from the Lares District, Porto Rico. By Bela Hubbard 79 Part 3. Fossil Corals of Porto Rico, with Descriptions also of a Few Recent Species, By H. N. Coryell and Violet Ohlsen 167 Part 4. The Tertiary Foraminifera of Porto Rico. By J. J. Galloway and Caroline E. Heminway 275 Dates of Publication of Parts and Editors Part 1, 1920 Ralph Winfred Tower Part 2, 1920 Ralph Winfred Tower Part 3, 1929 Herbert F. Schwarz ' Part 4, April 21, 1941 Erich Maren Schlaikjer (iii) PARTS ISSUED Volume I (complete in four parts, with index). Part 1. History of the Survey, by N. L. Britton. Geological Introduction, by C. P. Berkey. Geology of the San Juan District, with colored map, by D. R. Semmes. Part 2. Geology of the Coamo-Guayama District, with colored map, by E. T. Hodge. Part 3. Geology of the Ponce District, with colored map, by G. J. Mitchell. Part 4. The Physiography of Porto Rico, with colored map, by A. K. Lobeck. Volume II. Part 1. Geology of the Lares District, with colored map, by Bela Hubbard. Part 2. Geology of the Humacao District, with colored map, by Charles R. Fettke. Part 3. Geology of the Fajardo District, with maps and illustrations, by Howard A. MeyerhoCf. Volume III (complete in four parts). Part 1. Tertiary MoUusca from Porto Rico, by C. J. Maury. Part 2. Tertiary Mollusca from the Lares District, by Bela Hubbard. Part 3. Fossil Corals of Porto Rico, by H. N. Coryell and Violet Ohlsen. Part 4. The Tertiary Foraminifera of Porto Rico, by J. J. Galloway and Carohne E. Heminway. Volume IV. Geology of the Virgin Islands, Culebra and Vieques. Part 1. Introduction and Review of the Literature, by J. F. Kemp. Physiography, with colored map, by H. A. Meyerhoff. Part 2. Physiography (concluded), with two colored maps, by H. A. Meyer- hoff. Volume V (complete in four parts, with generic index). Part 1. Descriptive Flora— Spermatophyta (Part), by N. L. Britton and Percy Wilson. Part 2. Descriptive Flora — Spermatophyta (Continued). Part 3. Descriptive Flora— Spermatophyta (Continued). Part 4. Descriptive Flora— Spermatophyta (Continued). Volume VI (complete in four parts). Part 1 Descriptive Flora— Spermatophyta (Continued). Part 2. Descriptive Flora— Spermatophyta (Continued). Part 3. Descriptive Flora— Spermatophyta, with Appendix (Concluded). Part 4. Supplement to Descriptive Flora— Bibliography. Index to Volumes V and VI. Volume VII. ' Part 1. Plant Ecology of Porto Rico, by H. A. Gleason and Mel T. Cook. Part 2. Plant Ecology of Porto Rico (Continued). Part 3. Paleobotany of Porto Rico, by Arthur Holhck. Volume VIII. Part 1 Mycology, by Fred J. Seaver and Carlos E. Chardon, with contribu- tions by Rafael A. Toro; F. D. Kern and H. H. Whetzel; and L. 0. Overholts. Part 2. Supplement to Mycology, by Fred J. Seaver, Carlos E. Chardon, Rafael A. Toro, and Frank D. Kern. Revision of the Myxomycetes, by Robert Hagelstein, Myxophyceae, by Nathaniel L. Gardner. Part 3. The Diatomaceae of Porto Rico and the Virgin Islands, by Robert Hagelstein. Volume IX (complete in four parts, with index). Part 1. Mammals — Chiroptera and Insectivora, by H. E. Anthony. Part 2. Mammals — Rodentia and Edentata, by H. E. Anthony. Part 3. Birds — Colymbiformes to Columbiformes, by A. Wetmore. Part 4. Birds — Psittaciformes to Passeriformes, by A. Wetmore. Volume X (complete in four parts, with index). Part 1. Amphibians and Land Reptiles of Porto Rico, by Karl Patterson Schmidt. Part 2. The Fishes of Porto Rico and the Virgin Islands — Branchiostomidae to Sciaenidae, by J. T. Nichols. Part 3. The Fishes of Porto Rico and the Virgin Islands — Pomacentridae to Ogcocephalidae, by J. T. Nichols. Part 4. The Ascidians of Porto Rico and the Virgin Islands, by Willard G, Van Name. Volume XI, Part 1. Insects of Porto Rico and Virgin Islands — Diptera or Two-winged Fhes, by C. H. Curran. Volume XII. Part 1. Insects of Porto Rico and the Virgin Islands — Heterocera or Moths (excepting the Noctuidae, Geometridae and Pyralididae), by W. T. M. Forbes. Supplementary Report on the Heterocera of Porto Rico, by W. T, M. Forbes. Part 2. Insects of Porto Rico and the Virgin Islands — Moths of the Family Noctuidae, by William Schaus. Part 3. Insects of Porto Rico and the Virgin Islands — Moths of the FamiUes Geometridae and Pyralididae, by Wilham Schaus, Volume XIV. Part 1. Insects of Porto Rico and the Virgin Islands — Odonata or Dragon FUes, by Elsie Broughton Klots. Part 2. Insects of Porto Rico and the Virgin Islands — Homoptera (excepting the Stemomynchi), by Herbert Osborn. Part 3. Insects of Porto Rico and the Virgin Islands — Hemiptera-Heterop- tera (excepting the Miridae and Corixidse), by Harry Gardner Barber. Volume XV, Part 1. The Brachyuran Crabs of Porto Rico and the Virgin Islands, by Mary J. Rathbun. Part 2. Crustacea Macrura of Porto Rico and the Virgin Islands, by Waldo L. Schmitt. Amphipoda of Porto Rico and the Virgin Islands by Clarence L. Shoemaker. Volume XVI. Part 1. A Handbook of the Littoral Echiuoderms of Porto Rico and the Other West Indian Islands, by Hubert Lyman Clark. Part 2. Polychaetous Annelids of Porto Rico and Vicinity, by Aaron Louis Treadwell, Part 3. Bryozoa of Porto Rico with a Resume of the West Indian Biyozoan Fauna, by Raymond C. Osburn, Volume XVIII. Part 1. Porto Rican Archaeology, by Froelich G. Raiuey. Part 2. A Large Archaeological Site at Capa, Utuado, with Notes on Other Porto Rico Sites Visited in 1914-1915, by J. Alden Mason. Appen- dix. — An Analysis of the Artifacts of the 1914-1915 Porto Rican Survey, by Irving Rouse. The above parts may be procured at $2 each by addressing The Executive Sec- retai-y, New York Academy of Sciences, care of the American Museum of Natural Histoi-y, New York, N. Y. As now projected, this Series will include eighteen vol- umes of four parts each.